全 文 ：云南贡山异叶苣苔属(苦苣苔科)一新种
1 , 2　王 印 政　2李　恒
1(中国科学院植物研究所　北京 100093)　2(中国科学院昆明植物研究所　昆明 650204)
A New species of Whytockia W.W.Smith(Gesneriaceae)
from Yunnan , China
1 ,2　WANG Yin-Zheng　2LI Heng
1(Institute of Botany , the Chinese Academy of Sciences , Beijing 100093)
2(Kunming Institute of Botany , the Chinese Academy of Sciences , Kunming 650204)
Key words　Whytockia;W.gongshanensis Yin Z.Wang et H.Li;New species;Yunnan , China
关键词　异叶苣苔属;贡山异叶苣苔;新种;中国云南
贡山异叶苣苔　新种　图 1
Whytockia gongshanensis Yin Z.Wang et H.Li , sp.nov.Fig.1.
Affinis W.tsiangianae (Hand.-Mazz.)A.Weber , a qua sepalis latioribus , corolla pallide
caesio-purpurea , stylo ovario ulta 2-plo longiore , stigmate paene infundibulari differt.
Herba perennis.Rhizoma repens.Caules erecti , basi decumbentes , succulenti , 10 ～ 25 cm
alti , 6 ～ 8-nodes , saepe simplices , inferne glabrescentes , superne brunneolo-pubescentes et glan-
dulosi.Folia opposita , binatim valde inaequalia;ea normalia sessilia vel subsessilia , laminis mem-
branaceis , ovatis vel ovato-oblongis , 3 ～ 6 cm longis , 1.7 ～ 3 cm latis , apice acutis vel obtusius-
culis , basi in latere angusto cuneatis , in latere lato rotundatis , margine irregulariter remote repando-
serratis et glandulosis , utrinque sparse pubescentibus , nervis lateralibus in latere lato 6 ～ 11 , in
latere angusto 4 ～ 6 , petiolis 0.5 ～ 2 mm longis vel nullis;ea reducta sessilia , ovata , 3 ～ 10 mm
longa , ceterum ut in eis normalibus.Cymae axillares , 4 ～ 6 cm longae , 2 ～ 5-florae , pedunculis 2
～ 4 cm longis , cum pedicellis pubescentibus et glandulosis , pedicellis 7 ～ 15 mm longis.Calyx
campanulatus , 3 ～ 3.5mm longus , sepalis 5 , liberis , basi tantum connatis , ovatis vel ovato-orbic-
ularibus , apice obtusis vel acutis , extus puberulis , intus glabris , 3 ～ 3.3 mm longis , 2.2 ～ 2.4
mm latis.Corolla pallide caesio-purpurea , 8 ～ 10 mm longa , extus glabra , intus infra faucem inter
lobos anticos pilis unicellularibus claviformibus bifariis tecta , tubo circ.5 ～ 7mm longo , labio pos-
tico 1.5 ～ 2mm longo , ad medium 2-fido , labio antico 3 ～ 4 mm longo , 3-partito.Stamina 4 , fila-
mentis 2 mm supra basin corollae insertis , superne incrassatis et ciliatis , anticis 4.5 mm longis ,
prope medium geniculatis , posticis circ.3 mm longis , antheris 1.1 mm latis , pubescentibus.
Staminodium longe ovatum , circ.0.7mm longum.Discus circ.0.2mm altus.Pistillum 4 ～ 5mm
国家自然科学基金资助项目(39770063 , 39870056)。
1999-04-21收稿 ,1999-07-15收修改稿。
植 物 分 类 学 报　38(2):189～ 191(2000)
Acta Phytotaxonomica Sinica
图 1　贡山异叶苣苔　1.植株;2.花萼;3.花冠剖开;4.雄蕊;5.雌蕊;6.宿存花萼与不规则
开裂的成熟蒴果。(肖溶 ,孙英宝 绘)
Fig.1　Whytockia gongshanensis Yin Z.Wang et H.Li　1.Habit;2.Calyx;3.Opened corolla;4.Stamen;5.Pistil;
6.Persistent calyx and i rregular dehiscence of mature capsule.
190　 植　物　分　类　学　报 38 卷
longum , ovario ovoideo 0.8 ～ 1.1 mm longo , stylo 2.1 ～ 2.8 mm longo , ovario ultra 2 ～ 2.5-plo
longiore , stigmate infundibulari.Capsulae globosae , 3 ～ 3.4 mm diam., glabrae , irregulariter de-
hiscentes , in calycibus persistentibus omnino inclusae.
Perennial , succulent herbs.Rhizomes creeping , bearing roots at nodes.Stems erect , 10 ～ 25
cm tall with 6 ～ 8 nodes , simple and often decumbent at base , inferior glabrescent , superior covered
with brown-pubescences and glandular hairs.Leaves opposite , quite unequal.Large leaf blades in a
pair membranaceous , ovate and oblong-ovate , 3 ～ 6 cm×1.7 ～ 3 cm , sessile or with a short petiole
0.5 ～ 2 mm long , apex acute or obtuse , base oblique , rounded in wide lateral and cuneate in an-
gustate lateral , both surfaces pubescent , margins irregularly repand-serrate and glandular-serrate ,
nerves 6 ～ 11 in wide lateral and 4 ～ 6 in angustate lateral.Small leaf blades ovate , 3 ～ 10 mm
long , sessile , otherwise as larger leaf.Inflorescences pair-flowered cymes , 4 ～ 6 cm long , emerging
from the axils of the large leaves , with 2 ～ 5 pedicellate flowers , ebracteolate.Peduncles 2 ～ 4 cm
long and pedicels 7 ～ 15 mm long , covered with pubescences and glandular hairs.Calyx campanu-
late , 3 ～ 3.5 mm long.Sepals five and connate at base , ovate or ovate-orbiculate , 3 ～ 3.3 mm×
2.2 ～ 2.4 mm , outside pubescent and glandulose , inside glabrous , apex acuminate.Corolla blue-
purpureous , tubular and bilabiate , 8 ～ 10 mm long , and 5 ～ 7 mm long before limb , outside
glabrous , inside with two rows of the unicellar and clavate hairs under between lobes of the lower
lip;upper lips 1.5 ～ 2 mm long , 2 ～ lobed;lower lips 3 ～ 4 mm long , 3-lobed.Stamens 4 , at-
tached to corolla base for 2 mm long.Filaments incrassate on upper parts covered with sericeous
hairs;the anterior pair about 4.5 mm long , geniculate near middle;the posterior pair about 3 mm
long.Anthers connate , 1.1 mm wide , with sericeous hairs.Staminode obovate , 0.7 mm long.
Disc about 0.2 mm high.Pistiles 4 ～ 5 mm long;ovaries ovate , 0.8 ～ 1.1 mm long;styles 2.1 ～
2.8 mm long and 2 ～ 2.5 time longer than ovaries;stigmas sub-infundibulate.Capsules globate , 3
～ 3.4 mm diam., glabrous , irregular dehiscent , included in the persistent calyx.Seeds minute ,
many.
China.Yunnan(云南):Gongshan(贡山), Dulongjiang (独龙江), along Dulongjiang Riv-
er valley at alt.1350 m , by stream in shady and moist places , 1990-11-14 , Dulongjiang Exped.
(独龙江考察队)283(holotype , KUN).
Whytockia gongshanensis resembles W.tsiangiana on the basis of the small corolla (8 ～ 12
mm long)and the connate stigma.However , it is easily distinquished from W.tsiangiana by the
blue-purpureous corolla , the styles 2 ～ 2.5 time longer than the ovaries , and the sub-infundibulate
stigmas.
Additional discussion:W.gongshanensis is known only from the type locality in Dulongjiang
River valley up to date , growing by stream in shady and moist places at about alt.1350 m.Du-
longjiang River valley belongs to the Hengduan mountain.The region is well-known in the world for
its complicate and unique geological structure and floristic characteristics.The sub-infundibulate
stigmas of W.gongshanensis constitutes a new feature for the genus Whytockia.Correlatively , its
geographical distribution contributes to the genus not only a new geographical record of the genus but
also a peculiar pattern of distribution that is difficult to interpret with normal dispersal.
致谢　本文承蒙王文采 、李振宇和杨亲二先生审阅并提出修改意见 ,特此致谢。
2 期 王印政等:云南贡山异叶苣苔属(苦苣苔科)一新种 191