Abstract:The root-stem transition zone in Panicum miliaceum L. is from the base of hypocotyl to coleoptilar node. The general trend of transformation of vascular tissue in the transition zone is the same in Zea mays L. Triticum vulgate Viii., and Avena sativa L.. The first step in the transition is from the exarch to the endarch; and the first leaf above the coleoptile come from branches of cotyledon trace; and midrib of the first leaf come from vascular bundle in axis. But cotyledon trace branches at cotyledon node in Panicum miliaceum L. which is the same in Zea mays L. but differs from Triticum vulgare L. and Avena sativa L. There are inversion, splitting and mergence in the course of transformation. But splitting and mergence are different from traditional patterns. There were anticlinal and periclinal splitting of xylem bundles in transformation of xylem. The two neighbour xylem bundles are linked by vessel elements from new differentiation. The transformation of phloemis completed by mergence and resplitting. Adventitious roots develop after transition is completed.