Abstract:Macrospore metrocyte of Pinus sylvestris var. mongolica developed to female gametophyte containing female gamete-egg by a series of changes. Macrospore metrocyte formed on June 8~14 every year, then made a meiosis and macrospore formed on June 16~20 in Harbin. With the nucleus divisions of macrospore, macrospore went to the phase of free nuclei and the cell wall formed from May 28 to June 4 in the next year. The archegonial initial cell emerged in young female gametophyte and the neck cell and the central cell formed after it divided once. Central cell was divided to egg cell and ventralcanal cell on June 7~9, archegonia developped to mature on June 13~15. The mature archegonia contained neck cell, ventralcanal cell and egg cell, but neck cell and ventralcanal cell has degenerated.