全 文 ：鳝{欲雇
广 西 植 物 Guihaia 19(2)：105— 118
泡鳞 扁 豸
中国鳞毛蕨属泡鳞亚属的分类研究
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(云卣大学生态学与地植物学础宽所， 昆明 650091)
A taxonomical study on the genus Dryopteris subgenus
Erythrovariae from China
Lu Shugang
Institute Ecology andGeobotany．YunnunUniversity．K unmins 650091)
Abstract In this paper．a taxonomical revision on the genus Dryopteris subgen Erythrovariae(H．
Ito)Fraser—Jenkins from China is made,A new，revised s~tem is proposed．34 species and 2 varie-
ties are recognized and placed in 3 sections and 2 series．Of which，section Indusiatae S．G ．Lu and
series Chart1Pionianae Ching ex S．G．Lu are described as new．A key to the sections，series and spe -
c Jes is given．Each is provided with literature citations，synonymies and distributions．
Key words Dryopteris subgen．Erythrovariae；taxonomical revision；China
鳞毛蕨属
Dryopteris Adans．
泡鳞亚居
Subgenus Erythrovariae(H．Ito)Fraser-Jenkins in Bul1．Br．Mus．Nat．Hist．Bot．14(3)：195．
1986．
Type：Dryopteris erythrosora(D Eaton)Kuntze．
This subgenus more or less covered with bullate scales on the underside of costae and costules，or
at least lanceolate scales with$accaLe base．It contains about 45 species．distributed mainly in Asia．
China and Japan is its centre of distribution a few species extending to New Guinea，Indo-China and
E．Himalays．34 spe cies and 2 varieties ale recognized in China．
Key to sections，series and species
． Lamina [anceolate or ovate-[anceolate,usually pinnate to bipinnate；lowest basisco pic pinnute on the
[owest pinna not markedly longer than those above；p Jnnuleapex rounded，not caudate．
2．Upper parts of stipes bearing many sinai[sca~s：scales OU the base of stipes lanceo[ate，brown or light
brown rSect．I．Erythrovariae H．Ito)．
1998—05—13收藕
作者简介：陆树刚， ，1957年出生，硕士，副教授．植物分类学研究方向
国家 自然科学基金重大项 目 (项目编号393918001)
．
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06 广 西 植 物 】9卷
3．Scales on the rachis sparsely，linear—lanceolate，thick，entire(Set．1．Erythrovariae)．
4．Lamina pinnate，pinna serrate or pinnatifid at margin．
5．Lamina pinnate．pinna serrate at margin．shalowly cordate at base ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯⋯ ⋯ ⋯ ⋯
⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ 1．D．decipiens(Hook．】O．Kuntze
5．Lamina pinnate．pinna pinnatifd at margin．cordate at base⋯ ⋯ ‘⋯ ⋯ ‘⋯ ⋯ ⋯ ’⋯ ‘⋯ ⋯ ⋯ ‘⋯ ⋯ ‘
· ⋯ ⋯ ⋯ ⋯ ⋯ - -⋯ ⋯ ⋯ ⋯ ⋯ ⋯ - ·⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ · 2．D．deeigiens var．dipla~oides(Christ)Ching
4．Lamina bipinnate，pinnule with stalks at base．
6．Pinnule triangular-ovate，serrate or pirmatifida at margin obtuse at apex．
7．Pinnule serrate at margin；sod near to cesta ·⋯·⋯··⋯ ·⋯ ⋯ ⋯·⋯ ⋯ ·⋯ 一 3．D．fusdpes C．Chr．
7．Pinnule pinnatifida at ma rgin；soft close to ma rgin⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ 4．D．ryo—itoana Kurata
6．Pinnule lanceolate,pinnately lobed or pinnately parted at margin，acuminate at apex．
8．Pinnule pinnately lobed；bulate scales densely on the ce sta and rachis；indusia red dish-brown
on the centre ⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯ 5．D．erythrosora(Eato n1 O．Kuntze
8．Pinnule pinnately parted ；bullate scales spa rsely on the ce sta and rachis；indusia light reddish
on the centre⋯ ⋯ ⋯ ··⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ··⋯ ⋯ ⋯ ·⋯ ⋯ ⋯ ⋯ ⋯ ·⋯ ⋯ ·⋯ ⋯ ··6．D．hondoensis Koidz．
3．Scales on the rachis densely，ovate-lancce late or linear-lanceolate scales exist simultaneously，
thin，toothed or entire(Ser．2．Championianae Ching ex S．G．Lu
9．Sca les on the rachis ovate，toothed at ma rgin．
10 Lamina bipinnate pinnule entire or serrate at margin；scales on the rachis dark brown ⋯ ‘⋯ ·‘
⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ·⋯ ⋯ ⋯ ··7．D．1epidorachis C．Chr．
10．Lamina tripinnate or bipinna te．pinnules lobed ；scales on the rachis brown ⋯ ·⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ··
⋯ ⋯ ⋯ ··· ⋯ ⋯ · ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ·⋯ ⋯ · ⋯ ⋯ · ·⋯ ⋯ ·· ⋯ ⋯ · ⋯ ⋯ ⋯ ⋯ ·⋯ ·· · 8．D．tsoongii Ching
9．Scales on the rachis lanceolate or linear--lanceolate，entire or toothed at margin．
1 1．Upper pa rts ofstipes and rachis bearing only scattered scales‘⋯ ⋯ ‘⋯·9．D．kinkieusis Koidz．
1 1．Upper pa rts ofstipes and rachis densely covered with sca les．
12．Pinnae sessile，hardly adnate at base；scales on the rachis us ually entire．
13．Lamina tripinnate or bipinnate，pinnules pinnatifid ⋯⋯⋯ 10．D．simasakii(H．Ito)Kurata
13．Lamina bipinnate．pinnules pinnately lobed；scales much more densely⋯ ⋯ ⋯ ⋯ ⋯ ⋯⋯ ’‘’⋯
⋯⋯⋯⋯⋯⋯ ⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯ lL n simasakii var．paleaeea(H．Ito)Kurata
12．Pinnae with distinct stalks；scales on the rachis toothed at nlar n ⋯ ⋯ ⋯ ⋯ ⋯ ⋯⋯ ⋯ ⋯ ⋯ ⋯
⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ 12．D．clmmpionii(Benth．1 C．Chr．
2．Uppe r parts of stipe s glabrous ；sca les on the base ofstipes linear-lanceolate，black or blackish—
brown(Sect．2．1ndusiatae S．G．Lu)．
14．Soft exindusia~；pinna e opposite,hardly sessile；lowest basisce pic pinnule on the lowest pinna
not shortened ⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯ l3．I)．gymnosora(Makino)C．Chr．
】4．Sori indusiate．
15．Pinnae sessile or subsessile；lateral pinnae horizontally pa tent from rachis．
I6．Basal pinnule on lower pinnae shortened and parallel to the rachis，overlaping the rachis
⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯ 14．I)．indusiata(Makino)Yamamoto
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16．Basal pinnule on lower pinnae not shortened and not paralel to the rachis．
17．Lam ina bipinnate；serrates with long aristate on top；stipe8 usually purpure 。。。‘⋯ ‘。⋯ ⋯ ‘⋯ ‘。
⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ l5．D．tenuicula M atthew et Christ
17．Laminatripinnate；pinnules pinnatisect,serrateswithoutlong aristateontop；stipe s stramineous
⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ l6． D．submargina~ RosensL
15．Pinnae with distinct stalks；lower lateral pinnae ascending，not at right angles to rachis．
18．Pinnae pinnatisect．pinnules adnate to pinna rachis···⋯ ⋯ ·⋯ ⋯ ··17．D．guangxiensis S．G．Lu
18．Pinnules with stalks or subsessile，not adnate to pinna rachis．
19．Pinnules oblong，rounded at base，rounded or truncate at apex,usualy entire at margin．
20．Scales on stipe and rachis black；pinnules rounded at apex with spiny serrate；sori near to
COSta ⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯ l8．D．tenuipes(RosensL1 Serizawa
20．Scales on uppe r parts of stipe s and rachis brown，pinnules rounded or truncate at ape x，With—
out spiny serrate；soft media1．
21．Lamina broadly deltoid，basal pinnae much more longer than those above；pinnules
usua~y rounded at ape x⋯ ⋯ ⋯⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ 19．D．subtriangularis rHope 1 C．Chr．
21．Lamina ovate—lanceolate；pinnules usually truncate at apex ⋯ ’⋯ 。⋯ ⋯ ⋯ ’。。⋯ 。⋯ ‘’。。⋯ ‘’’。。
⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ 20．I)．assamensis(Hope、C．Chr．et Ching
19．Pinnules deltoid-ovate，broadest at base，acuminate or obtuse at apex，pinnatilobate to
pinnatisect at m argin．
22．Pinnules cordate at base with 2～ 3 mm long stalks．
23．Scales on rachis and pinna rachis black；soft slightly near to margin·⋯ ⋯ ⋯ ·⋯ ⋯ ⋯‘‘‘⋯ ‘’‘。
⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ·⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ··21．D．integriloba C．Chr．
23．Scales on rachis and pinna rachis brown；sori slightly near to costa ⋯ ⋯ ·⋯ ⋯ ⋯ ·⋯··⋯ ·⋯ ·
⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ 22．n tahmingensis Ching
22．Pinnules rounded or cuneate at base，sessile or subsessile．
24．Scales on base of stipe s black；lowest posterior pinnules on the lowest pinnae much
longer than those above pinnatipartite to pinnatisect ·⋯ ···⋯ ·⋯ ······⋯ ····⋯ ⋯ ··⋯ ··⋯ ·
⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯ 23．D．1abordei(Christ)C．Chr．
24．Scales on base ofstipe s dark brown；lowest posterior pinnules on the lowest pinnae
slightly longer than those above pinnatilobate to pinnatifd⋯ ···⋯ ⋯ ⋯ ·⋯ ⋯ ·⋯ ⋯⋯ ·⋯ ⋯
⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯⋯ ⋯ ⋯ ⋯⋯ ⋯ ⋯⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ 24．n pseudosparsa Ching
1．Lamina pentagonal-ovate，us ually tripinnate；lowest basiscopic pinnule on the lowest pinna
markedly longer than those above；pinnule apex caudate and acutely po inted rsecL 1 Varhe
Fraser—Jenkins)．
25．Basal pinnaewith 3～ 4 cm long stalks．
26．Sori induslate．
27．Scales on stipe ．rachia and pinna rachis black；indusia entire at margin ⋯ ···⋯ ⋯ ·⋯ ⋯ ⋯ ·⋯ ⋯·
⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ 25．I sordidipes Tagawa
27．Scales on stipe．rachis and pinna rachis brown；indusia with ciliate at margin ⋯⋯ ⋯ ⋯ ⋯ ⋯ ⋯
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⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ 26．D．xungwuensis Shing ot J．F．Cheng
26．Sori exindns iate ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ 27．D．dehnaensis Ching et Shing
25．Basal pianne with about l Gnl long stalks．
28．Lamina pentagonous
．
as long as broad,tripinnate．ultimate pinnules pinnatipartite．
29．Scales on stipes light brown；pinnae and pinnules with distinct stalks⋯ ⋯ ·⋯ ·⋯ ⋯ ⋯ ‘⋯ ⋯ ‘⋯ ”
··⋯ ⋯ ⋯ ·⋯ ⋯ ··⋯ ·⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ·⋯ ⋯ ··⋯ ⋯ ⋯ ·⋯ ⋯ ·⋯ ⋯ ···⋯ ⋯ ⋯ ·28． D amurensis Christ
29．Scales on stipes black；pinnae and pinnules subsessile⋯⋯⋯ 29．D forMosBila fChrist)C．Chr．
28．Lamina deltoid~oblong or ovate，much longer than broad，bipinnate or tripinnate，ultimate
，
pinnules entire or pirmatilobate．
30．Ultimate pinnules or segm ents ovate；stipes densely clothed with brown，linearlancco late
scales at base ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ 30．D．yongdeens~ w _M ．Chu ex S．G．Lu
30．U ltima te pinnules or segments lanceolate；stipes densesly covered with black
，
dark brown or
bico lor scales at base．
3 1．Scales on the top of rhizome and on the base of stipes co mpletely black⋯ ⋯ ⋯ ⋯⋯ ⋯ ⋯ ⋯ ⋯
⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯ 31．I)．padfica(Nakai Tagawa
31．Sca les onthetoPof rhizome and 0nthe base ofstipes dark brown or bico lor．
32．Scales on stipes brown,hair-like at apex．
33．Lamina bipinnate，ca udate-acuminate at apex；sod larger,usually in one row on either
side of pinna rachis·⋯ ···⋯··⋯ ⋯ ··⋯ ⋯··⋯ ·⋯⋯ ⋯ ··⋯ ··⋯ ⋯⋯ ···32．D immixta Ching
33．Lamina bipinnate to tripinnate
， acute at apex；sori sma ller，scattered 0n either side of
pinna tachis⋯ ⋯⋯⋯⋯⋯ ⋯⋯⋯⋯⋯⋯⋯⋯·，⋯⋯⋯⋯⋯⋯·33．D．varia(L．)O．Kuntze
32．Scales oR stipe s bicolor,black on centre，brown towards margins．
34．Scales on stipes and rachis deflexed ⋯ ⋯ ⋯ ⋯ ⋯ ⋯⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ 34．D．saxifraga H．Ito
34．Scales on stipes and rachis not deflexed．
35．Rachis and pinna rachis densely covered with bulate sca les ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯⋯ ⋯ ⋯
⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯⋯ 35．D．setnsa(Thunb．1 Akasawa
35 Rachis and pinna rachis parsely clothed with bullate sca les ⋯ ·····⋯ ⋯ ⋯ ·⋯ ⋯ ⋯·⋯ ·····
⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ 3 D．[~terosflnela Koid7．-
组 1．红盖鳞毛蕨组 ．
SecL 1．Erythrovariae H．[tO in BoL Mag．Tokyo 50：32．1936；Fraser-Jenkins in Bul1．Br．Mus．Nat．
Hist．Bot．14：(3)：196．1986．
Type：Dryopteris erythrosora(Eato n1 Kunt~．
This section contains 2 series．
系 1．红盖鳞毛藏系
Ser．1．Erythrovarlae
Type：Dryopteris erythrosora(Eato n1 Kuntze．
This series contains 5 species and 1 variety in China．
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2期 陆树刚： 中国鳞毛蕨属泡鳞亚属的分类研究 109
1．迷人鳞毛蕨
Dryopteris deciplens(Hook．)O．Kuntze in Rev．Gen．PI．2：8 12．1891：C．Chr-．Ind．Fil．1906；Supp1．3．
84．I934；ChinginBul1．FanM em．Inst．Bio1．BOL 8：475．1938：W U．W ang etPongin Bul1．DepLBio1．Sun
Yatsen Univ．3：24．p1．3．I932；Nakaike．New FI．Jap．Pterid．407．t．407．I992；FI．Fujian．1：I97 I982；FI．
Anhui．1：146．￡ 140．1985；FI．Zhejiang．1：408．t．I一255．1993：FI．Jiangxi．1：278． 282．I993．
— — 印 ，口 圳 decipiens Hook． Sp．Fil．4：86．￡243．186Z——一Aspidium decipiens(Hook．)Luers．in
Eng1．Bot Jahrb．4：360．1883．
Sichuan，Guizhou,Guangxi，Guangdong Fujian，Jiangxi，Anhui，Zhejiang and Hunan；Japan．Type
from Fuj Jan．
2．深裂鳞毛蔗
Dryopteris decipiens var．dlplazioides(Chr~t1 Ching in Bul1．Fan Mere．Inst．Bio1．Bot 8：476．1938，
Nakaike，New FI．Ja Pterid．408． 408．1992；F1．Jiangxi 1：279 1993 b f M diplazioides
Christin Bul1．Acad．Geogr．Bot．1902：260．1902；C．Chr．
， Ind．FiL 581．1906．— — Dryopte7 mscipes
diplazioides(Christ)Ching，FI．Jiangsu l：66．1977；FI．Anhui 1：149．1985；FI．Zh@ang 1：252．1993．
— — Dry。pter integripinna Ching in Wuyi Sci．Journ．1 t、、 8．198 1．— — D opter decipie#Is vat．
subbipinnata Ching in Wuyi Sci．Journ．1 t、、1 8．1981．— — Dryopteris cordipinna Ching et Shing in FI．
Fujian l：197．I982．— — D 印fP山 metqfus~ipes Ching et C． Zhang in Bul1．Bot．Res．3(3)：19． 15．
1983；FI-Zhej．1：25 1．1993．— — D 印fPr relroSo-pateacea Ching et C．F．Zhang in Bul1．Bot．Res．3(3)：
20．1983；FI．zhej．h 254 1993 D 印lPr mCmeticaChing etC．F．ZhanginBul1．BOLRes．3(3)：2I．
I6．1983；FI．zhej．1：249． 卜250．1993．
Sichuan，Guizhou，Jiangxi
，Fujian，Anhui，Jiangsu and Zhejiang；Japan．Type from Guizhou．
3．黑足鳞毛藤
DryoptedsfuscipesC．Chr．．Ind．Fil．Supp1．2：14．1917：ChinginBul1．FanM ere．Inst．Bio1．Bot．8：476．
1938；Nakaike，New FI．Jap．Pterid．417． 417a．417b．I992；S．G．Lu in Actz Phytotax．Sin．28(6)：474．
1990；Ic-Corm．Sin．1：239． 477．1992；FI．Jiangsu 1：65． 96．1977；FI．FujJan h 199．￡185．1982；FI．
Anhui h I48． 143．1985；FI．Zhej．1：251． 卜259．1993；FI．Jiangxi l：280． 284 1993 D 叩fP，
confertipinna Ching et Shing，FI．Fujian 1：199．600 1982 D 印tP， muh~ugata Ching et Shing in
Bul1．Bot．Res．3(3)：12．￡10．1983．— — D 印lPr r 胁 Ching et C．F．Zhang in Bul1．Bot．Res．3(3)：
16． 13 1983 D，)．印fP parafuscipes Ching et Z．Y．Liu in Bul Bot．Res．4(3)：23． 26．1984(non
Ching et Chiu，1987)．— — D仰 fPr mediaBsora Ching et Chiu in Bot．Res．2：6．PI．2 ￡4．1987．
— — D 0 Pr stenochlamysChing etChiu exShing et J．G．Chengin JiangxiSci．Journ．8(3)：49．1990；FI．
Jiangxi l：280- 285—1993 D 叩fP， ^scipes form．major Ching ex Shing et J．F．Cheng in Jiangxi
Sci-Journ．8【3)：49-1990：FI．Jiangxil：280．1993．— — D 卵fPr y∞ ChinginBul1．Res．2(2) 64．PI．1．￡
3．1982．
Yunnan，Sichuan，G uizhou
， Guangxi，Guangdong,Hunan，Hubei，Jiangxi． Fujian，Jiangsu，Zhejiang,
Anhui and Taiwan；Japa n
． Korea and Indo-China．Type from G uizhou．
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O 广 西 植 物 9卷
4．宽羽鳞毛蕨
Dryopteris ryo-ltoana Kurata in Journ．Geobot．15：84．]967；Nakaike
， Enum．Pterid．Jap．Fil．227．
1975；New FI．Jap．Pterid．441． 441．1992．—— _D opteris triangula o Ching in Bul1．Bot．Res．2(2)：
64． 】-4．】982．—— _D y叩 Pr dispar Ching et C．F．Zhang in Bul1 Bot．Res．3(3)：】7．￡ 】4．1983；FI．
Zhejiang l：255．1993．一——D o ，括kaihuaensis Ching et C．F．Zhang in Bul1．Bot．Res．3(3)：24． 】8．
1983：FI．Zhejiang l：255．1993．一——D 叩 P， pseudofuscipes Ching et Chiu in Bot．Res．2：3．PI．】． 3．
】987．— — D 叩 搪，括lushanensisChing et Chlu in Bot．R ．2：20．PI．8． 】．】987；FI．Jiangxi l：286．￡ 292．
】993．
Zh~iang and Jiangx~；Japan．Type from Japan．
5．红盖鳞毛蕨
Dryopteris erythrosora(Eaton)O．Kunt~e in Rev．Gen．PI．2：812．189l，C．Chr．．1nd．Fil．263．1906；
Ching in Bul1．Fan Mere．Inst．B；0I_BOL＆ 481．1938；S．G．Lu in Acta Phytotax．Sin．28(6)：473．1990；
Nakaike．New Ft．Jap．Pterid．4 L0．￡410a．410h 1992：Ft．Jiangsu l：66．￡99．]977；F】_Anhui l：]49
． ￡ 145．
1985：F】_Zhejhng l：253．f l-260．1993；FI．Jhngxi l：279． 283．1993．— — 州eryhrosorum Eaton
jn Perry’Narr．Exp．China 2：330．】856．— — 工 ” erythrosora(Eaton)Moore．Ind．Fil．91．】858．
— — Ⅳ ，o “ erythrosorum(Eaton)Hook．，Sp．Fil．4：120．￡253．1862．— — DrJ，叩fPr bulligeraChing
in Bul1．Fan M em．1nsL Bio1．BOL＆ 483 1938 Dryo Dleris remotlpinnuta Ching et C．F．Zhang in Bul1．
Bot Res一3(3)：15一 】2．1983；FI．Zhejhngl：255．1993．— — D 叩fP sino-erythrosoraChing et Shingin
FI．Fujian l：200，600．1982．———D 括pseudo-erythrosora Ching et C．F．Zhang in Bul1．Bot．R ．3
(3)：27． 20：1983(non kodama，1913l———口 印搪， lfnyingen~isChing etC．F．Zhangin Bul1．BOL R ．
3(3)：30．C 22．1983；FI．Zhejiang l：254．1993．— — D y叩 Pr oppositlpinna Ching et Z．Y_Liu；n Bul1．Bot．
Res．4(3)：25．L 28．1984．— — D 叩f squaraistipes Ching et Z．Y．Liu in Bul1．Bot Res．4(4)：7． 35．
1984．— — |!) 叩 f即 o6如碍 “ Ching et Ch；u in Bot．Res．2：5．PI．2．￡ 2 1987．— — DrvD口fP，
paraerythrosora Ching et C．F．Zhang in FI．Zhejiang l：250． l一258．1993．
Yunnan，Guizhou，Sichuan
， Guangxi，Guangdong,Hunan，Hubei，Jiangxi，Fujhn，Jiangsu，Zhejiang
and Anhui；Japan and Korea．Type from Japan．
6．桃花岛鳞毛藤
Dryopterls hondc~nsis Koid~ In Acta Phytotax
． Geobot．1：31．1932；C．Chr．．1nd．Fi]．Supp1．3：88．
1934；Nakaike，New FI．Jap．Pterid．424．C 424．1992；1watsuki
． Ferns and Fam Allies ofJapan 199．PI．126．
￡4-5．]992；FI Zhejiang l：269．￡1—271．1993．
Zhejiang and Sichuan；Japan and Korea．Type from Japan．
系2 阔鳞鳞毛蕨系
Ser．2．Championianae Ching ex S．G．Lu seL nov．
Stipite uti rhachi paleis rufo—brunneis
，
ovatis vel late]anceolatis
， margine erosis vel integris densissime
obtecto．
Type：Dryopteris championii(Benth．)C．Chr．
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2期 陆树刚： 中国鳞毛蕨属泡鳞亚属的分类研究
This series contains 5 species and 1 vadety in China．
7．轴鳞鳞毛蕨
Dryopteris lepidorachis C．Chr．，Ind．FiL 274．1906(based on Pob,podium rheosorum Bak．，1891)；FI．
Jiangsul：65．￡95．1977：FI．Fujian1：201．1982；FI．Anhui1：147． 14Z1985；Ching etChiuinBot．Res．2：
19．P1．7． 4．1987．— — P0 0 “ rheosorium Bak．In Ann．BOL 5：457．1891(non Bak．1844)．
— — Ⅳ印 r^0 “ t rheosorium(Bak．)Hand．-Mazz．，Symb．Sin．6：24．1929．一 一 D 叩fem championii var．
rheosora(Bak．)Shing in FI．Jiangxi 1：282．1993；C．F．Zhang in F1．Zhejiang 1：250．1993．— — 口 叩fP山
ba~tr innatifda Ching et Z．Y．Liu in Bul1．Bot．Res．4(4)：6．f 34．1984．— — D 卵fPr lepidocauton
Ching etChiuinBot．Res．2：19．PI．￡3．1987．
Zhejiang．Jiangsu,Jiangxi，Anhui and Fujian．Type from Zhejiang．
8．观光鳞毛葳
Dryopterist$oolgiiChinginBot．Res．2：14．PI．6． 1．1987；FI．Jiangxil：282．L 287．1993；F1．Zhejiang
l：260．1993．— — D 叩 ， wuyuanensis Ching in Bot．Res．2：15．P1．6．￡2．1 987．
Anhui，Zhejiang and Jiangxi．Type from Anhui．
9．京鸡鳞毛葳
Dryo~eris kinkiensis Koidz．ex Tagawa in Acta Phytotax．Geobot．2：200．1933：C．Chr．．1nd．Fil．
Supp1．3：89．1934；Nakaike，New FI．Jap．Pterid．428．1992；FI．Jiangxi 1：273．￡275．1993；FI．Zhejiang 1：
260． l一262旷1993 D r叩 fP，打 zhenanensis Ching et Chiu in BOL Res．2．1 1．PI．5．￡ 1．1987；FI．
Zhejiang 1：258．1993．— — D 叩 ， championi vaT．tenuifrons(H．1to)H．1to in Nakai et Honda，Nova
FI．Jap．4：48．1 939．— — D 印 fP， neoassamenMs Ching in Bull Fan M em．1nsL Bio1．Bot．8：480．1938．
Sichuan，Zhejiang，Jiangxi，Fujian and Guangdong；Japan．Type from Japan．
10．高鳞毛葳
Dryopteris simasakii rH．Ito)Kurata in Jo~rrL Geobot．18：5．1970；Nakaike,New FI．Jap．Pterid．447．
447．1992；1watsuki，Ferns and Fern Atlies of Japan 196．P1．122．￡4．1992．— — D 叩 fP indusiata var．
simasakii H．1to in Journ．Jap．Bot．9：57．￡6．1933．— — D 叩fP s labordei var．simasakii(H．1to)H．1to in
Nakai et Honda，Nova F1．Jap．4：46 1939 D 叩 excelsior Ching et Chiu in Bot．Res．2：8．PI．3．￡
4．PL 4．f．1．、987．— — Dr，0pterts laodianensis Ching et Chiu in BoL Res． 16．P1．6，t．3．1987．
— — D 叩fPr eryhrochtamysChing etZ．Y．LiuinBul1．BOLRes．4(4)：5．￡33．1984．
Yunnan，Guizhou，Sichuan
，Guangxi and Zhejiang；Japan．Type from Japan．
I1．密鳞毛葳
Dryopteris sim saldi var．peleacea(H．1to)Kurata in Journ．Geobok 18：5．1970；Nakaike New FI．Jap．
Pterid．448．L 448．199Z — — D 叩 f州 indusiata var．pateacea H_lto in Joum ．Jap．Bot．9：57．￡7．1933．
— — D 印fP， labordei vat．simasaki form．paleacea(H．1to)H．1to in Nakai et Honda，Nova F1．Jap．4=
46 1939 Dryopter rufosquamosa Ching et Chiu in Bot．Res． 6．PI．3．E．I．1987．— — Dryo Bleris
paomowanensi$Ching etZ．Y．Liu in Bul1．Bot．Res．4(4)：10．￡38．1984．
Yunnan，Sichuan，Guizhou and Guangxi；Japan．Type from Japa n．
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2 广 西 植 物 9卷
12．阔鳞鳞毛蕨
Dryopteris championi(Benth．1 C．Chr．apud Ching in Sinensia 3：327．1933；C．Chr．，Ind．Fil．Supp]．3：
83．1934；Ching in Bul1．Fan M ere．Inst．Bio1．Bot．8：486．1938；Ic．Fi]．Sin．4：181．PI．181．1937：Ic,Corm．
Sin．1：239． 478．1972；F1．Jiangsu 1：66． 97．1977；F】．Fuj]an 1：197．f．183．1992；F／．Anhui h 152．f．14g．
1985：Ching et Chiu in Bot．Res．2：12．PI．5． 2．1987；S．G．Lu in Acta Phytotax．Sin．28(6)：473．1990；
Nakaike．New FI．Jap．Pterid．402．￡402a．402b．1992；FI．Zhejiang 1：249． 1—257．1993．H．Jiangxi 1：281．
￡ 286．1993．— — 础 “ l championi Benth__FI．Hongk．456．186】．— — D 叩 f changii Ching in
Wuyi Sci．Journ．1：7．]98 1．— — D 叩fPⅢ wuyishanensis Ching in F】．Fujian 1：198．￡ 1 84．]98Z
— — Dr，opter bulatipateaceaChingin Bul1．BoLRes．2l ：65．t．2-1．1982．— — Dryopter yandongensis
Ching et C．F．Zhang in Bul1．BoL Res．3(3)：13．L 11．1983；F1．Zhejiang l：256．1993．一——D 叩fPr打
linganensis Ching et C．F．Zhang in Bul1．Bot．Res．3(3)：23．e 17．1983：FI．Zh@ang 1：250．1993．
— — D 叩fP gutishanensisChing et C． Zhang in Bul1．Bot．Res．3(3)：29． 21．1983；F1．Zhejiang 1：
257．1993．— — D 叩fP， nanehuanensis Ching et Z．Y．Liu in Bul1．Bot．Res．3(4)：20． 18．1983．
— — D 叩 faoshanens~s J．X．Li et S．￡Ma in Bul1．BoL Res．3(4)：139—14](cum．fig．)．1983．
— — Drvopter infrahirtetla Ching et Z．Y．Liu in Bul1．Bot．Res．3 、1 22．t．、e．1983．一 Dryopte7{s
neofusc P Ching etZ．Y-Liuin Bul1．Bot．Res．4(3)：22． 25．1984．— — D 叩fPr patefie Ching et己
Y．Liu in Bul1．BoL Res．4(4)：2． 30 1984 D PfPr huanglungensis Ching in Bot．Res．2：10．P】．4．
3旷1987 D 叩fe，括,eofuscipesChing etChiuin Bot．Res．2：4．PI．1． 4．1987(nonChing etZ．Y．Liu，
1984)．— — Dry0pter qinyuanensis Ching et Chiu in Bot．Res．2：1．PL 3．f．2．1987(err．D．chinyuanensis)．
— — D7yo p{erts wangii Ching in Bot．Res． 13．PL 5．f．3．1987．— — Dfyoptern conferta Ching et Chiu in
BoL Res．2：1 7．P1．7．￡ 1 1987 D 印 f zunyiensis Ching in Bot．Res．2：18．PI．7．￡ 2 1987．
— — D 印 f grandiosa Ching et Chiu in Bot．Res．2：8．P】．3． 3．1987；FI．Zhoiang l：259．1993．
Yunnan，Sichuan，Guizhou,Guangxi，Guangdong，HongkonK Hunan，Hubei，Jiangxi，Fuj Jan，Jiangsu，
Zhejiang，Shandong and Henan；Japan and Korea．Type from Hongkon~
组 2．黑鳞鳞毛蕨组
Sect．2．Indusialae S．G．Lu
， sect．nov．
Basi stipitis pa leis nigris vel atrobrunneis lineari一]anceolatis dense obtecto，sursum uti rhachi subglabra．
Scales onthebaseofstipes black ornearly black
，
upper parts of stipa s and rachis subglabrous．
Type：Dryopteris indusiata(Makino)Yamamoto．
This section contains 1 2 species in China．
13．裸果鳞毛蕨
Dryopte~s gymnosora(Makino)C．Chr．．Ind．Fil．269．1906；Tagawa,Co1．1]lustr．Jap．Pterid．105．PI．
39．￡2I9．I959；FI．Fujian 1：203． 190．1982；S．G．Lu in Acta Phytotax．Sin．28(6)：472．1990；Nakaike，
New F】．Jap．Pterid．4I9．￡4】9．1992：F1．Zhejiang 1：262． 1-264．1993；F1．Jiangxi 1：284．￡289．1993．
— — Nephrod m gymnosorum M akino in Bot．M ag,Tokyo 13：64．1 899．— — D ryopter tabordei aucL．
non C．Chr．．1906：lc．Corm．Sin．1：240． 480．I972；F】．F iang l：203．￡ 189．1982；FI．Anhui h 153．C
l 5】．1985；FI．Zh~iang1：261．￡1—263．1993：F】．Jiangxi1：283．￡288．1993．
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2期 陆树刚： 中国鳞毛蕨属泡鳞亚属 的分类研究 1 13
Yunnan
， Sichuan，Gu Jzhou,Hunan，Hubei，Jiangx J，Zhejiang，Fujian and Anhu J；Japan．Type from
Japa n．
14．平行鳞毛藤
Dryopterls indusiata(Makino)Makino et Yamamoto cx Yamamoto Supp]．Ic．P1．Formos．5：3．1932；
Tagawa．Co1．Ilustr．Jap．Pterid．104．209．PL 39．￡218．1959；S．G．Lu in Acta Phytotax．S Jn．28(6)：474．
1990；Nakaike New FI．Jap．Pterid．425．￡425．1992；FI．Jiangxi l：285．￡291．1993；FI．Zhej Jang l：264．￡
1-266 1993 N2ph0d m gymnosorum var．indusiatum M akino in BoL M ag．Tokyo 13：65-1899．
— — Dryo碑ens gy删∞s。rⅡvar-indusiata(Makino)Makino cx Bonaparte in NoL Pterid． 49-1918-
Yunnan，Sichuan，Guizhou，Guangxi，Hunan，Jiangxi，Zhejiang and Fuj Jan；Japan．Type from Japan．
15．华南鳞毛蕨
Dryopteris tenuicula M attew et Christ in Lecomte．Not．Syst．1：5 1．1909；C．Chr．．1nd．Fi]．Supp]．1：40．
19 13；Ching in Bul1．Fan Mem．1nst．Bio1．Bot．8：484．1938；FI．ZhejJang l：26 1—1993 D 叩 e
purpureta Tagawa in Acta Phytotax．Geobot．1：307．1932；Co1．1llustr．Jap．Pterid．104．21Z PI．38．￡216．
1959；C．Chr_lnd．F JL Supp1．3：96．1934；Nakaike，New FI．Jap．Pterid．440．￡ 440 1992 D 叩 e
rubristipes Ch Jng et z-Y-Liu in Bul1．BoL Res-3 t4) 19-t-17—1983 Dryo erlsji shd ns Ch Ju
et Yao in Bul1．Bot．Res．2(2】：62．P1．1． 1．198Z— — D 叩 r ，d Ch Jng in Bul Bot．Res．2(2】：64．PI．
1．￡ 3．1982．— — D ，Pr neglecta Ching et Z．Y．L Ju Jn Bul1．Bot．Res．4(4)：4． 32．1984．
— — Dryopterts subtenuicuta Ching et Ch Ju in Bot．Res． 29．PI． 10．f．3 1987 Dryopferts
subchampioni Ching in Bot．Res．2：14．PI．5．￡4．]987．
Guangdong，Guangxi，Gu Jzhou,S Jchuan，Hunan and Zhej Jang；Japan and Korea．Type from
Guangdong．
1‘ 无柄鳞毛蕨
Dryopteris submarginata Rosenst．in Fedde，Repert．Sp．Nov．13：132．1914；C．Chr． lnd．Fil．SuppL 2：
17．1917；FI．Fujian l：200．1982；FI．Zhejiang l：262 1993 D 叩 em sessitipinna Ch Jng et Chiu in
Bot．Res．2：29．PI．10． 4 1987 一 D Pf nudistipes Ch Jng et Chiu in Bot．Res．2：30．PI．1 1． 1．
1987．
Sichuan，Guizhou，Guangxi，Hunan，J Jangx J，Zhejiang and Fuj Jan．Type from Guizhou．
17．广西鞠毛蕨
Dryopterls guangxiensis S·G．Lu in Guiha Ja 19(1)：27—28一]999 D 叩 parafuscipes c舶 et
Chiu Bot Res．2：4．PI．2l￡1．]987(nonChing etZ．Y．L Ju,1984)．
Guangxi．Type from M t．Daming．
18．落叶鳞毛蕨
Dryopteris tenuipes(RosensL)Serizawa in Journ．Jap．Bot．46：19．1971：FI．Talwan l：329,1994．
— — D ，。 ，Pr括erythrosoravar．tenuipesRosenst．inHedwigia 56：341．1915；Hayata
． 1c．PI．Formos．8：147．
1918；H．1toinBoL M ag．Tokyo 50．69．1936．
Taiwan．Type from Ta Jbei．
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l】4 广 西 植 物 19卷
l9．三角鲷毛蕨
Dryopteris subtriangularls(Hope~C．Chr．，Ind．Fil．296．1906；Chingin Bul1．FanMem．[nst．BioLBut．
8：485．1938；Tard．一Blot．et C．Chr．in Lecomte
， F1．Gen．Indo-Chine 7(2)：315．P1．35．￡卜2．1940；Ching
et Y．X．Ling in Acta Phytotax．Sin．22(3)：202．1984；Frmser-Jenkins in Bul1．Br．Mus．Nat．Hist．But．18
(5)：430．C 53．1989．— — Ⅳ印 r^0dj subtriangulare Hope in Journ，But．Loud．28：327．1890．— — LⅡ 口
fitix—mas var．subtriangutaris(Hope)Bedd．，SuppL Fems Brit．[nd．56．1892．— — D 叩 r subassamensis
Ching in Bul1．Fan M em．[nsL Bio1．Bot．＆ 451．1938．— — Dryopterts uropinna PriceinGard．Bul1．Sing．
30：244． ．1．1977；C．M．KuoinTaiwania 30：31．1985(exc1．Syr1．)．— — Dryop№r{s erythrosora auct．．Wu，
Wang et Pong in Bull DepL Bio1．Sun Yatsen Univ．3：38． 10．1932 (non O Kuutze．t89t),
— — D 叩 ， acutiden Ching in Acta Phy[0tax．Sin．8：141．P1．21．￡ 24．1959；F1．Hainan l：144．1964
(non C．Chr．，1905)．— — Dr op№rts caudifronsChing in Acta Phy~otax．Sin．敏 148．P1．22．t．25．1959 FI．
Hainan 1：】44．1964．
Yunnan，G uizhou，Sichuan，Xizang~Guangxi，Hainan． Taiwan；Vietnam，Philippense，Thailand，Burma
and India．Type from India．
20．阿萨姆鳞毛蕨
Dryopteris assamens~s(Hope1 C．Chr．et Ching in Bol1．Dept．BioL Col1．Sci．Sun Yatsen Univ．6：4_
1933；Tard．一BIoL et C．Chr．in Lecomte,FI．Geu．Indo--(2hine 7(2)：316．1941；Fraser-Jenkins in Bul1．Br．
Mus．Nat．Hist．BOL 18(5)：428．L 52．1989．— — Ⅳ印hrodium assamen~s Hope in Journ．BOL Lond．28：
326．1890．— — n —州 vaL assamensis(Hope)Bedd．，Supp1．Fems Brit．[nd．57．1892．
— — D，yop r 矾 n搬 suhsp．a$$~／rte／lsi~ op巴)C．Chr．，[nd．FiL 266．1906．
Yunnan，Guizhou,Guangxi
， Guangdong and Hongkong；Vietnam,Sikkim and India．Type from
India(Assam)．
21．羽裂鳞毛蕨
DryopterisintegrflobaC．Chr．inBul1．Dept．SunYatsen un Jv, 5．1933：Ind．Fil．Supp1．3：87 1934；Ch—
ing in Bul1．Fan M ere．InsL Bio1．BOL 8：479．1938：F1．Hainan．h 145．1964；C．Chr．et Tard．一Blot．in
Lecom te
， FI．Gen．Indo-Chine 7(2)：3 14．1941．- D 。PfPr htrtosparsa aucL：WU,Wang et Pong in
Bul1．DepL Bio1．Sun Yatsen Univ．3．40．P1．II．1932(non Christ,19o8 — — D 叩fP subintegitoba
Serizawa in Journ．Jap．BOL 46：17．￡ 1 1971：F1．Taiwan l：328．1994．
Yunnan，Guangxi，Guangdong，Hainan and Taiwan；Vietnam,Type from Guangdong(Dinhua)．
22．大明鳞毛蕨
Dryopteris tahmingem~s Ching in Bul1．Fan MerrL Inst．Bio1．鲁0t．8：480．1938；Pie．Ser．
． [rid．Fil．Supp1．
4：128．1965．— — D fP hwangiChingin F1．FlIj nl：200．600．1982．
Yunnan
， Guangxi，Guangdong and Fujmn．Type from Guangxi(ML Daming)．
23．齿头鳞毛蕨
Dryoi~ ris labordei fChrist)C．Chr．Ind．Fil．273．1906；Ching in Bul1．Fan Men [nst．Bio1．But．s：488．
1938；【。Corm．Sin．1：240．1972(exc1．descr．et￡480)；FL Fujian l：203．1982 rexc[．descr．et 189)；F1．
—
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2期 陆树剐： 中国鳞毛蕨属泡鳞亚属的分类研究 1 15
Anhui l：153．1985(exc]．descr．e【 15 1)；S．G．Lu in Acta Phytotax．Sin．28(6)：474．1990；F1．Zhejiang 1：
261．1993(exc]．descr．et 卜263)；F1．Jiangxi l：283．】993(exct．desc~et￡288)．— — “ labordei
Christ in Bul1．Soc．Bot．Franc~52：M em．1：40．】905．
Yunnan，Guizhou，Sichuan，Guangxi，Guangdon~ Hunan，Hubei
， Anhu~Jiangxi，Zhejiang，Fuj／an
and Taiwan；Japan．Type from Guizhou．
24．假稀羽鳞毛蕨
Dryopte~s pseudosparsa Ching in Bul1．Fan M em．Inst．Bio1．Bot．8：489．1938；Pic．Ser．．Ind．Fil．Supp1．
4：124．I965；S．G．Lu in Acta Phytotax．Sin．28(6)：475．1990．— — D 叩f cuneatiloba Ching et Chiu in
Bot．Res．2：】1．P1．4． 4．1987．— — D 叩 f盯如 scabr 盯 ChinginBor．Res．2：9．P1．4．￡ 2．1987．
Yunnan
，
Sichuan，Guizhou and Guangxi．Type from Yunnan．
组3．变异鳞毛蕨组
S—ect．3．Vafiae Fraser-Jenkins in Bul1．Br．Mus．Nat．Hist．Bot．14(3)：196．1986．
Type：Dryopteris varia(L_)O．Kuntze．
This section contains 12 speci~ in China．
25．藩鳞鳞毛蕨
Dryopteris sordidipes Tagawa in Acta Phytotax．Geobot．3：29．1934；Co1．Ilustr．Jap．Pterid．101．213．
PI-37． 208．1959；Naka Jke．New Ft．Jap．Pterid．449．￡449．1992；FI．Taiwan 1：327．1994．
Taiwan；Japa n．Type from Japan．
26．寻乌鳞毛蕨
Dryopteris xungwuens~Ching et Shing ex Shing et J．￡Cheng in Journ．Sci．J~ngxi 8(3)：48．1990；F1．
Jiangxi 1：275．￡278．1993．
Jiangxi and Guangxi．Type from Jiangxi(Xungwu)．
27．德化鳞毛蕨
Dryopteris dehuaensis Ching in F1．Fujian 1：209．601．￡197．1982；F1．Zhejiang 1：265．￡1—267．1993．
— — Dry。pterts gushanica Ching et Shing in F1．Fujian 1：209．601．1982．— — D 。口ler neosordidipes
Ching ex Shing et J． Cheng in Journ．Sci．Jiangxi 8(3)：48．1990；FI．Jiangxi 1：275．￡277．1993．
Fujian，Zhejiang and Jiangxi．Type from Fujian(Dehua)．
28．黑水鳞毛蕨
Dryopteris amurensis Christ in Bul1．Geogr．BoL M ans．1909：．35．1909；C．Chr．
． Ind．FiL Supp1．1：29．
1913；Ching in Bul1．Fan Mem．Inst．Bio1．BoL 8：494．1938：Tagawa．Co1．I[1ustr．Jap．Pterid．204．PI．35．￡
】98．】959；Nakaike．New F1．Jap．Pterid．400． 400a．400h 4000 1992；Ft．Herb．NJChina1：61．￡48．I958．
Liaoning，Jilin，Heilongjiang and Neimongol；Japan,Korea and U．S．S．R．Type from U．S．S．R．
(Saghalin)．
29．台湾鳞毛蕨
Dryopteris formosana(Christ)C．Chr．Ind．Fil．266．1906；Ind．Fil．Supp1．3：86．1934；Hayata，MaL FI．
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l16 广 西 植 物 9卷
Formos．416．19】1；Tagawa，Co]．Iflustr．Jap．Pterid．104．2138．PI．38．t 215．1959；Nakaike，New F1．Jap．
Pterid．415．￡415a．415b．1992；FI．Zhejiang l：265．￡1 8．1993：FI．Taiwan l：322．PI．】33．1994．
— — ㈣ fo mo阳n“，l Christ in Bul1．Herb,Boiss．II．4：615．1904；Kawakami，List．P1．Formos．152．
1910． — — Dryo*er phaeolepis Hayata Ic． P1． Formos． I^ 169．t． 、09． 1914． — — Po stichum
constanlissimum Hayata Ic．P1．Formos． ．^]91．t．129—1914 Dryopter constantissima tHayata)
Hayata，Ic．P1．Formos．4：19】 1914 D teris kodamai·Hayata
， Ic．PI．Formos．4．I56．￡97．19】4．
— — PD fc^Ⅲ varium vat-．eurylepidotum Rosen．st．in Hedwigia 56：340．1915；Hayata．Ic．PI．Formos．8：
156—1918 D 印fPr liuiChinginBul1．Bot．Res．4(4)：12． 41．1984．
Taiwan，Sichuan and Zhejiang；Japan．Type from Taiwan．
30．永德鳞毛蕨
Dryopterls yongdeens~W ．M ．Chu ex S．G．Lu in Acta Phytotax．Sin．28(6)：473．1990
Yunnan(Yongd Type from Yongde．
Endem
3l_太平鳞毛蕨
Dryopteris pacifca(Nakai)Tagawa，Co1．Illustr．Jap．Pterid．100．21】．PI．36．￡204．1959；F1．Jiangsu h
69．￡104．1977；F1．Fujianl：202．￡187．1982；FI．Anhuil：152． 150．1985；1watsuki,Fernsand．FernAllies
of Jap．197．PI．124．￡4．1992；FI．Zhejian8 1：266．1993：FI．Jiangxi l：287．￡294．1993．— — 。“ fc “ l
pacifcum Nakai in BOL Mag．Tokyo 39：1 19—1925 PD，y “^m hototepis vat．hikonense H．Ito in
Journ．Jap．Bot．10：45 I旷1934 ) teris varia vat．hikonensfs(H．Ito)Kurata in Namegata et Kurata，
Enum．Jap．Pterid．304．341．1961；Nakaike，New FI．Jap．Ptefid．457．￡457a．457b．】992．— — D 叩 rPr
n由t嚷em Ching et Chiu in Bot．Res． 27．P1．10．t．1．1987(err．D t搋t ngen ．— — Dry。ple s
yushanens~sChing etChiuinBot．Res．k 28．PI．10．t．2一]987 Dryopleris pudouens ChinginBullBot．
Res．3(3)：1 1．￡9．1982．— — D 叩fPr quadrifda Ching ex Shing et C．F．Cheng in Joum．Sci Jiangxi 8
(3)：49．19913；F1．Jiangxi l：290．￡298．1993．
Zhejing,Jiangsu，Anhui，Jiangxi，Fujian and Hunan；Japan and Korea．Type from Japan．
32．假异鳞毛蕨
Dryopteris immixta Ching in F1．Tsin1．2：165．225．P1．1 1．￡ 1-2．1974；FI．Jiangsu l：68．￡ 102,]977：FI．
Fujian l：203．￡】88．1982；FI．Anhui l：149．￡144．1985：FI．Zhejiang l：268．￡1_270．1993；F1．Jiangxi l：
288． 295．1993．
Yunnan．Sichuan，Guizhou,Gansu，Shannxi，Hubei，Hunan
， Jiangsu，Zhejiang，Fujian and JiangxL
Typefrom Gansu．
． 变异鳞毛蕨
Dryopteris varia(L-)0·Kuntze,Rev．Gen-PI．2：8 14-189]；C．Chr．Ind．Fil．Supp1．3=10】．1934；Ching
in Bull-Fan Mem-InsL Biol-Bot．8：490．1938(pro parte)；Tagawa，Co1．1lustr．Jap．Pterid．100．214．P1．36．
￡203-1959；FI-Jiangsu l：67·c 100．1977；FI-Fujian l：201．￡186．1982；FI．Anhui l：】5 1．￡】47．1985；
Nakaike,New FI-Jap_Pterid．456-￡450．1992；1watsuki
， Ferns and Fern Alies of Jap．196．Pl_123． 3．
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2期 陆树刚： 中国鳞毛蕨属泡鳞亚属的分类研究 117
1992；FI．Zhejiang l：266．1993：FI．Jiangxi 1：286．￡ 293．1993；FI．Taiwan 1：329-PI-134-1994·
— — PD 。 “州 varium L，Sp．PI．2：1090．1753．— — 枷“州 varium (L-)Sw．in Schrad．，Journ．BOL
1800—2：35．1801：Hook．．SIx Fil．4：80．￡226 1862．— — PD stichum~arium (L_)Presl，Epim．Bot．57．
1851．— — 且” 口opaca Hook．in Journ．Bot．9：339．1857．— — “m opacum (Hook．)Benth_F1．
Hongk．456 1861 D 叩 f m yabei Hayata，M aL Formos．424．191 1；I。 P1．Formos-4：187-￡ 125-
1914．— — D “^坍 holoepis Hayata，lc．P1．Forrnos．5：332． 1915．— — D fPr mats~izoana Koidz．
In Bot．M ag．Tokyo 39：15．1925．— — D 叩 f ogawai H．1to in Nakai，lc．PI．As．Or．1：18．P1．9．1935-
— — D 叩 r caudifblia Ching et Chiu in Bot．Res．2：21．P1．8．￡2．1987；FI．Zhejiang 1：266-1993-
— — D， 叩fer consimitis Ching in Bot．Res．2：25．Pl_9．￡3．1987．— — D 印 fuyangensis Ching et
Chiu in Bot．Res．2：26．P1．9．￡4．1987；FI．Zhejiang l：268．1993．— — D 印f gtabrescensChing et Chiu
ex Shing et J．￡Cheng in Journ．Jiangxi Sci．8(3)：48．1990；FI．Jiangxi 1：274．￡276 1993 D 叩fⅢ
[ingi in BoL Res．2：23．P1．8．￡4．1987；F1．Zhejiang 1：267．￡1-269．1993．— — D 叩r m nanctmanensis
Ching etZ．Y．LiuinBut1．not．Res．4(4)：9．￡37．1984[101 3(4)：20．L18．1983，n∞4(3)：19．￡21．1984]．
YuBBaB，Sichuan，Guizhou，Guangxi,GUBBgdong，Hunan,Hubei，Henan，liangxi，FuiJan，Zhejiang,
Jiangsu，Anhui and Taiwan；Japan,Korea，Vietnam,Philippines and India．Type from Guangdong．
． 虎耳鳞毛蘸
Dryopteris saxifraga H．Ito．in BOL M ag．Tokyo 50：125．1936；Tagawa．Co1．1llustr．Jap．Pterid．101．
213．P1．37．￡ 206．1959：1watsuki．Ferns and Fern Allies of Japan．198．P1．125．￡ 1-2．1992；FI．Herb．N．
China 1：58．1958．— — D 印f ， varia var．saxifraga(H．1to)H．Ohba in Sci．Rep．Tohoku Univ．(B)36：
1I1．1971；Nakaike，New F1．Jap．Pterid．459．￡459 1992 D 印teris varia subsp．sax~fraga(H．1to)
Sugimoto．Keys Herb．PI．Jap．Pterid．282．405．1966．
Liaoning and Jilin；Japa n and Korea．Type from Japa n．
35．两色鳞毛蕨
Dryopterls setosa(Thunb． Akasawa in Bul Kochi worn．Univ．’：27．1959．— — Po}ypod m setosum
Thunb．，FI．Jap．337．1784．— 印 setos~m(Thunb．)Sw．in Schrad．，Journ．not．1800—2：39．1801．
— — D 印f r varia subsp． setosa (Thunb．)Sugimoto，Key Herb． PI．Jap．Pterid．405． 1966．
— — D 印fPr varia var．setosa(Thunb．)Ohwi，FI．Jap．Pterid．88．1957；Nakaike，New F1．Jap．Pterid．
460． t． 400a． 400b． 1992． — — N印 hrodfum bissetianum Baker in Journ． BOL 1877： 366． 1877．
— — D 印fP bissetiana(Bak．)C．Chr．，lnd．Fil．245．1906；Tagawa，Co1．1lustr．Jap．Pterid．100．205．PI．
36．L 205．1959；lc．Corm．Sin．1：241．￡481．1972；FI．Tsinling 2：165．1974；FI．Jiangsu 1：69．C 103．1977；
F】_Anhui 1：150． 146．1985：1watsuki．Ferns and Fern Allies of Japan 197．PI．123． 4-5．1992；FI．
Zhejiang 1：270．L卜272．1993；FI．Jiangxi 1：288． 296．1993．— — 尸。 ml舯bissetianum(Bak．)Nakai
in not．M ag．Tokyo 45：102．1931．— — Dryo p{efb thunbergii Koidz,In not．M ag．Tokyo 38：106．1924．
— — DTyo p{er shanghaiensis Ching et Chiu in BoI．Res． 24．PL 9．t．1 1987 Dr opter paravaria
Ching et Chiu in Bot．Res．2：22,P1．8．￡3．1987．— — D 印 fP sinobi．~setiana Ching et Z．Y．Liu in Bul
Bot．Res．4 ：8．t．36．1984．— — Dryopt州 s pⅢ 曲i3sett。M Ching ex Shing et J．t．Chengin JiangxiSci．8
(3)49．1990；F1．Jiangxi 1：290．￡297．1993．
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l18 广 西 植 物 9卷
Yunnan，Sichuan，Guizhou
, Hunan，Hubel,Shandong,Shanxi，Henan，Shannxi，Jiangxi，Anhui，Fujian，
Zh~iang and Jiangsu；Japan and Korea．Type from Japan．
36．棕边鳞毛蕨
Dryopteds SaCTOSaBC~I Koidz．in Bot．M ag．Tokyo 38：】O8．1924；Tagawa．Co1．IIlnstr．Jap．Pterid．】01．
212．PI．36．￡ 207．】959；1watsuki．FernsandFernAllies ofJapan197．PI．124．￡2-3．1992；FI．Jiangsu1：68．
￡】01．1977；FL Anhui 1：152．￡149．1985．— — P0lystichum sacrosanctum(KoidL)Koidz．In Bopt．Mag．
Tokyo 43：388．1929．— — Dr op托r varia 敲．sacrosancta(KoidL)Ohwi FI．Jap．Pterid．88．1957；
Nakaik~．New FI．Jap．Pterid．458．E．458．1992．— — P。 {c 竹1 bissetianum vat．sacrosanctum(Koidz 、 i
Nakai in BoL Mag．Tokyo 45=103．193 1．— — D 印fP bissetiana vat sacrosancta(KoidL)H．Ito in Bot．
Mag．Tokyo 5 o：36．1936．— — D 叩fP varia subsp．sacrosancta(KoidL)Sugimoto，Keys Herb．F1．Jap．
Pterick 281．405．1966．— — DrJ 叩 fP tieanzuensisChing etChiuinBot．ReS．2：24．PI．9． 2．1987．
Shandon~Liaoning and Zhejiang；Japan and Korea．Type from Japan．
本文在研究过程中，得到中国科学院 中国植物志编辑委员会的支持，得到中国科学院植物
研究所标本馆的帮助；杭州大学张朝芳教授借阅部分研究标本 在此一并致谢。
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