Abstract:Following a sequence of field research and observation for many years, the above and underground stratification of the seasonal rainforest is hereby discussed. The result shows that there are seven above-ground strata and also seven underground strata corresponding respectively to the environmental gradient of climate and soil in the seasonal rain-forest. Different ecophysiological features, above-ground canopies, and the morphology of underground root systems are formed because of the effects of environmental gradient. The young shrubs and herbs (including epiphytes) exist in the layer where they are poorly illuminated, but with Little change of tempereture and humidity. Above the young tree and shrub layer is a layer of environmental gradieni with strong sunshine, and variable temperature and humidity. Rapid flow and energy transfor as well as circulation and storage of substances occur in the soil at 0–100 cm in depth where is an active bio-activity functional absorption of the fine roots. Under the above-mentioned layer, there is less exchange of energy and substances because of the function of the layer roots is mainly to fix the plant. Measurement of carbon dioxide concentration soil indicate that rainy season is the best season for the flow of substance and energy transfer and for the growth and development of rain-forest during the year. The highest growth speed of rainforest occurs in August and the lowest in January. In addition, in accordance with the structural features of rain-forest, some noteworthy problems on the reconstruction of artificial economic forest after deforestation of the seasonal rain-forest were discussed.