全 文 ：热带亚热带植物学报 2004，12(6)：541—55 1
Journal of Tropical and Subtropical Botany
广东石门台自然保护区藓类植物初报
何祖霞 ，张 力 ，谢国忠2，黄林生2，李剑雄2
(1．中国科学院华南植物园，广东广州510650；2．广东石门台自然保护区管理局，广东英德513000)
摘要：石门台自然保护区位于广东省中北部，面积82 260 hm ，是全省面积最大的保护区。根据我们过去三年的初步调
查和研究，石门台保护区共有藓类植物33科89属 158种(含 1亚种3变种)，其中广东新记录属3属，分别是Eccremidium、
Phylodon和Stereodontopsis；另外，Eccremidium brisbanicum等23种为广东新记录种。
关键词：藓类植物；名录；新记录；石门台自然保护区；广东
中图分类号：Q949．35 文献标识码：A 文章编号：1005-3395(2004)06—0541—1 1
A Preliminary List of M osses from Shimentai
Nature Reserve，Guangdong
HE Zu．xia ，ZHANG Li卜，XIE Guo．zhong ，HUANG Lin-sheng ，LI J ian．xiong
(1．South China Botanical G~dem,the Chinese Academy of Sciences，Guangzhou 510650，China；
2．A dministrative Bureau of Shimentai Nature Reserve。Yingde 5 1 3000，China)
Abstract：Shimentai Nature Reserve IS situated in the center of north Guangdong，24。l 7’49一24。3 l’02”N，1 1 3。
01’1 1”一1 13。46’22’IE．It occupies about 82 260 hectares．and is the largest nature reserve in Guangdong．According
to our field surveys and studies in the past three years， 1 58 species of mosses in 89 genera and 33 families are
recognized from Shimentai Nature Reserve．Among them,3 genera(Eccremidium,Phylodon,and Stereodontopsis)
and 23 species are newly recorded for Guangdong．
Key words：Mosses；List；New records；Shimentai Nature Reserve；Guan gdong
Shimentai Nature Reserve(abbreviated hereafter
as Shimentai) lies in the center of north Guangdong，
administratively belongs to Yingde，Qingyuan．It locates
just outside the north border of Tropic of Cancer(23。
27q~1)，24。17’49”-24。3l’O2”N，ll3。Ol’ll”-1 13~46’22”E，
with a land area of 82 260 hectares． It is the largest
nature reserve in Guangdong,Geographically,Shimentai
borders Hengshishui on the east，Boluo on the west，
and is adjacent to Shigutang,Hengshitang,Yunling and
Dazhen on the south，and Wongyuan，Qujiang and
Ruyuan onthenorth(Fig．1、．
Shimentai is generaly rugged topographically．
Mountainous landscape occupies about 83．6％ of the
tOtal land area,and the average altitude ofmountainous
Received：2003—-10—-09 Accepted：2003——10—-28
Corresponding author
regions ranges 800—1 400 m．The basic rocks generally
compose of granite，san dy shale，and limestone．The
northern part of Shimentai is higher than the southern．
The highest peak is Chuan diding on the noah border
with Qujiang，reaching 1 5 86 m above sea leve1．
Shimentai has a subtropical monsoon climate，with
annual mean temperature 20．9~C，mean annual rainfall
1 882．8 nl／n，and mean annual non—frost time 317 days．
The major vegetation types are monsoon valey
evergreen broadleaved forest， montane evergreen
broadleaved forest，subtropical evergreen coniferous—
broadleaved forest． Th e soil types are latosolic red
earth(below 300 m al1．)，montane red soil(300-600 m
alt．)and montane yelow soil(above 600 m alt．)[1-31．
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Fig．1 Map ofShimentai Nature Reserve，Guangdong．Numbers show the localities ofsites surveyed
． 1 Genggu，2 Huawu，
3 Ludong，4 Zhongkong，5 Huangdong，6 Jinzikeng，7 Lianshan，8 Chuand／ding,9 Laowuchang,10 Shuitou，
1 1 Huangdong Waterpower Station，12 Shakou，13 Chaitoushan，14 Huashuishan，15 Daxishan．
Sporadic collections and studies have been
carried out in Shimentai previously． Ⅵ checked the
bryophyte colections in the Herbarium of South
China Institute of Botany， the Chinese Academy of
Sciences 0BSC)，and found that the first colection WaS
made by Mr．Liang Bao—han in 1 943．Three specimens
were collected．Afterward,Prof．Xu Xiang—hao collected
31 bags ofmosses in 1956． and Mr ． Chen Xue—xian
collected 3 in 1 957．At the same year．South China
Institute of Botany organ ized an expedition team
(msc Geo．Dept．)for phytogeographical survey，3 l
bags of bryophyte specimens were collected．All these
specimens were colected from Huashuishan．Some of
these collections were studied and reported in various
references t4-~．Recently,Forestry College ofSouth China
Agricultural University and Biology Department of
Guangzhou University reported 65 species of mosses
in 38 genera and 23 families from Shimentai in an
unpublished paper”， un fortunately they cited no any
specimens．Obviously,the knowledge on the diversity
of Shimentai bryophytes was far to complete．
1 Materials and methods
Two field surveys have been carried out in
November．200 l and March，2003． In total， 532
bags of bryophytes were collected． Based on the
microscopical identifcation of our collections，the
previous collections deposited in IBSC，and consulting
relevant references,we recognized 1 58 species
(including 1 subspecies and 3 varieties) in 89 genera
and 33 fam ilies of mosses occurred in Shimentai．
Among them，three genera(Eccremidium,Phylodon，
and Stereodontops is) and 23 species (Barbela
flagelifera，Bryum paradoxum,B．pseudotriquetrum,
Campylopus ，e taiwanensis，Dicranodontium
didictyon, Duthiella speciosissima, Eccremidium
brisbanicum,Ectropothecium obtusulu~Homalothecium
laevisetum，Isopterygium minutirameum，Leucobryum
aduncum vat．8co~are，Macromitrium angustifolium，
Phyllodon lingulatus， Plagiothecium eurphyllum，
Polytrichastrum alpinum，Ptychomitrium dentatum，
obryum latifolium,Rhynchostegium contractum，
1)ForestryColege ofSouthChinaAgriculturalUniversity，BiologyDepartmentofGuangzhouUniversity．Alist ofhigherplantsfrom Shimentai
Nature Reserve，Guangdong-In：Pang X F，Zhang J Q．Colected papers for investigation in Shimentai Nature Reserve，Guangdong
． The Administative
Bureau ofShimentai Nature Reserve，Guan gdong，2001，229—293
． (In Chinese)
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第6期 何祖霞等：广东石门台自然保护区藓类植物初报 543
SVhognum cuspidatum，S．ovattm,Stereodontopsis
pseudorevoluta,and Vesicularia fer／ei)were new
additions to Guangdong．
The liverwort flora of Shimentai will be reported
in a separate paper．
2 A preliminary list of mosses from
Shimentai Nature Reserve
(1)In this list,the family arangement folows Chen ．
Genera and species are arranged alphabetically．
The Latin binomials generally were checked by
consulting Redfeam et a1． m．in addition to update
references．
(2)For saving space， specimen colectors are
abbreviated as Z for Zhang Li and ZH for Zhang
Li& He Zu．xia． Other collectors were not
shortened．
(3)Al specimens examined are deposited in
Herbarium of South China Insititute of Botany，
the Chinese Academy of Sciences(msc)．
(4)The species and genera new to Guangdong are
indicated by an asterisk( )and double asterisks
(料)，respectively．
Sphagnaceae
1． Sphagnum cuspidatum Enrh． ex Hofm．～
Midway from Lianshan to Jinzikeng，1 000 m，on
soil in sparse forest，Z2785．
2．Sphagnum junghuhnianum Dozy & Molk．～
Zhongkong,510 m，on rock in forest，Z2626．
3． Sphagnum ovatumHamp．- Chuandiding．1 350m,
on rock in sparse forest，Z2840；1 540 m，on rock
in bamboo forest，Z2827．Midway from Jinzikeng
to Huangdong，940 m，on rock in forest，Z2843；
950 l on rock in forest,Z2849．Zhongkong,465 l
on rockin forest，Z2631．
4， Sphagnum palustre L．——Zhongkong，465 m，on
soil in forest，Z2641．
Ditrichaceae
5，Ditrichum dificile(Duby)Fleisch．～ Shigutang．
580 m，on rock in forest，Z2742．
6．Ditrichum heteromalum(Hedw．)Brit．一 Shakou,
36 m，on rock in grassland，ZH3277，
7．Ditrichum palidum (Hedw．)Hamp．一 Midway
from Jinzikengto Chuandiding，1 250 m，on trunk,
Z2800．
8．}}Eccremidium brisbanicilm (Broth．) Stone&
ScoR．—— Huangdong Waterpower Station，408 m，
on soil in forest， ZH3333， ZH3335， ZH3337．
Shuitou， 220 m， on soil by a rice field， ZH3223．
Previously，this genus or species was known for
China only from Hong Kong㈣．Guangdong is the
second locality．
9．Garckea flexuosa(Grif．)Marg．&Nork．一
Huangdong Waterpower Station，270 m，on rock
by a stream ， ZH3258； 300 m， on soil in forest，
ZH3264．
Dieranaeeae
10．Campylopus atrovirens De Not．———Jinzikeng
1400 mon rock by a path， Y．-H． Yan59．
Shigutang,535 IT1，on rock in forest,Z2730；595 n
on rock in forest，Z2744．
1 1．Campylopus ericoides (Grift．、 Jaeg．～
Huashuishan，1 000 m，on trunk，IBSC Geo．Dept．
6100 5】fWith C graeilis)．Midwayfrom Jinzikeng
to Chuandiding，1 160l11onthink,Z2788；1 235l11
on rock in forest,Z2796．
12． Campylopus fragifs(Brid．、B．S．G．一 Midway
from Jinzikeng to Chuandiding，1 240 m，on soft
in grassland，Z2792．
13．Campylopus gracilis(Mit)Jaeg——Huashuishan,
l 000 m，on trunk，mSC Geo．Dept．6100『4 (with
C．ericoides)．
14．Campylopus hemitrichius(C．Muel1．)Jaeg．一
Chaitoushan,1 70 n1．on rock by a stream,ZH3290．
15． Campylopus taiwanens~ Sak．一 Chaitoushan．
170 m，on rock in forest，ZH3296；410 m，on rock
in forest．ZH33O2．
16．Campylopus umbelatus(Am．)Paris-Huangdong
Waterpower Station， 350 m， on rock in forest，
ZH3239．Shuitou，279 m，on concrete in sparse
forest,ZH3179；350m，on soilinforest，ZH3211．
1 7． ranella aus~o．sinensis Herz．& Dix．～
Huangdong Waterpower Station，300 m，on soil by
apath，ZH3232．
18．Dicranela coarctata(C．Muel1．)Bosch&Sande
Lac．——Huangdong Waterpower Station，270 m，
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on rock，ZH3256，ZH3260．Shakou，90 m，on soil
by a path，ZH3289．Shuitou，260 m，on soil by an
agricultural field，ZH3215； 300 m， on soil by a
path，ZH3195．
1 9．DicraneUa varia(Hedw．)Schimp．一 Huangdong
Waterpower Station，270 m，on rock by a stream，
ZH3257．
20．*Dicranodontium didictyon (Mit) Jaeg ——
Jinzikeng,800 m，on thin soil over rock，Y．一H．
Yan56．Midway from Jinzikeng to Chuandiding，
1 335 m，on rock in forest，Z2820，Z2821．
21．Dicranodontium filifoliumBroth．——Huashuishan,
IBSC Geo．Dept．6172t4,~．
22．Dicranodontium papilifofium C． Gao——
Zhongkong，510 m，on rock in forest，Z2622．
23．Dicranodontium uncinatum (Harv．)Jaeg．一
Laowuchang，665 m， on rock in forest， Z267 1，
Z268 1．Midway from Lianshan to Jinzikeng,830 n=L
on rock in forest，Z2777．
24．Dwrano~ma dicarpum (Nees) Paris．—
Chuandiding，1 570 m，on humus in forest，Z2832．
Jinzikeng， 1 000 m， on rock by a path， Y．一H_
Yan60． Midway from Lianshan to Jinzikeng，
1 000 m，on rock in forest，Z2780；1 330 m，o
．
n
rock in forest，Z2817．
25．Dicranoloma fragile Broth．——Midway from
Jinzikeng to Chuandiding， 1 340 m， on rock in
forest， Z2822． Zhongkong， 510 m， on rock in
forest，Z2620．
26．Holomitrium densifolium(Wils． Wijk．&Marg．
— — MidwayfromJinzikengtoChuandiding，1 165 m，
on rock in forest， Z2790； 1 235 m， on rotten
branch，Z2795．
27．Leuco／oma okamurae Broth．———Midway from
Lianshan to Jinzikeng，855 m，on rock in sparse
forest，Z2778．
28．Trematodon longicollis Michx．———Huangdong
Waterpower Station，270 m，on rock by a stream，
ZH3252，ZH3255，ZH3259；408 m，on soil in
forest， ZH3334． Midway from Lianshan to
Jinzikeng，1 020 m，on soil in sparse forest，Z2784．
Leucobryaceae
29． Leucobryum aduncum var．scalare fC．Muel1．
ex Fleisch．)A．Eddy— Midway from Lianshan to
Jinzikeng，785 m，on rock in forest，Z2775．
30．Leucobryum boninense SulL ＆ Lesq． 一
Huangdong Waterpower Station，300 m，on rock
by apath，ZH3233．
3 1．Leucobryum bowringii MitL — Laowuchang．
5lO rI1，on rock in forest，Z2679．Ludong，275 m，
on rock in forest，Z2602．Zhongkong，650 m，on
roten wood in forest．Z2604．
32．Leucobryum chlorophyllosum C． Muel1．———
Huashuishan， 274 m， on rock by a stream，
ZH332 1．Shuitou，279 m，on thin soil over rock in
sparse forest，ZH3 1 80．
33．Leucobryum javense(Brd．)Mit．一 Daxishan，
3 10 m，on branch，Z2703．Jinzikeng，l 000 m，on
rock by a path，Y．一H．Yan45．Laowuchang，665 m，
on trunk， Z2669． Midway from Jinzikeng to
Chuandidin~，l 260 m，on roten branch，Z2799．
34．Leucobryum juniperoideum(Brd． C．Muel1．一
Chaitoushan，400 m，ZH3297．Huawu，525 m
， on
rotten tree base，Z2556．Shuitou，300 m，on soil by
a stream．ZH3l98．
35．Leucobryum scabrum Sande Lac．——Huashuishan,
H．X．Xu ll(2)[4,51．Shuitou，300 m，on rock by
stream，ZH3213．Zhongkong，465 m，on rock in
sparse forest，Z2634．
36．Leucophanes octoblepharioides Bricl—— Ludong，
295 m，on rock in forest，Z2584．
37．Octoblepharum albidum Hedw．——Laowuchang，
345 m，on tree base，Z2677．Ludong，290 m，on
trunk,Z2601．Zhongkong，515 m，onrockin forest，
Z2624．
Fissidentaceae
38．F sidens crispulus Brd．一 Daxishan．395 m．on
rock by a stream in forest。Z2722．
39．F sidens dubius P． Beauv．一 Midway from
Jinzikeng to Huangdong，505 m，on rock in forest，
Z2864．Shuitou，300 m，on rock in forest，32 l O．
Zhongkong，525 m，on rock in forest，Z2614．
40．Fissidens geminiflorusDozy&Mole —Midway
from Jinzikeng to Huangdong，505 m，on rock in
forest，Z2862．Shuitou，283 m，on rock in sparse
forest．ZH3l87．
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546 热 带 亚 热 带 植 物 学 报 第 12卷
290 m，on rock by a stream，ZH3272．
68．Pohlia nutans(Hedw．)Lindb．一 Shuitou，300 m，
on concrete，ZH3 192．
M niaceae
69．Plagiomnium acutum (Lindb．) T．K叩．一
Chaitoushan， 400 m， on thin soil over rock in
forest，ZH3298．Huawu，430 m，on soil in sparse
forest，Z2571．Shakou，36 m，ZH3275．Shuitou，
270 m，on rock in sparse forest，ZH3207．
70．Plagiomnium cuspidatum (Hedw．1 T．Kop．一
Huashuishan，274 m，on soil by a stream，ZH33 19
．
71．Plagiomnium succulentum (Mit．)T．Kop．一
Huawu，435 m，on soil in forest，Z2577．Ludong，
275 m，on rock in forest，Z2593；275 m，on thin
soil over rock in forest， Z2594． Midway from
Jinzikeng to Huangdong，360 m，on rock in forest，
Z2867． Shuitou， 220 m， on rock by a stream，
ZH3217．
72．Plagiomnium vesicatum (Besch．)T．Kop．一
Huangdong Waterpower Station，270 m，on rock
by a stream，ZH3247．Shuitou，280 m，on rock by
a stream．ZH3183．
Leptodontaceae
73．Caduciella guangdongens~ Enroth——Huashui—
sh矶 760 m，on bark，IBSC．Geo．Dept．6070[7J．
Rhizogoniaceae
74． Pyrrhobryum latifolium (Bosch&Sande Lac．)
Mitt．——Midway from Jinzikeng to Chuandiding，
1 260 m，Z2804．Zhongkong，625 m，on rock in
forest，Z2607．
75．Pyrrhobryum spiniforme (Hedw．) Mit．一
Huashuishan,274 n=L on rock by a stream,ZH3323．
Midway from Jinzikeng to Huangdong，905 m，on
rock in forest，Z2851．Midway from Lianshan to
Jinzikeng， 555 m， on rock in forest， Z2755．
Shigutang 545 m， on rock in forest， Z273 8．
Zhongkong，500 m，on rock in forest，Z2629．
Bartramiaceae
76．philonotis lancifolia Mit．——Chaitoushan,400 n=L
on rock，ZH3295．
77,Philonotis thwaitesii Mitt．———Huangdong Water-
power Station，300 m，on rock in forest，ZH323 1．
78．PhUonotis turneriana (Schwaegr．) Mit．一
Daxishan,300 mon rock,Z2697．Jinzikeng,300 m，
on soil by a path，Y．一H．Yan53．
Orthotrichaceae
79． Macromitrium angustifolium Dozy&Molk．一
Huashuishan，IBSC Geo．Dept．6098，IBSC Geo．
Dept．6099．Laowuchang，665 m，on log in forest，
Z2667．
80．Macromitrium ferriei Card．＆Th臼．一 Genggu，
325 m， on tree base in sparse forest， Z2652．
Huashuishan，IBSC Geo．Dept．6 1 80．
81．Macromitrium formosae Card．一 Huashuishan，
500 m，on wet rock，X．一X．Chen 8001 1．
82．Macromitrium gymnostomum Sul1．& Lesq．一
Huawu， 430 m， on log in sparse forest， Z2572．
Midway from Jinzikeng to Chuandiding，1 280 m，
on roten log in forest， Z2806． Midway from
Lianshan to Jinzikeng，985 m，on branch in forest，
Z2781．
83．Macromitrium japonicum Dozy& Molk．一
Huawu，525 m，on trunk，Z2554．Shuitou，295 m，
on trunk ofBischofia po carpa，ZH3203．
84．Macromitrium nepalense (Hook． & Grev．)
Schwaegr．一 Huawu，555 m，on trunk，Z2549．
Midway from Lianshan to Jinzikeng， 5 85 m， on
rotenlogin forest，Z2762．
85．Schlotheimia greviUeana Mit．——Huashuishan，
870 m，on beech，X．-H．Xu l O(4)．Huawu，43 5 m，
on trunk in sparse forest， Z2569． Midway from
Jinzikeng to Huangdong，725 m，on roten log in
forest，Z2855．Shigutang，600 m，on roten log in
forest，Z2746．
86．Schlotheimia pungens Bartr． ———Huashuishan,
IBSC Geo． Dept．6173， IBSC Geo． Dept．6179．
Midway from Jinzikeng to Chuandiding，1 230 m，
on trunk， Z2794； 1 3 10 m， on trunk， Z2808，
Z281 1．
Racopilaceae
87．Racopilum aristatum Mitt——Huashuishan,218n1．
onhumusbyastream,ZH3316．Laowuchang，670m,
on log in forest，Z2670．
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第6期 何祖霞等：广东石门台自然保护区藓类植物初报 547
Trachypodaceae
88．*Duthiella speciosissima Broth． ex Card．——
Midway from Jinzikeng to Chuandiding，1 3 10 m，
on rock in forest，Z2809．
Pterobryaceae
89．Garovaglia elegans(Dozy&Molk．)Hampe ex
Bosch& Sande Lac．一 Midway from Jinzikeng to
Huangdong，600 m，on trunk，Z2857．
M eteoriaceae
90．Aerobryidium levieri(Ren．&Card．)S．H．Lin—
Daxishan， 360 m， on roten branch in forest，
Z27l6．
91．Aerobryopsis parisirCarct)Brotk- Chuandiding
l 400 m，onbranch，Z2836．
92．Aerobryopsis subdivergens (Broth．) Broth．一
Ludong 270 I on trunk,Z2588．Shigutang，595 m，
on branch，Z2743(with Barbela q f r ．
93．Aerobryum speciosum Dozy & Molk —
Chuandiding，l400m，ontrunk，Z2838．
94． Barbe／a flageUifera(Card．)Nog．一 Daxishan，
310 m，on branch，Z2701；325 m，on branch by a
stream in forest，Z2712．Genggu，325 m，on tree
base in sparse forest， Z2651． Midway from
Jinzikeng to Chuandiding l 250札 on branch,
Z2801．Shigutang 610 rnonbranch,Z275l；580I
一
on roten branch，Z2740．Zhongkong,5 1 5 m，on
branch．Z2623．
95．Chrysocladium retrorsum (Mit)Fleisck —一
Huawu,520 m，on rock in forest，Z2543．
96．Meteoriopsis reclinata(C．Muel1．)Fleisch．in
Broth．——Midway from Jinzikeng to Chuandiding，
l 305 m，on trunk，Z2812．
97．Meteorium subpolytrichum (Besch．)Broth．一
Huawu， 425 m， on log in sparse forest， Z2573．
Midway from Jinzikeng to Huan gdong，720 m，on
roten log，Z2854．
98．Neobarbela comes(Gri劬Nog．-Daxishan,360m,
on roten branch,Z2714；375n1onbranch,Z2719．
99．Neodicladiela pendula(Sul1．)Buck．一 Midway
from Jinzikeng to Chuandiding，l 310 m，on trunk，
Z28l0．
1O0．Pseudobarbela atenuata (Thwait．&Mit．、
Nog．——Laowuchan g，455 m，on trunk，Z2696．
Ludong， 280 m， on trunk，Z2581．Shigutang，
535 m，on branch，Z2735．
Neckeraceae
101．Homaliodendron exiguum osch&SandeLac．)
Fleisch．——Daxishan，360 m，on roten branch，
Z27 1 7． Huashuishan， IBSC Geo． Dept．6045．
Laowuchang，670 m，on bran ch，Z2662．
102．Homaliodendron flabelatum (Sm．1 Fleisch．——
Chuandiding，l 3 80 I on trunk,Z2825．Daxishan,
305 m， on rock in forest，Z2700； 310 m， on
branch， Z2705； on rock in forest， Z2706．
Laowuchan g，480 m，on rock in forest，Z2695．
Ludong， 290 m， on rock in forest， Z2583．
Midway from Jinzikeng to Huangdong，945 m，on
roten trunk， Z2848． Midway from Lian shan to
Jinzikeng， 600 m， on rock in forest， Z2766．
Shigutang，535 m，on rock in forest，Z2727．
103．Pinnatela anacamptolepis(C．Muel1．)Broth．
— — Huashuishan，on trunk，X．一H．Xu 23(3)In．
Ludong，280 m，on rock in forest，Z2579．
104．Thamnobryum subseriatum (Mit．ex Sande
Lac．)Tan— Midway from Lianshan to Jinzikeng．
590 m，On rock in forest，Z2769．
Hookedaceae
105．Calicostela prabaktiana(C．Muel1．)Bosch&
Sande Lac．———Midway from Lianshan to Jinzikeng,
610m，on rockin forest，Z2771．
106．Calyptrochaeta japonica(Card．&Ther．)1wats．
& Nog．一 Midway from Lianshan to Jinzikeng．
600 I on rock in forest，Z2770．
107．Distichophyrllum collench) matosum Card．var．
coUenchy’matosum — Shuitou，283 m，on rock
by a stream，ZH3 1 88．Zhongkong，500 m，on rock
in forest，Z2628．
1 08．Distichoph) llum coUench~ matosum Card．var．
pseudosinense Tan&Lin— Laowuchang 355札
on thin soil over rock in forest，Z2655(with
Pallavicinia subciliata)．
109．Distichophyllum maibarae Besch．———Midway
from Jinzikeng to Huan gdong，505 m，on rock in
forest，Z2863；580 m，on rock in forest，Z2859．
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热带 亚 热 带植 物 学 报 第 12卷
Shigutang，630 m，on rock in forest，Z2753．
Hypop~rygiaceae
1 10．Cyathophorum adiantum (Grif．) Mit．
(Cyathophorela subspinosa Chen，C_japonic
Broth．，C．tonkinens~(Broth．&Par．)Broth．) 玛一
Daxishan，310 m，on branch，Z2704；310 m，on
rock in forest，Z2708．Laowuchang，490 m，on
mink，Z2693；635 m，on rock in forest，Z2686．
Ludong，280 m，on rock in forest，Z2578．Midway
from Jinzikeng to nuan~ong，600 ITI，on trunk，
Z2858．Shigutang，610 m，on trunk，Z2747．
Theliaceae
1 1 1．Fauriel& tenerrima Broth． ———Midway from
Lianshan to Jinzikeng，575 m，on roten log in
forest，Z2673；455 m，on branch，Z2761．
Leskeaceae
112。Pseudoleskeopsis zippeli(Dozy&MolE)Broth．
— — Chaitoushan, 480n=L on rock by a stream,
ZH3308．Midway from Lianshan to Jinzikeng，
600 m，on rock in forest，Z2772．Shuitou，220 m，
on rock by a stream，ZH3220．
Anomodontaceae
113．Haplohymenium pseudo-triste(C．Muel1．)Broth．
— — Huawu，495 m，on trunk，Z2561．
1 14．Herpetineuron toccoae(Sul1．&Lesq．)Card．一
Huashuishan，X．一H．Xu 1 1(1)．Shuitou，280 m，
on rock in sparse forest，ZH3185．Huangdong
W aterpower Station，300 m，on roten log by a
stream ．ZH3238．
Thllidiaceae
115．Claopodium aciculum(Broth．)Broth．一 Huawu,
490 m，on soil in forest，Z2565．Shuitou，220 m，
on rockbya stream ．ZH3218．
116．Claopodium assurgens(Sul1．&Lesq．)Card．一
Huashuishan,200 m on rock,X一14_Xu 1 9．Shuitou,
300 m，on rock in sparse forest，ZH3193．
1 17．Haplocladium angustifolium(namp．&C．MuelL)
Broth．。_——Huangdong Waterpower Station,300 rn,
on rock，ZH3237．Shakou，36 m．on rock in
grassland，ZH3274；90 m，on rock by farmland，
ZH3285．
118．Haplocladium microphyHum(Hedw．1 Broth．一
Chaitoushan,410 rn,onrock,ZH3301．Huangdong
Waterpower Station,270 n=L on concrete,ZH3245；
350 m，on rock，ZH3243．Shakou，36 m，on rock
in grassland，ZH3276，ZH3280．Shuitou，300 m，
on soil by a stream．ZH3194．
119．Thuidium assimile(Mit)Jaeg— Huashuishan，
IBSC Geo．Dept6177a；X—H Xu9,x—I-[Xu 210)．
120．Thuidium glaucinoides Broth．——— Daxishan,
305 n1，on branch,Z2699；3 l 0 n1，on branch,Z2702；
375 Il1，on rotten log in forest，Z2726．Huangdong
Waterpower Station，270 m，on rock by a stream，
ZH3246； 408 m， on rock in forest， ZH3326．
Huashuishan ，274 m，on soil in forest，ZH3322．
Laowuchang，610 m，on rock in forest，Z2688．
Ludong．295n=L on rockin foresg Z2585．Midway
from Jinzikeng to Huangdong，840 m，on rock in
forest，Z2853．Shuitou，220 m，on rock by a
stream ，ZH3219．Zhongkong，615 m，on rock in
forest．Z2608．
121．Thuidium pristocalyx (C．Muel1．)Jaeg．一
Huangdong Waterpower Station，388 m，on roten
log in forest，ZH3324；388 m，on soil in forest，
ZH3325．Huashuishan,IBSC Geo．Dept6 1 74；x-H
Xu25(a)；274 m，on rock by a stream ，ZH3317．
Huawu，435 m，on tree base in forest，Z2570．
Jinzikeng,1 000m on soilbyapath,Y．一14_YaIl58．
Laowuchang， 665 m， on tree base in forest，
Z2663．Midway from Lianshan to Jinzikeng,445 m
onbranch，7_2759．
Brach3 theciaceae
122．Eurhynchium savatieri Schimp．ex Besch．———
nuan~ongWaterpowerStation,300m,onconcrete
bya stream ．ZH3265．
123． 丑_omalothecium laevisetum Sande Lac．一
Genggu，365 m，on crevice，Z2649．
124．Platyhypnidium riparioides (Hedw．)Dix．一
Chaitoushan,4 1 0 m,on rock,ZH3299．Huashuishan,
266 m，on rock by a stream ，ZH3314．
125．*Rhynchostegium contractum Card．- Huashui—
shan，150 m，on soil by a path，ZH3312．
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第6期 何祖霞等：广东石门台自然保护区藓类植物初报 549
Entodontaceae
l26．Entodon macropodus(Hedw． C．Muel1．一
Genggu， 295 m， on rock in forest， Z2642．
Huangdong Waterpower Station，290 m，on rock
by a stream。ZH3266．Huashuishan，388 m，on
rotten log in forest，ZH33 1 5．Huawu，495 m，on
roten log in forest。Z2560．Shuitou。280 m。on
concrete by a stream in sparse forest． ZH3 l 84．
Zhongkong，500 m，on rock in forest，Z2627．
127．Entodon viridulus Card．——Shuitou。300 m。on
rock in sparse forest．ZH3201．
1 28．Mesonodon flavescens(Hook．)Buck— Genggu,
3l0m。ontreebaseinforest．Z2645．
Plagiotheciaceae
129． Plagiothecium eurphylum (Card．&Ther．、
1wats．——Midway from Jinzikeng to Chuandiding，
l 255 m，on rock in forest，Z2802．Midway from
Jinzikeng to Huangdong，6 1 5 m，on thin soil over
rock in forest，Z2856．Midway from Lianshan to
Jinzikeng，8 l 5 m，on roten branch，Z2776．
130．Pseudotaxiphylum pohliaecarpum (Sul1． &
Lesq．)1wats．——Chaitoushan，1 70 m，on rock，
ZH3293．Huangdong Waterpower Station，290 m，
on rock in forest，ZH3248，ZH3270；300 m，on
rock by a path，ZH3225，ZH3236．Huashuishan，
l50 n1’on rock in fores~ZH33 10．Huawu,525 n
on soil in forest，Z255 1．Laowuchang。on rock in
forest，Z2666．Shakou，36 m，on rock，ZH328 l：
90 m．on soil in bamboo fores~ZH3288．Shigutang,
545 m，on rock in forest，Z2736．Shuitou，260 m，
on soil in fores~ZH32l6：290 n on soil in fores~
ZH3l9l：350 m，on soil in forest，ZH32l2．
Sematophylaceae
131．Acroporium secundum (Reinw．& Homsch．、
Fleisch．——Shigutang,625 n1．on branch by a stream
in forest，Z2752．This species has been reported
for Guangdong once by Jia et aL(2001)．Unfor—
tunately,they did not include it in their bryophyte
list【 ． Its occurrence in Guangdong is now
verifled．
132．Brotherela falcatata(Dozy&Molk．1 Fleisch．
- Chuandiding, 1 550 n1．on rock in bamboo
fores~Z2829．
133．Brotherela fauriei(Card．)Buck——Chuandiding,
l 360 onrotenbranchinfores~Z2826；l 540
on rock in bam boo fores~Z2828．Daxishan,305 n1．
on branch， Z2698． Midway from Jinzikeng to
Chuandiding，l 1 65 m，on rock in forest，Z279 1；
l 300m，on rockinforest，Z2807．
134．Brotherela henoni (Duby) Fleisch． ——
Chuandiding，l 565 m，on humus in forest，Z2834．
Midway from Jinzikeng to Chuandiding，l 330 m，
on rock in forest，Z28 1 8．Midway from Jinzikeng
to Huangdong，870 m，on rock in forest，Z2852．
Midway from Lianshan to Jinzikeng，l 020 m，on
soil in sparse forest，Z2783．Shigutang，605，on
branch，Z2750．Zhongkong，470 m，on rock in
forest．Z2636．
135．Gammiela tonkinensis(Broth．&Par．、Tan—
Huashuishan，S．-F．Shul 8(2)阎，IBSC Geo．Dept．
6182；on branch,X．一H．Xul8(2)．Shigutang,600
on log in forest，Z2749．
136．Isocladiela surcularis(Dix．、Tan&Mohamed
— — Daxishan，3 10 m，on roten branch in forest，
Z2707；400 m，on branch，Z2720．
137．Palisadula chrysophyla rCard．)Toy．一 Midway
from Jinzikeng to Chuandiding，l 260 m， on
branch，Z2803．
138．PapiUidiopsis ma~rosticta(Broth．&Par．1 Buck
& Tan— Laowuchang， 410 m， on rock by a
stream in forest，Z2657；465 m，on rock in forest，
Z2694．Shigutang，640 m，on trunk，Z2754．’
139．Sematophylum pulcheUum (Card． Broth．一
Huangdong Waterpower Station，408 m，on roten
log in forest，ZH3327．
140．SematophyUum subhumile(C．MueU．)Fleisch．
一 Chuandiding， l 315 m， on roten in forest，
Z2823； l 450 m， on leaves in bamboo forest，
Z2837．Huawu，550 m，on rock in forest，Z2550．
141．SematophyUum subpinnamm (Brid． Brit．一
Huangdong Waterpower Station，320 m，on log
bya stream，ZH3240．
142．Trichosteleum lutschianum (Broth．& Par．1
Broth．——Ludong， 280 m， on rock in forest
，
Z2586．Shigutang，6 1 0 m，on branch，Z2748
．
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550 热 带 亚 热 带 植 物 学 报 第 l2卷
Hypnaceae
143．Ctenidium capilifolium (Mit．) Broth． 一
Chaitoushan，470 m，on rock，ZH3305．Shuitou，
220 m，on rock by a stream，ZH3222：
144． Ectropothecium obtusulum (Card．、1wats．一
Huangdong Waterpower Station，270 m，on rock
by a stream,ZH3249，ZH3250．Huashuishan,266 m
on rockbya stream，ZH3313．
145．Hondaela caperata(Mit．)Ando，Tan&1wats．
— — Zhongkong，635 m，on trunk，Z2605
．
146．Hypnum plumaeforme Wils．——Chuandiding，
1 370 m，on rock in forest，Z2824．Huangdong
Waterpower Station，300 m，on rock，ZH3228；
320 m，on rock in forest，ZH3262．Laowuchang，
545 m，on rock in forest，Z2660；670 m，on log
in forest， Z2668． Midway from Jinzikeng to
Chuandiding，1 330 m，on rock in forest，Z28 15．
Shuitou，220 m， on soil by farmland， ZH3221；
280 IrL on thin soi1 over rock by a stream, ZH3204．
147．*Isopterygium minutirameum(C．Muel1．)Jaeg．
— — Laowuchang，355 m，on tree base，Z2654．
148． *Phylodon lingulatus(Card．)Buck- Daxishan,
390 m。 on rock by a stream in forest， Z2724．
Previously， this genus and species was known
only from Hong Kong， Taiwan，Xizang， and
Yunnan in China．
149． *Stereodontopsis pseudorevoluta(Reim)Ando
— — Midway from Jinzikeng to Chuandiding，
1 240 m, ontrunk，Z2793．
150．rax／phyUum alternans (Card．、 1wats． 一
Laowuchang，670 m，on rock by a stream in
forest,Z2673．
151．Taxiph)’Uum taxirameum (Mit．、Fleisch．一
Shakou，36 m，on thin soil over rock，ZH3279．
152． Vesicularia ferriei(Card．&Ther．、Broth．一
Chaitoushan ，4 1 0 m，on rock，ZH3303；480 m，on
rock by a stream，ZH3306．Huashuishan，274 m，
on rock by a stream，ZH3320．
Diphysciaceae
’153．Diphyscium fulvifolium Mit．一 Laowuchang，
490 m，on rock in forest，Z2659．
Polytrichaceae
1 54．Atrichum yakushimense(Hofik．、Mizushima—
Chaitoushan，400 m，on rock in forest，ZH3294．
155．Pogonatum cirratum(Sw．、Brid．ssp．fuscatum
(Mit．) Hyv~nen — Huangdong Waterpower
Station，300 m，on rock，ZH3227．Huashuishan，
X．一H．Xu3；230 m，B．一H．Liang84010，B．H．
Liang 84011． Huawu, 490m on soilinforest,
Z2568．Ludong,280 II1，on rock in forest，Z2580；
525 m，on rock by a stream in forest，Z2613
，
Z26 1 6．Shigutang，535 m，on rock by a waterfall，
Z2734．Shuitou，280 m，on rock by a stream，
3206．
156．Pogonatum inflexum (Lindb．)Sande Lac．一
Huangdong Waterpower Station，270 m，on rock
by a stream ，ZH3254．Huashuishan，150 m，on
，
soil in forest，ZH3309；450 m，IBSC Geo．Dept．
6048~Shuitou，220 m，0n soil by a path，ZH3208．
Zl~gngkong，465 m，on rock in forest，Z2633．
157． Polytrichastrum alpinum(Hedw．1 G．L．Smith．
一 Jinzikeng，1 000 m，on soil，Y．一H．Yan47．
158．Polytrichastrum formosum(Hedw．)G L Smith．
— — Huashuishan,IBSC Geb．Dept6097．Jinzikeng，
1 400 m， on crevice ofa peak， Y．一H． Yan46．
Midway from Jinzikeng to Chuandiding，1 1 75 m，
on rock in sparse forest，Z2786．This species had
been reported from Dinghu Shan in 1982 but had
no specimen citation[ ． W e treated it as a
doubtful record for Guan gdong in the update
checklist of Guangdong bryophytes ．Therefore，
the occurrence of this species in Guangdong is
now confirmed．
Acknowledgements
We WOUld 1 ike to thank Prof． Xing Fu—WH，
Mr． Yan Yue—hong， Mr．Chen Hong—feng， Mr．Zhang
Hua，Mr． Li Yuan—qiu，Mr．Luo Jian．ming and
other co1 1eagues for their invaluab1e assi s—
tance during field surveys． Partia1 support
was received with gratitude from Shimentai
Nature Reserve and the Director Fund of
South China Inst i tute of Botany． the Chinese
Academy Of Sciences．
2)He Z)(，Zhang L．A checklist ofbryophytes ofGuangdong Province，Chma．Submited to the Journal ofHatofi Botanical Laboratory,2003
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第6期 何祖霞等：广东石门台自然保护区藓类植物初报 551
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