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Effects of Elevated CO2, Drought Stress and Nitrogen Deposition on Photosynthesis Light Response Curves of Bothriochloa ischaemum

CO2浓度升高、干旱胁迫和氮沉降对白羊草光响应曲线的影响



全 文 :!24" !1#
 Vol.24  No.1
! " # $
ACTA AGRESTIA SINICA
   2016$  1%
  Jan.  2016
犱狅犻:10.11733/j.issn.10070435.2016.01.010
犆犗2x€y{、åæz{wK|}$t~Œ.
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2.&*+,QÏúQÙУ/01,‹ ƒ‘712100;3.‹:(¢*¾Y+“”Hl+,,‹ ƒ‘712100;
4.‹ùÃY+‹:‰Ž9\QÏÃĬ•­^C˓J®_,‹ ‹710048)
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:20141113;LMJK:20150331
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Âj
(1987),—,z:ЩÉ,ÊË,²Y,Ì«ÍÎÙC?9KÒÓ/0,Email:xiaosha525@163.com;<Ôc™ Au
thorforcorrespondence,Email:xuesha100@163.com
78

n—˜¦v-
(犅狅狋犺狉犻狅犮犺犾狅犪犻狊犮犺犪犲犿狌犿 (L.)Keng.)DàĊ‹<ðT‘š9ù8·VŠ£),Ôd³Pb
BC

/0¡¦v-L6@CO2 ×A、Q=ÍÖ6äiQRœX´TX•u»`6‘VŠéf8·。efgh:
ÀCO2 ×Aês6äi`íî,¦v-X´犛犘犃犇,ÝY%‘šØÝ(犘犿犪狓),gD`m:Ý(犃犙犈)6‘W6­
(犔犛犘)©„íîóI;š‰ú5€¡犛犘犃犇,犘犿犪狓6犔犛犘。5uÒØ=•gh,CO2 ×A,Q=Óù6äi
QRD犛犘犃犇,犘犿犪狓6犔犛犘 Jú…†UV,äiQRD犃犙犈Jú…†UV。CO2 ×A6Q=ÓùD犘犿犪狓,犃犙犈
6犔犛犘J…†Tæöcd;CO2 ×A6äiQRD犘犿犪狓,犃犙犈6‘ž­(犔犆犘)Jú…†Tæöcd;Q=Óù
6äiQRD犘犿犪狓,犃犙犈6犔犛犘 Jú…†Tæöcd。RDA=•ghiuÚUV¦v-‘š8·Tݕ«5
u

àÄCO2 ×Aês6iÑíîDš‰ú5ʪT¦v-‘šØݜJ…†TŒ”cd。
9:;

¦v-
;CO2 ×Aês;š‰ú5;iÑ;‘VŠéf
<=>?@
:Q945.11;S543.9    ABCDE:A     AFG@:10070435(2016)01006907
犈犳犳犲犮狋狊狅犳犈犾犲狏犪狋犲犱犆犗2,犇狉狅狌犵犺狋犛狋狉犲狊狊犪狀犱犖犻狋狉狅犵犲狀犇犲狆狅狊犻狋犻狅狀狅狀
犘犺狅狋狅狊狔狀狋犺犲狊犻狊犔犻犵犺狋犚犲狊狆狅狀狊犲犆狌狉狏犲狊狅犳犅狅狋犺狉犻狅犮犺犾狅犪犻狊犮犺犪犲犿狌犿
XIAOLie1,4,LIUGuobin1,2,ZHANGJiaoyang3,YANGTing3,XUESha1,2
(1.StateKeyLaboratoryofSoilErosionandDrylandFarmingontheLoessPlateau,InstituteofSoilandWaterConservation,
NorthwestA&FUniversity,Yangling,ShaanxiProvince712100,China;2.InstituteofSoilandWaterConservation,Chinese
AcademyofScienceandMinistryofWaterResources,Yangling,ShaanxiProvince712100,China;3.ColegeofResourceand
Environment,NorthwestA&FUniversity,Yangling,ShaanxiProvince,712100,China;4.StateKeyLaboratoryBase
ofEcohydraulicEngineeringinAridArea,Xi'anUniversityofTechnology,Xi'an,ShaanxiProvince710048,China)
犃犫狊狋狉犪犮狋:Inordertostudythephotophysiologicalcharacteristicsofoldworldbluestem (犅狅狋犺狉犻狅犮犺犾狅犪
犻狊犮犺犪犲犿狌犿 (L.)Keng.)fromloesshilygulyregionunderthebackgroundofglobalclimatechange,the
chlorophylcontent(犛犘犃犇)andphotosynthesislightresponsecurvesweredeterminedunderpottedcon
ditionswithdifferentatmosphericCO2conditions,soilmoistureconditionsandnitrogendepositiontreat
ments.TheresultsshowedthatelevatedCO2andnitrogendepositionincreasedthe犛犘犃犇value,maxi
mumphotosyntheticrate(犘犿犪狓),apparentquantumefficiency(犃犙犈),lightsaturationpoint(犔犛犘);
droughtstressdecreasedthe犛犘犃犇value,犘犿犪狓and犔犛犘.Multifactoranalysisofvarianceindicatedthat
CO2concentration,soilwaterconditionsandnitrogendepositionlevelalhadhighlysignificantinfluence
onSPADvalue,犘犿犪狓and犔犛犘;andnitrogendepositionlevelhadsignificantinfluenceon犃犙犈.CO2con
centrationandsoilwaterconditionshadsignificantinteractioneffectson犘犿犪狓,犃犙犈and犔犛犘;CO2concen
trationandnitrogendepositionlevelhadsignificantinteractioneffectson犘犿犪狓,犃犙犈andlightcompensa
tionpoint(犔犆犘);soilwaterconditionsandnitrogendepositionlevelshadsignificantinteractioneffectson
犘犿犪狓,犃犙犈and犔犛犘.RDAanalysisindicatedthatnitrogenwasthemostimportantfactorthataffectedthe
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photosynthesischaracteristicsof犅.犻狊犮犺犪犲犿狌犿.ElevatedCO2concentrationandnitrogendepositionhad
compensationeffectonthephotosyntheticreductionof犅.犻狊犮犺犪犲犿狌犿inducedbydroughtstressunderthe
backgroundofglobalclimatechange.
犓犲狔狑狅狉犱狊:犅狅狋犺狉犻狅犮犺犾狅犪犻狊犮犺犪犲犿狌犿;ElevatedCO2concentration;Droughtstress;Nitrogendeposition;
Photosyntheticresponsecurves
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Y`KLðè’ýi>CO2,YŠCO2 ×Aj
v1öJTØAíî
[1],
d2014$àÄYŠCO2
×AþÀíîÍ395.95μmol·mol
-1[2]。
ÏIPCC


d21ð"yàÄYŠCO2 ×AÐWd700
μmol·mol
-1[3]。
.e

èéCO2 ×Aíî1¤9
TÛË:Š"#àÄ
DàÄT9\Hl6Š
‹<ð¤9¡‚4UV
[4]。
ÛATês6sàF
T;<

"#š‰’nð÷<=2wx+LT„
þ
[2]。
é.@2

ð”dTY`íî+’iÑ
`TgØíî

µv

&þÀ’n«µ’


_ð÷!ñYiѯ&Ž
[56]。
LàĊ‹<ð
Tnoœ

YŠCO2 ×Aês、š‰ú56iÑ
íîˆö59

ˆöaš

t@DÜ_9\Hl¤9
…†UV
[7]。
‘šcdÚJK9Ÿ°JT9ù®¯

LàÄ
gGH&¶ç•«6p
[8]。
JK‘šcdT‘VŠ
éf·ðTڑ`m<`†A6JK%‘šØÝã
%T‰]

<=‘VŠéfÕÛLj—kJKTÝ
Y%‘šØÝ

gD`m:Ý

Í5^ØÝ

‘W6
­6‘ž­?9ùžë

.}žëÚ/0JK‘
šcd?9ù9\=ÄDàĊ‹<ðVŠTÿ
Ï
[912]。
JKT‘šcd67c¶B*+8TU
V

e÷Hl5uAÚUVJK‘š9ù8·T•
«ÒÓ
。CO2 ÚJK‘šcdT.ý,s×AT
CO2 ÿ[\JKT‘šØÝ,IAJ/0ghŸ#
9ŸLs×ACO2 œÿJK‘š§¥œ,OP
‘šKŠPR
[1314]。
QÚJK\‘‘šcdT.
ý

Q=ú5Ð"#JK‘šØÝ

Š¸"A6°ð
ØÝ?ŠNæہ‚u

Æò"#JKTÝY%‘
šØÝ6gD`m:Ý?‘VŠéf8·žë
€
[1517]。
iuÚJKNX•uT䒒=


Rubisco?‘š½j|p’@ð¥T ATP?KF
T䒒=

iÑTíîÐ[\JKX´X•u
Tš’

›sX´T‘š§¥
[1819]。
µvDéàÄ
Š‹<ðœ÷5u

j|Y5uæöcdœJKT
‘š8·z¡Y`T/0
[2021],


DCO2 ×A
ês

š‰ú56iÑñM5uæöcdœJK
T‘š9ù8·/0]¼ri½

8¾ÚwÙs.
.G8ß_ùHlœTèÙ-G¦v-
(犅狅狋犺狉犻狅
犮犺犾狅犪犻狊犮犺犪犲犿狌犿 (L.)Keng.),ZD‘DñG5u
ӚcdœT9ùVŠ]öri½

»ŠjwÙs
.EF-»JK¦v-n/0DR

ÔdË2³P
bÂ&BCç/0CO2 ×Aês、š‰ú56i
ÑD‘X•u»`6‘VŠéfTUV

(Ln
/0öçàĊ‹<ðnoœJK‘š9ù9ð8
·TVŠ›œ*+ÿÏ

1 UVWXY
1.1 jkÛµ
»/0Ôd³PbBC

é2014$6%‘

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³PŒ¹n¶TPVC<º(15cm×20cm,2Â
ן),<º&¹>Â500µTKœ3Qcn}wQ
(9cm×18cm,"Âן),³PÙCn:»ü
Tw-Ù

ŒjL}wQ2¸ƒÐšTw-Ù

·
_L}Qe=<ºžú¼Jè

LJèS>Âß}
sO½Ó2cm,2Ân2cmTPVCøcnïQ<
½

ã_L}Q>¹îƒw-Ù

O=œQ³&
TÙCœàŸë
,2014$6%9òL}wQ2Ôd
¾bTÒâbG¦v-Gm

¿³­f3M¾,¿
¾b3]Gm,O=œQ。
2014$8%1ò,¿³¿¾Ð@1rŸIˆ
äTÈÉ

¿³Ð@3rÈÉ),·_гPރ
ÉʋË
(AGCD003NtlF,/4ÚÉÃ
HlJAºp

&\‘CO2 ×A±í,6@äiQ
R6š‰ú5Óù

BC²2MCO2 QR,ˇ
PCO2 ×A(400μmol·mol
-1)
6±íCO2 ×A
(800μmol·mol
-1),
d¿O¸CO2 cneúCO2
œŠ”

¿á24h6%ÿœŠ;3M NuQR,Ë
0gN·m-2·a-1,2.5gN·m-2·a-165.0
gN·m-2·a-1;3MQ=@AÓù,˕Aš‰
ú5
(35%~40%Tã%£Q`),iAš‰ú5
(55%~60%Tã%£Q`)6Q=O2(75%~
80%Tã%£Q`)。t²18MÓù(g1)。B
CÔdà5m²²

¿GÓù5M•*。CO2 ×
Aè2MÉʋËb,=¾²©n4006800
07
!1#  j?:CO2×Aês、š‰ú56iÑD¦v-‘VŠéfTUV
μmol·mol
-1,
‘XÍֈ@

ŸA
:55%;‘Ò500
μmol·m
-2·s-1;ÛA:28/22℃(À/Á))。iu
QRèe”¼îÖW×b

g2),àúiu=6
“=¾é8%18ò,9%2ò,9%17ò,10%2
ò
,10%17ò611%1òäîé³P&。¿“
äi2–0.004205g·mL-1TÖW×Ûá,d
ÞáÂ=¾Ð0,5,10mLTÖW×Ûá©QTÝ
ÃLN1,N26 N3ÓùT³P&,·_dÝÄL
³PgÓÝQ

"ÐÖW×ÛáЃd³PÙC
&

nÜ[ô#ÙCýQD¦v-9ŸTA

é2014$9%17ò¼½\‘³PbQÓù。³
PÙC»Q`ÔdS•â\‘éb

Ë¿
áœÅ\‘S•

OKLTQ=

‘¢£Lß
©QR

`1 ‚no`
Table1 Pottreatment
äi`
Nitrogenapplication
rate/gN·m-2·a-1
CO2×ACO2concentration/μmol·mol-1
400 800
Q=»` Watercontent Q=»` Watercontent
35%-40%FC 55%-60%FC 75%-80%FC 35%-40%FC 55%-60%FC 75%-80%FC
0 C1W1N1 C1W2N1 C1W3N1 C2W1N1 C2W2N1 C2W3N1
2.5 C1W1N2 C1W2N2 C1W3N2 C2W1N2 C2W2N2 C2W3N2
5.0 C1W1N3 C1W2N3 C1W3N3 C2W1N3 C2W2N3 C2W3N3
`2 ‚ƒK1`
Table2 Nitrogenapplicationrate
äNÓù Nitrogenapplicationtreatment N1 N2 N3
äN(Ú`)Nitrogenapplicationrate/gN·m-2·a-1 0 2.5 5.0
¿³äN` Nitrogenapplicationrateforeachpot/g 0 0.044 0.088
äNH4NO3`(Ö×&ÚN»`(35%)AmountofNH4NO3/g 0 0.126 0.252
¿“äNH4NO3` AmountofNH4NO3foreachtime/g 0 0.021 0.042
1.2 ²³PQWXY
1.2.1 犛犘犃犇 ûbc 2014$10%10ò(š‰
ú5Óù23d_,¦v-¼xv#)ÀÁf¿M
Óù&T3³dò»¤SPAD-502X•uTé
X´犛犘犃犇 ,é2(Ýäœà±¼TÂX,
¿³•*é3“。
1.2.2 üýþÿ!bcs"#†$ Lé6@Ó
ù¦v-X´犛犘犃犇T@2鑑VŠéf。
=¾é2014$10%11ò610%12òDñMÉÃ
Š‹Ë&T¦v-X´‘VŠéf\‘é

dLi
-6400¶Üºr‘”(Li640002BLED)Â&‘–@
A2000,1600,1200,1000,800,600,400,300,200,160,
120,80,4060μmol·m
-2·s-1t14M,²©;<
‘Ò–A_

Ýu¨©2%60s,•*3“。
/0gh¦v-‘VŠéfTÝnÂFnÈ"
6YéfÂF
[20],
‘gWân
犘狀 =犪犘犃犚+犘max-
(犪犘犃犚+犘max)2-4θ犪犘犃犚犘槡 max
2θ -
犚犱
  â&:犘狀 n%‘šØÝ(μmol·m
-2·s-1);θ
n‘šéf9éÄATÆA
;犪(AQE)ngD`m

(μmolCO2·m
-2·s-1);犘犿犪狓nÝY‘šØÝ
(μmolCO2·m
-2·s-1);犘犃犚 n‘šJ:Çb
(μmol·m
-2·s-1);犚犱 nÍ5^ØÝ(μmolCO2
·m-2·s-1)。
1.3 lmno
ÔdPhotosynthesisWorkBenchÕÖD‘V
ŠéfTžë\‘²ß

ÔdSPSS16.0÷5uÒ
Ø=•
(ANOVA)D6@Óù%SPAD6‘VŠ
éfžë\‘غ…†QRT×C
(犘=0.05),
5uÒØ=•dé¸tCO2 ×A、Q=ÍÖ6ä
iQRj|XÛã%L§+LTæöcd

Ôd
SigmaPlot12.5cë。ÔdCanocoÕÖ\‘CO2
×A

Q=ÍցäiQR6¦v-‘VŠéfž
ëTÈ/=•
(RDA)。
2 pqW>r
2.1 ´„犆犗2 x€y{、åæz{wK|}$t
~Œ/…犛犘犃犇Ç2|u
¦v-X´犛犘犃犇Téef(ë1)gh,
CO2×A±í›s¡X´T犛犘犃犇,òLš‰ú
5$6äiÍ֜
(W1N1,W1N26 W3N1)Wd
…†QR
(犘<0.05)。@2,Àìäi`Tíî,¦
17
! " # $ !24"
v-X´犛犘犃犇 ©„íîóI。nÀìÙCš
‰ú5TîŸ

¦v-X´犛犘犃犇6q‚u。
5uÒØ=•efgh6@CO2 ×A,Q=ÍÖ6
iÓùD犛犘犃犇TUVغ©Wdú…†QR,
ICO2 ×A,Q=ÍÖ6iÓùD犛犘犃犇 ï…
†Tæöcd

g3)。
=1 c~犆犗2 x€、ó>wKno$t~Œ/…
犛犘犃犇Ç2|u
Fig.1 EffectsofCO2concentration,waterconditions
andnitrogentreatmenton犛犘犃犇value
ü

6@;Çg˜Øº…†
(犘<0.05)
Note:Differentlettersindicatesignificantdifferenceatthe0.05level
2.2 ´„犆犗2 x€y{、åæz{wK|}$t
~Œ.uv€2|u
6@CO2 ×A、Q=ÍÖ6äi`Í֜¦v
-X´‘VŠéf~ë21˜。èëLã,L‘Ò
–An0~400μmol·m
-2·s-12,X´T%‘š
ØÝgØSê

-‘–Wd400μmol·m
-2·s-1
_

%‘šØÝSêìA‚o

ã_6q¨©
。CO2
×Aês›s¡¦v-TÝY%‘šØÝ
(犘犿犪狓),
gD`m:Ý
(犃犙犈),Í5^ØÝ(犚犱),‘W6­
(犔犛犘)6‘ž­(犔犆犘)(g4)。Lˆ@CO2 ×A
6ÙC»Q`Í֜

Àäi`Tíî¦v-T
犘犿犪狓,犃犙犈,犚犱 6犔犛犘 „íîóI。L@? CO2
×A6äiQRœ

Àš‰ú5Tv-T
犘犿犪狓6犔犛犘 „‚uTóI。
5uÒØ=•efgh
,CO2 ×AD犘犿犪狓,
犚犱,犔犛犘6犔犆犘TUV©Wdú…†QR(犘<
0.01),D犃犙犈ïh…UV;š‰ú5D犘犿犪狓,犔犛犘
6犔犆犘 Jú…†UV,D犚犱 J…†UV,ID
犃犙犈UV;äi`D犘犿犪狓,犃犙犈,犚犱,犔犛犘6
犔犆犘 ©Jú…†UV。CO2 ×A,ÙC»Q`6ä
iQRD‘VŠéfžëT8@‰]gPn
:CO2
×A6ÙC»Q`D犘犿犪狓,犃犙犈,犔犛犘6犔犆犘 J…
†Tæöcd
;CO2 ×A6äiQRD犘犿犪狓,犃犙犈
6犔犆犘 Jú…†Tæöcd;ÙC»Q`6äiQ
RD犘犿犪狓,犃犙犈,犔犛犘 Jú…†Tæöcd,D
犔犆犘J…†Tæöcd;CO2 ×A,ÙC»Q`6
äiQRD犚犱 6犔犛犘 Jú…†Tæöcd,D
犘犿犪狓6犃犙犈 æöcd。RDA=•efgh
CO2 ×Aês6äi`íYÿ›s¦v-X´T
犛犘犃犇,犘犿犪狓6犔犛犘,nš‰ú5ÿ€¦v-
X´T犛犘犃犇 ,犘犿犪狓6犔犛犘,‘&äiQRÚU
V¦v-‘š8·Tݕ«T5u

ë3)。
=2 c~犆犗2 x€、ó>wKnoP
t~Œ/….uv€
Fig.2 Photosyntheticresponsecurvesof犅狅狋犺狉犻狅犮犺犾狅犪
犻狊犮犺犪犲犿狌犿underdifferentCO2concentration,
waterconditionsandnitrogendepositiontreatment
27
!1#  j?:CO2×Aês、š‰ú56iÑD¦v-‘VŠéfTUV
`3 c~犆犗2 x€、ó>wKno$t~Œ犛犘犃犇Çw.uv”l2aø†X}>r
Table3 Resultsofmultifactorvarianceanalysisof犛犘犃犇valueandlightresponsecurveofphotosynthesisparametersof
犅狅狋犺狉犻狅犮犺犾狅犪犻狊犮犺犪犲犿狌犿underdifferentCO2concentration,waterconditionsandnitrogendepositiontreatments
Óù
Treatment
犛犘犃犇
value
ÝY%‘šØÝ
犘犿犪狓/μmol
CO2·m-2·s-1
gD`m:Ý
犃犙犈/μmol
CO2·m-2·s-1
Í5^ØÝ
犚犱/μmol
CO2·m-2·s-1
‘W6­
犔犛犘/
μmol·m-2·s-1
‘ž­
犔犆犘/
μmol·m-2·s-1
犉 犘 犉 犘 犉 犘 犉 犘 犉 犘 犉 犘
CO2 18.567 0.000 32.900 0.000 1.299 0.262 17.921 0.000 95.551 0.000 20.512 0.000
W 58.875 0.000 181.529 0.000 0.563 0.574 4.035 0.026 95.843 0.000 7.895 0.001
N 26.850 0.000 302.990 0.000 54.291 0.000 23.958 0.000 86.614 0.000 7.826 0.002
CO2×W 0.303 0.740 4.938 0.013 4.257 0.022 3.736 0.034 4.271 0.022 7.570 0.002
CO2×N 2.982 0.059 6.750 0.003 7.064 0.003 0.377 0.689 0.590 0.560 13.012 0.000
W×N 0.718 0.583 7.627 0.000 5.411 0.002 2.043 0.109 7.527 0.000 3.151 0.026
CO2×W×N 0.208 0.933 2.555 0.055 2.056 0.107 4.663 0.004 2.142 0.096 6.140 0.001
  ü:CO2:CO2×A;W:Q=;N:äi`
Note:CO2:CO2concentration;W:Waterconditions;N:Nitrogendepositiontreatment
`4 c~犆犗2 x€、ó>wKnoPt~Œ.uv€”l
Table4 Lightresponsecurveofphotosynthesisparametersof犅狅狋犺狉犻狅犮犺犾狅犪犻狊犮犺犪犲犿狌犿under
differentCO2concentration,waterconditionsandnitrogendepositiontreatment
Óù
Treatment
ÝY%‘šØÝ犘犿犪狓
/μmolCO2·m-2·s-1
gD`m:Ý犃犙犈
/μmolCO2·m-2·s-1
Í5^ØÝ犚犱
/μmolCO2·m-2·s-1
‘W6­犔犛犘
/μmol·m-2·s-1
‘ž­犔犆犘
/μmol·m-2·s-1
C1 C2 C1 C2 C1 C2 C1 C2 C1 C2
W1N1 9.10±0.18aA 9.13±0.36aA 0.030±0.001aA 0.040±0.003aB 0.60±0.15abA 0.62±0.37aA 547±44aA 621±15aB 16±4abA 20±12aA
W1N2 10.27±0.68abA12.28±0.71bB 0.050±0.005cdA0.063±0.007cdeA1.12±0.23cdA 1.57±0.35bA 565±23aA 669±79abA 23±6bcA 24±4abA
W1N3 12.72±2.00cA 14.54±0.44cA 0.062±0.013deA0.070±0.008deA1.15±0.42cdA 1.70±0.27bA 667±44bA 705±27abA 19±6abcA 20±0aA
W2N1 10.55±0.26abB 9.53±0.24aA 0.037±0.003abA0.053±0.005bcB0.97±0.22bcA 0.86±0.26aA 547±58aA 711±27abB 19±2abcA 23±6abA
W2N2 12.91±1.08cA 12.74±0.35bA 0.052±0.002cdA0.061±0.003cdB1.23±0.11cdeA 1.52±0.12bB 652±45bA 753±12bB 20±4abcA 31±2bcB
W2N3 15.89±0.70dA 18.16±0.31dA 0.058±0.014deA0.058±0.007bcA1.54±0.19deA 1.54±0.45bA 800±54cA 975±83cB 27±6cA 20±7aA
W3N1 11.60±0.47bcA12.98±0.89bA 0.042±0.001abcA0.049±0.010abA0.56±0.14aA 1.85±0.11bB 660±35bA 749±51bA 12±0aA 45±2dB
W3N2 15.14±1.12dA 17.98±0.40dB 0.044±0.001bcA0.056±0.003bcB1.15±0.27cdA 1.42±0.43bA 760±45cA 984±7cB 27±6cA 39±2cdB
W3N3 18.92±0.96eA 20.58±0.22eB 0.065±0.005eA 0.072±0.002eA 1.60±0.18eA 1.76±0.37bA 884±52dA1039±59cB 21±2bcA 17±2aA
  ü:@j6@uU;Çg˜6@Óùã%غ…†(犘<0.05),@‘6@YU;Çg˜6@CO2×Aã%غ…†(犘<0.05)
Note:Differentlowercasesinthesamecolumnindicatesignificantdifferencebetweendifferenttreatmentsatthe0.05level;Differentcap
italletterinthesamerowindicatesignificantdifferencebetweendifferentCO2concentrationatthe0.05level
=3 c~犆犗2 x€,ó>wƒKóôW
.uv€”l2犚犇犃>r
Fig.3 RDAanalysisofCO2concentration,water
conditions,nitrogentreatmentwiththelightresponse
curveofphotosynthesisparameters
3 ˆ‰
3.1 ´„犆犗2 x€y{、åæz{wK|}$t
~Œ/…犛犘犃犇Ç2|u
犛犘犃犇ÚJKX´TˆDX•u»`Éë,
‘TYuX´TX•u»`„‡ˆ‰

Ljd
çuvJKX´X•u»`
[2223]。Bloom?[24]/
0…˜Ÿ#s×ACO2 ÓùÿUVJKX´X•
u»`

xÊ?
[25]
°PYŠCO2 ×AêsLj›
s„xX´X•u»`

L»/0&
,CO2 ×Aê
s›s¡¦v-X´TX•u»`

iÚJK9Ÿ
=Ä&ݕ«TD=A5m

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X•uTš’

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íî

JKX´X•u»`…†íî
[2627]。
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,犛犘犃犇 ÚuvJKX´‘
š§¥T•«»F

3.2 ´„CO2 x€y{、åæz{wK|}$t
~Œ.uv€2|u
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。CO2
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›sJKT‘šØÝ

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CO2 ×Aês›s¡JKTÝY%‘šØÝ6‘
W6­

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[28]
DCO2 ×A±íœÊä
6,ËQ?
[29]
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iuT¼î§[\JKX•uTš’

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CO2 ±ícK.%CO2 ×A¢£Lß©TQR,
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‚iš‰ú5DJ/‘š
cdT6ÏUV
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‰ú5œCO2 ×Aês…†›s¡¦v-TÝY
%‘šØÝ6gD`m:Ý
。CO2 ×Aêsÿ…
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œacd
[34]。
én?
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6pJÐ?
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í?
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6‘W6­ÀÙCQ=€nœ

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œìAóY

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©D¦v-T%‘šØÝJ…†Tæöcd

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iÑTíîLj…†›sš‰ú5œ¦v-‘š
§¥

4 p‰
  LàĊ‹<ðTnoœ,CO2 ×Aês6i
ÑTíîÿ…†›s¦v-X´T犛犘犃犇 ,Ý
Y%‘šØÝ6‘W6­

nš‰ú5ÿ"#¦v
-X´犛犘犃犇,ÝY%‘šØÝ6‘W6­T
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T‘š§¥J…†Tæöcd

niuQRÚUV
¦v-‘š§¥Tݕ«T5u

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Lj…†‚iš‰ú5+’T¦v-‘š§¥œ
TUV

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%&( )*+)
57