免费文献传递   相关文献

Effect of ABA on transcriptionally active Ty1-copia retrotransposons in Dendrobium officinale

脱落酸对铁皮石斛Ty1-copia类反转录转座子转录活性的影响



全 文 :ÙÚ ÛÜÝÞ
?65!#$%
ßàáâ
áVŽáâã³™š
$s
!
tuv
!
wxy
!
!
z.{
!
|}

ÁY[0B ½ÂÃ^[Ĉ¼€‚ƒÄˆÅy ÁY“ÆÇ\Èɑ
ÊË̱ÍÎÏÐ
!
ÁY ÑË
))$$#

de
#!
ƒ„Ã
?65!#$%
ßàáâáVŽàáâ£L‰ùǼ»
$

$$
!
/.V-F
g
ÙÚ 
!3D3
‰zÛÜÝÞ
…dàáâ
@<>!>?5@<>

$
°g
*#$ ^*äOCR
$
ËÉ
$$
!
/.V-F
g

3D3
º«¬
?65!#$%
ßàáâáVŽ
áâ㳧¨
)
&OCRÛ
&
¤¥•¿1}˜©°g
&*
C
?65!#$%
ßàáâáVŽàá⣉U1Ì
$
>1Ì@
ƒ³Ò
$
I٠1̱
*&% j*## ^\$
l@³Ç
&#c)i j4(c4i)
w
\VN1Q<3>X
ÙD˜©cò‡ÛÜÝÞxþ ˜T
?65!#$%
ßàáâáVŽAÿ
$
‹¸4I¡éêCDŠ‹ÚuwD
&
4I󭎲ú3Å
?3?3^.7
•
<33?^.7
‰U
1̦ŠA[ÚuwD
*
]Dáâã³ÛÜÝÞ
?6V5!#$% >?
1Ì$‡éêCDŠ‹ÏçÚuwDÉʱÍ
)
yz—
2x }
$
D

C1Ì&ò~l9fˆ‡TŽ@
$4
C1Ì&òtT@
$
‰U1Ì
8FLKGa(
m1c”~l9
@
)
Š2x 1Ìw´ÿ{ß}G˜Ç

ß
*
³‡xþ
?6V5!#$% >?
1̏ß}cò4?c”áâ
?6V
þàáâá

>?
1̇xþ
?6V
þàáâáVŽ
>?
1Ì.$çÿÒO
$
ËÉw
3D3
«¬}c”áâ
?6V
þàáâáVŽ
>?
1
̇xþ
?6V
þàáâáVŽ
>?
1ÌDT‚ƒ
)

fgh
#!
ÛÜÝÞ
*
ÙÚ 
!3D3* ?65!#$%
ßàáâáVŽ
*
àáâ£
!>?*
áâã³

ij)(
#!*$)5$5*4

klmP
#!
ÕÜç/&(xN
!*$*<*4*$*$<*$$*

nopq
#!
!
´?+
$
XPY
$
ê*º»”»()•‡@ŒQ
$
X5/NHV#MN.6N1OWH&$(2*#c-./

pqrs
#!
œ¯
$
ZWƒ„”
$X5/NHV# VT1HV-2#)c-./
!!
àáâáVŽ=»‹¸yº»xþ$
+,
$
º¨±Íxþ:
$
º»xþ…¢p+v
”/Ί‹
+*,
)
èyàáâáVŽýÃ
Y;35
>;35Y;3
…d±ÑáVwD
$
á⸗Çuø
Šã³§¨ç„Fl
)
A‰¶·
$
àáâá
VŽÃéˆóD‹¸yº»$
$
]1˜àáâá
VŽjkDáV|º
$
º¨žn@”»•õ”»
éê Xã
+),
$
|‹Œ6”;ϘT
&
þ•ç~
&

&
ÙÚ 
!3D3&
jE ÆT
!9TC3
•¬Õ
 
!83
Cõ”»é꺨)´
?1QV
•
?Q.V
à
áâáVŽ§¨
+&5,
)
lQ©4”»ëpG
ì“àáâáVŽáâXã
+#,
) ?N1M
C
+%,
¸
:<.$)$!/ %67)$5/Fc
$còàáâáVŽ
?N>?5
º¨ž
9TC3$83
¦›èÔÌ
! \.+UTS6/HV5
UT+RW1MWP
«¬
$
Š§¨¬¯ÉÊ)´
)
§%
$
‹

$9TC3$83$3D3
C‡éêÏçl<˜Ž‰
z1º¨XãŠã³
)
ÛÜÝÞ
P61:.C$/+$!$1%(6GH/WSNTQ9HM.
ýY{p7!$,™
$
òR,zƒ„§ÉkDi
xӓ
&
Q”×–Cr²¹×
+(,
) *$
³Ê
4$
oR
Ãz
$
ÛÜÝÞþ•ç~
&
’©ðçCAÿÌç„(
)°‡/‚
$
ÛÜÝÞv+°‡k{_c
†
+45,
)
]ÛÜÝޒ©ð狸”v—a´
&
O
ŒõÚô
&
Ôdjc”Càá
)
P:Eàá
$
ÕÜ
çÛÜÝÞv+()±²n²cɇÛÜÝÞ6”
Dðç()
$
`ŒQû³‡@ýÆò6”Dðç
ç„
+*,
)
§
$
aëB´ÛÜÝÞÙÚ 
!3D3
Šxþ8
?65!#$%
ßàáâáVŽáâã³
™š¦kDáâã³>ßàáâáVŽ²ú
ƒ„
$
‡ÄàáâáVŽû²ø
$
ÃvÇÛÜÝÞ
ŒQû‡@)Id4
)
!
™6
I«ÛÜÝÞ@;
<$
wÕÜòÁ°M
,»ƒc$UâC¯PYfÐÇÛÜÝÞ
Pc+$!$?
1%(6$
6vÕÜè<
!)$q)o;$4q*oX$
ÿÛÜÝ
ޘÅ2$
$
kDû[(À
)
Y;3X7<%N
£
$9M*
$$ sDWRTS$U;?@P$
¤
-((%-
t
)4
Öt
$
v
*$&
o

p
!!!!!! ! ! ! ! ! !!!! ! ! ! !
A.Vc)4$:PPWT!$
9N6$ *$&
¥tÏ
\9Y
.
(5?AT-Q.S$>;3HP.@VWP!?.QNV>;3
)«ü

«ük
$-Y;3
“—«ük–$L
?NGN5
>N
FG
*
¤¥‹Ì
YB
#ÇaÆÇe‰‹
*
|ñÛ«ük$L
_LKX;
”»()
!
³™

DE
FG
*
¼»è:J”r”»r9
!
:J

´qDEF
G“—
)
(!
}~
(_!
™6‰z
ÂI«@;@Ž
$
¸ÕÜòÁ0–RµÃ
çQ%K/RÆÇeçQxHþçeþ•ç~
!
’
òv
# O-U
g
$
¾
* n$
ç~
($ U
Q”±
óô
&
cQóDA“þç°
$
ê^x1fõõ3
$
&þç°¶Hdƒ¬$
* O
déê
$
M}
&þç°îyÔÇ
$$
!
/.V-F
g

3D3
ö1
$‰z
$
‰zQb˜gÇ
)$$ #$$ *$$ ($$ *&$
/H1
+),
)
Âòw´
3D3
‰zþ簊Ç,›
Õ
)
M}&ÛÜÝÞ[:î1‚fõŠ·
$
‹y
>;3

)
(_(!
ÛÜÝÞ
>;3
)Â
Ê՜5¸C
+&,
îó
}~)ÂÛÜ
ÝÞ
>;3$
¹•Ô‹˜’’|
!;N1.US.*$$$
¿
$
øû³‹
c$i
míâ|ñW’¿
)
(_&!>?5@<>
(_&_!
àáâ
!
&Õþ
>;3
•
3D3
«¬þ
>;3!
&
$$
!
/.V-F
g
3D3
~lQb8‰z}
™6 )Âk
>;3
C(𓠰
$
˜g‹
?N[N5
SN
FGàáâ«ük
>;3
…dàáâ
)
(_&_(!>?5@<>
àÀ
!
ƒ4
KT1DN1[
$
?6?!#$%
ß
F?>
àáâáVŽ
>?
x‰UL²R
$
Œw
‡
GW/NS
C
+,

?6?!#$%
ß
F?>
àáâáVŽ
>?
1̒|ùǼ»
?65I!o53<;K<;?5
?LL?;<3LKK5)o
•
?65>!o53><3?>?<>?<;35
<>?35)o*
8ÊxÇÛÜÝÞ
3-QH1

# N-QH15I
!o5??K?K??KK3??•
N-5
QH15>!o5???<<¼
»–è:J”r”»r9
!
:J

´qDEFG
“—
)
>?5@<>
àÀÏÿ
!*$
!
F#$ sDWRTS*
!
F**
//.V-F
g
9M
* h
c*
!
F**c //.V-F
g
U;?@P$c(
!
F*$
!
/.V-F
g
¼»
!?65I
•
?65>$c(
!
F* E
-
!
F
g
<%N
£
$c
!
F*-Y;3
—q
&
!
F) >?5@<>
r±àÀ91Ç
#4& n
x@³
/H1$4& n


/H1$&#c n
a~
c& /H1$%* n
Ês
/H1$%* n
Ês
/H1)
Š$O5x
)
IwO
$
8Êx
*
IwO
)
(_)!@<>
v»Û
&
¤¥¦¿1
‹
i
míâ|ñ¿r±v»
$
Nj|ñ
Û«ükÛǹ¢
@<>

$
6”&Ûv»z
”y
\9Y
.
(5?

!?NGN>N
FG

:
$
ᢁ
Ì
YB
#á¢u¡DËr
$)% n
ç~
* j# O
…
d®›Ó‹Œ
$
u›…¤¥yFD
$ /M-F
g
2
º±eë
FD
1Ïç~x$ç~
& j# O
}
$@<>
fТ³¤¥
)
&fÐÇ¢³Ì1˜gÂ
)$$
!
F$
è:J”r”»r9
!
:J

´qDEFG¿
1
)
(_$!
àáâ£1̘©
¿13ØB´
;$
DF38?
91…dl@³
я
$
€‹
ÙD…d4/я
$
‹
9XK3
c$
ÙDƒ”~Åbÿ{cQ…¢
)
&!
3؇˜©
&_!3D3
‰z
F?>
àáâáVŽ
>?5@<>

ê
>?5@<>

?6V
þàáâáVŽ
>?
x
3Øzî
$
 °r±+RÉÊ
&
’-R
$
:8:¸
*#$ j*%$ ^\$
‡xvO:8:A“
*
w
$$
!
/.V
-F
g
3D3
«¬}r±+RÉÊ>y,›Õ
r±+R
$
ËÉ
$$
!
/.V-F
g
3D3
ÛÜÝÞ$
?65!#$%
ßàáâáVŽáâDÉÊ«¬Š‹
)
&w
3D3
«¬ °
>?5@<>
v»w:8Û
&
¤
¥Ç¿1
$
w
DVNPQ
я•
Y;3PQNS
˜©}
$
š:°
&*
CDáâã³àáâáVŽàá⣉U1Ì
!;?4{Yâ<
GC&%$) jGC&%&&$
g
)
9cY;39NS[TS*òw
3D3
‰zÛÜÝÞ
>;3

*3D3c
w
3D3
‰z
)$$#$$*$$($$*&$ /H1
ÛÜÝÞ
>;3
ð“ÿr±
)
g
!
ÛÜÝÞàáâáVŽàáâ£1Ì
@<>

!3
•
$$
!
/.V-F
g
3D3
ÛÜÝÞ$
?65-.\HN
ßàá
âáVŽ§¨™š
!D
IHMc!@<>N/\VHRH-NQH.1 .RSTkTSPTQSN1P-SH\QNPTPTbWT1-TRS./
P61:.C$/+$!$1%(6!3N1U T7\STPPH.1PN16V6PHP.R?6V5!#$%VH[T
STQS.QSN1P\.P.1PT7\.PTU Q.$$
!
/.V-F
g
3D3QSTNQ/T1Q!D
-4(%-
t
)4
Öt
$
v
*$&
o

p
!!!!!! ! ! ! ! ! !!!! ! ! ! !
A.Vc)4$:PPWT!$
9N6$ *$&
&_(!3D3
«¬}kDáâã³ÛÜÝÞ
?65-.5
\HN
ßàáâáVŽ1̘©
&_(_!
ÛÜÝÞ$kDáâã³
?65!#$%
ß
>?
IÙ 1ÌÑÒ
!
&:°
&*
CkDáâã³
àáâáVŽàá⣉U1Ìy
;?4
{…d1Ìя
$
3اÉj‰–‡ºtº»
?65
!#$%
ßàáâáVŽ
>?
1ÌDT´ϓ³
$
1́:Ç
*&% j*## ^\!
§
$
¹‡xþ
?65!?
#$%
ßàáâáVŽ
>?
1ÌxaA“
+#,
)
 D
1̖$F
3?
Jx
$

9XK3c$
ÙD|ê& 
D1Ì
3?=K<
ъÇ
c* jc(4 !
§
*

Y;393;
ÙD|ê & I Ù   1 Ì l @ ³ Ç
&#c)i j4(c4i$
Š$1Ì
<53D35&
‡
<53D35
$<53D35$<53D35)*
l@³­´
$
–Ç
4(c4i$

<53D354
‡
<53D35*
l@³
­ñ
$
£Ç
&#c)i)
ËÉÛÜÝÞxþ$GºD
4I
?65!#$%
ßàáâáVŽKñáâ㳺
ž
3D3
«¬
)
§
!
ÛÜÝÞ$kDáâã³
?65!#$%
ßàáâ£1Ìþ—
?N^VT!?OT1W-VT.QHUT-./\.PHQH.1 .RSTkTSPTQSN1P-SH\QNPTQSN1P-SH\QH.1NV6N-QHkNQTU .R?65!#$% STQS.QSN1P\.P.1PH1 P61:.C$/+?
+$!$1%(6
1Ì< :8:
3?=K<
1Ì< :8:
3?=K<
1Ì< :8:
3?=K<
<53D35 *#) c%& <53D35 *#) c <53D35*4 *# c#
<53D35* *#$ c%& <53D35# *# c&( <53D35)$ *# c*4
<53D35) *## c#4 <53D35% *#& c4 <53D35) *#& c#
<53D35& *#* c#* <53D35( *## c&4 <53D35)* *4 c(
<53D35 *#& c4 <53D354 *#* c* <53D35)) *#& c#%
<53D35# *#) c( <53D35*$ *#& c&4 <53D35)& *#& c#
<53D35% *# c <53D35* *#& c#& <53D35) *#* c)*
<53D35( *#* c)# <53D35** *#* c% <53D35)# *#& c#
<53D354 *#$ c* <53D35*) *# c&* <53D35)% *# c%
<53D35$ *#& c# <53D35*& *## c# <53D35)( *# c&(
<53D35 *# c* <53D35* *#) c(# <53D35)4 *# c%)
<53D35* *#) c(4 <53D35*# *#* c#* <53D35&$ *# c%
<53D35) *# c) <53D35*% *&% c# <53D35& *#& c&4
<53D35& *# c <53D35*( *#& c#% <53D35&* *# c*
!!
y‹ÙD
\VN1Q<3>X
˜©
$
cò:°w
3D3
«¬}Dáâã³
&*
CÛÜÝÞ
?6V5!#$% >?
1
Ì$‡xþ:°
?6V5!#$% >?
1ÌAÿ
$
‹¸
4I¡éêCDŠ‹ÚuwD
&
4I󭎲
ú3Å
?3?3^.7
•
<33?^.7
‰U1̦Š
A[ÚuwD
*
]‡xþ:°
?6V5!#$% >?
1
ÌLgý
#
¸Dáâã³ÛÜÝÞ
?6V5!#$% >?
1Ì$‡éêCDŠ‹ÏçÚuwDÉʱÍ
$
Š$FD™7•éêo_ÚuwD1ÌC? M
ÑÈèxþ$
&c%#i
±Íg
&c*&i$
¹˜˜
ËÉ
3D3
GÃXãÛÜÝÞ$A[F‡éêÏç
ÚuwD
?6V5!#$%
ßàáâáVŽ
)
&_(_(!
ÛÜÝÞ$kDáâã³
?65!#$%
ß
>?
2x 1ÌÑÒ
!
&¤¥°gkDáâã³

&*
C1Ìyz—2x 
$
ÊÕ
?65!#$%
ßàá
âáVŽ
>?
x2x ‰U1Ì
$
còD
4
C
1Ìc”‡tT@
$
C1Ìc”‡fˆ‡TŽ
@
$
Š$
<53D35*
•
<53D354
˜gc”‡

(

4
Ifˆ‡TŽ@
)
è§23
$
t
T@•fˆ‡TŽ@Gºý¬“ÛÜÝÞàá
âáVŽƒ;³/Î6¦A
)
‰U:8
8FLKGa(
+%,
!
t
)
I2x ¤k

.c”‡@
ƒ
$
Š$D
)(
C‹¸Jx»R@
$
š%
&
C
!<5
3D35*$<53D35&$<53D35))$<53D35&*
–
¸‰U:8¦Ëc”tT@`c”@ƒ
$
£D

C1Ì
!<53D35*%
‹¸{å@
!
g
*)
è:Gt
$
è
3D3
«¬kDáâã³
?6V5
!#$%
ßàáâáVŽ
>?
1̇xþ:°>1Ì
³;ϓ
$
ƒ;³>
$
1˜1Ì$‹¸fˆ‡TŽ
&
tX
&
{å¦Jx»RC@
)
-$4%-
t
)4
Öt
$
v
*$&
o

p
!!!!!! ! ! ! ! ! !!!! ! ! ! !
A.Vc)4$:PPWT!$
9N6$ *$&
À‹ÙD
1̅dя
$&
@~l9”™1˜§<1̉U9
*
҅1˜‰U³Ç
$$i*
Gã…1˜‰U³Ç
%i*
¼ã
…1˜‰U³Ç
$i*
^D”^1˜‰U³Ç:y
$i*_c
fˆ‡TŽ
*-
Ç(¢nv”{ó
$
äÆ?s§<1Ì$2x ?(
)
g
*!
ÛÜÝÞ$4Dáâã³
?6V5!#$%
àáâáVŽ
>?
x2x 1ÌÒ
IHMc*!?OTN/H1.PTbWT1-TNVHM1/T1Q.RSTkTSPTQSN1P-SH\QNPTQSN1P-SH\QH.1NV6N-QHkNQTU .R?6?!#$% STQS.QSN1P\.P.1PH1 P61:.C$/
+$!$1%(6
&_&!
ÛÜÝÞ
?6
þàáâáVŽ
>?
1Ìÿ
{cQ…¢˜©
&_&_!
ÛÜÝÞ$c”áâ
?6
þàáâáV
Ž
>?
1Ìÿ{cQ…¢˜©
!

9XK3
c$
ÙDƒ”~& °ÛÜÝÞkDáâã³
&*
C
?65-.\HN
ßàáâáVŽ
>?
2x 1ÌÅ
bÏÀÿ{cQ…¢
!
g
)$
˜—z@y
$$$
I/€L†¶
$
ƒ4…¢
^SN1-O
±G
½¹
&*
CkDáâã³
>?
1̘—

ß
)
Š
$-ߋF
*
I1Ì
$
ߋF
)
I1Ì
$
%ߋ
F
*
I1Ì
$
/ߋF
*
I1Ì
$
0ߋF
&
I

)
&_&_(!
ÛÜÝÞ$
-Y;3?65!#$% >?
1̕x
þ
Y;3?65!#$% >?
1Ìÿ{…¢˜©
!
&
 °
&*
CÛÜÝÞ
<
$˜T&kDáâã
³
?65-.\HN
ßàáâáVŽ
>?
1̇xþ
˜T°g
*
C2x 1ÌB´
9XK3c$
…¢
˜©ÙD…dÿ{…¢˜©
$
°g
;THMO^.S50.H1H1M
ÿ{cQ…¢
$
˜—z@y
$$$
I/€
L†¶
!
g
&)
c”áâ
?6V
þàáâáVŽ
>?
1Ì %
$
/
$
0߇xþ
?6V
þàáâáVŽ
>?
1Ì.$çÿÒO
$
t߇xþ$
<5
ÇAß
$
t-߇xþ$¯
<5
Ã%Š
1̖¸A†
)
ËÉw
3D3
«¬}c”áâ
?6V
þàáâáVŽ
>?
1̇xþ
?6V
þàáâ
áVŽ
>?
1ÌDT‚ƒ
)
)!
3f‡)f
º»ÏÕ7²øv”ÎÎd]yº»u¡X
X
&
v”¦YZl<
$
­f¼†”¢î@¦JR
TÚû
)
¸ûIl$
ÙÚ 
!3D3
ŠÇA@/ÎÚuŽ
$
j¸º»ø¡”»•õ
”»éêÚu$†ç„Š‹
)
oº»¡gn@
-4%-
t
)4
Öt
$
v
*$&
o

p
!!!!!! ! ! ! ! ! !!!! ! ! ! !
A.Vc)4$:PPWT!$
9N6$ *$&
!!
˜U9ƒ4
$$$
ì/€L†¶
$
—±R§Ãƒ”~
±ê&zUY|T
!
g
&
l
)
g
)!
ÛÜÝÞkDáâã³
?65!#$%
àáâáVŽ
>?
x2xx 1Ìÿ{…¢
IHMc)!@O6V.MT1TQH-QSTT.RN/H1.PTbWT1-TP.RSTkTSPTQSN15
P-SH\QNPTQSN1P-SH\QH.1NV6N-QHkNQTU .R?65!#$% STQS.QSN1P\.P.1P
H1 P61:.C$/+$!$1%(6
OŒéê|
#
Øä
&
´ï•[ŠCQ
$
ŠÏ8+fõ
“—•¢ë
3D3$
³2éê®å`!F¾€g
6Dñ¬¯
+(5*,
)
ƒ„§É
$3D3
º«¬(x
§¨
$
¸÷9¿$ÀwfÐ&
)$$
4@x
¡
3D3
ÚÛ
$
Š$A-§òÇ
3D3
ªø§¨
+**,
)
?6V5!#$%
ßàáâáVŽý
Y;35>;35Y;3

áVwD
$
áâýŠáV㳧¨qÎFl
$
¸Š
F?>P
$‹¸áâ†k‡fˆl<懫¬
§¨ÏçÚuwD
+*),
)
ŒéêºXãáV
Žã³
$
Gº‡º»Õ7²øDAÐçÿ
+*&,
)
áVŽ¸AÐCDžXãÇc”áV
$
pý
áVŽ-ÎxþA@%À
$
B´…¢:°•
-ÎÏçOŒ«¬xó­ŽÏ“ó­Ž
$
ê
`º¨¸éêCDc”áV
+*,
) ?OHTS6
C
+*,
ƒ
„3اÉ
#
ÎQ$XãàáâáVŽ
?1Q3$
?1QD$?1Q<
áâã³CDÇ~Ïl
$
¹ýèy
!!
g
&!
ÛÜÝÞkDáâã³
?65!#$%
àáâáVŽ
>?
x2x 1̇xþ
?65!#$%
àáâáVŽ
>?

2x 1Ìÿ{…¢
IHMc&!@O6V.MT1TQH-QSTT.RN/H1.PTbWT1-TP.RSTkTSPTQSN15
P-SH\QNPTQSN1P-SH\QH.1NV6N-QHkNQTU N1U STkTSPTQSN1P-SH\QNPT.R
?6?!#$% STQS.QSN1P\.P.1P.RMT1./TH1 P61:.C$/+$!$1%(6
)
@àáâáVŽ
F?>

E)
L1Ì~l
!
ŠÂ1Ì
xaA“
$
¸ÎQ
?1Q

F?>E)
LFDÚu>à
áâáVŽ§¨o_ÚuwD
)
Š$
$?1Q<

â㳺ž¬Õ •º»”±X뫬
)
a«Ç$
$
ÛÜÝÞ@;
<
w
3D3
«¬}¿g‡
?65!#$%
ßàáâáVŽáâã³Éʱ>
$
¹ýA§ÉÛ
ÜÝÞàáâáVŽ§¨‡ŠÕ7àÀpDAÐ
nÿ
$
D¸…AF«Ç€É
)
àáâáVŽ‡-θ
±v…¢´9$.^—‡ADyyL¶”‹‡ì
h²ø
)
àáâáVŽ+€Ân@}_…Šá
VdÇc”
$
¸a«Ç$˜T°gÛÜÝÞx
þ8kDáâã³
&*
C
?65!#$%
ßàáâáV
Žàá⣉U1Ì@ƒ³Ò
$
Š$
*)
C1Ì
kDtT@
&
fˆŽ
$
C‹¸{å@
$
]¹[@
ƒÇ~™šŠáâã³
$
›ÝDGº~™šŠáV
ã³
)
¹¸Š”»Ï8pcò´ß“¶·
+*#5*4,
)
ÑÒ³\Ȏý¬ûxþ8
F?>5
àáâáV
Ž
D8:$D8:
{å‡àáâáVŽ…dáV qý
þ——˜
5
àáâ£x
$
]ŠjGö„Šàáâ
áVŽÏÀ¹º…dáV
+*#,
)
§%
$
àáâáV
ŽVÕ

—)
(
éêCD-ÎxÚu§¨—_
pý…ŠL¶ìÑ78¦A
)
-*4%-
t
)4
Öt
$
v
*$&
o

p
!!!!!! ! ! ! ! ! !!!! ! ! ! !
A.Vc)4$:PPWT!$
9N6$ *$&
&*
CkDáâã³ÛÜÝÞ
?65!#$%
ß
àáâáVŽ
>?
1Ìÿ{ß˜©G˜Ç

ß
!
g
)$
~lßgàáâ£1Ì?(~l
$
േ
~lKñàáâáVŽáâ´9‹¸‚ƒ
$
Š‹
¸úŸHUpGº~l
+)$,
$
ÛÜÝÞxþ…
¢Š‹pGºnƒ
)
w‡xþàáâ£2x
 ÑҘ©
$
kDáâã³ÛÜÝÞ
?65!#$%
ß
àáâ£1Ì$%
$
/
$
0ßjM˜gL—A—
$
t
߇xþ$
<5
ÇAß
$
t-߇x
þ$¯
<5
Ã%Š1̖¸A†
)
«Ç3
Ø~£Çƒ„ÛÜÝރ„õ”»ë«¬‡áV
Žã³¦bçÿ)I‡AIJõHR
$
`³Ç
…AFGHƒ„àáâáVŽ¸ÛÜÝޏŒàÀ
²ø$Š‹ƒú‡JÁ²
)

tuvw
#
+ ,!A.6QNPYI$ 9H-ONTV@<$ 31USZT0G$TQNV,8#$%?($Q6STQS.5
QSN1P\.P.1 NSTW^HbWHQ.WPN/.1M\VN1QP+C,,@S.-;NQV3-NU 8-H
E83$ 44*$ (4#%*&,
+ * ,!GW/NS3$DT11TQZT1 CF,@VN1QSTQS.QSN1P\.P.1P+C,,311W >Tk
KT1TQ$444$)) !#&%4,
+ ) ,!KSN1U^NPQHT1 9 3,3-QHkNQH.1 .R\VN1QSTQS.QSN1P\.P.1PW1UTS
PQSTPP-.1UHQH.1P+C,,?ST1UP@VN1Q8-H$44($)#(,
+ & ,!@TNS-T8 >$ GW/NS3$ IVNkTV3C,3-QHkNQH.1 .RQOT?6V?!#$%
MS.W\ STQS.QSN1P\.P.1P.R\.QNQ.!@(%)$/)C6.7/UWSH1M\S.5
Q.\VNPQHP.VNQH.1+C,,@VN1QT\$44#-$!*#4&4,
+ ,!GH/WSNL$?.PNL$8OH/NUN8$TQNV,]3>X5$N?65!#$% STQS.5
QSN1P\.P.1 H1 .NQN-QHkNQTU ^6N^H.QH-N1U ^H.QH-PQSTPPTP+C,,@VN1Q
+ # ,!9OHSH<$ YTJHQ@C$ KSN1U^NPQHT1 93,3-QHkNQH.1 .RQOT\S.5
/.QTS.RQOT?1QVSTQS.QSN1P\.P.1 H1 Q./NQ.NRQTSH1.-WVNQH.1 +HQO
QOTRW1MNV\NQO.MT1 8(%:7#.$/+(5/+C,,9.V@VN1Q9H-S.^T
:1QTSN-Q$444$*!%#4*,
+ % ,!?N1ML9$ 9NL $` FHF<$ TQNV,:UT1QHRH-NQH.1 N1U -ONSN-QTS5
HZNQH.1 .RSTkTSPTQSN1P-SH\QNPTU./NH1 .RQSN1P-SH\QH.1NV6N-QHkT
STQS.QSN1P\.P.1PH1 +OTNQMT1./TP+C,,C:1QTMS@VN1QDH.V$
*$$$&%!##$&,
+ ( ,!
$%,8
,
A1
+8,,*$$,
+ 4 ,!
ªÂ±
$
âC¯
,
ÛÜÝÞv+òó¦GVtc†‡)
+C,,
$%$,-.
$*$$$)!#*$)),
+$,!
âC¯
$
ދ
$
ÃKÄ
,
ÛÜÝÞó@ŒQ…†‡4
+C,,
$%$,-.
$*$)$)(!&#&%,
+,!
âC¯
$
ÃKÄ
$
[.ƒ
,
ÄQ¦pÛÜÝÞ~”ÖÔ
+9,,
23
#
¢r+&·E
$*$*,
+*,!
âC¯
$
ދ
$
ÃKÄ
,
ÛÜÝÞÉrðç—_
+C,,
$%
$,-.
$*$)$)(!&#&(,
+),!

$
 É1
$
ÃÃ
,3D3
:Ú¬Ô[:$
Y7>7+
x§¨
+C,,
93ò+a
$ *$$$))!#,
+&,!
œ5¸
$
´?+
$
âC¯
$
C
,
ÛÜÝÞ
>@,%$
x¤¥¦
Šéꧨ˜©
+C,,
$%$,-.
$ *$)$ )(!*$#)&&#,
+,!GW/NS3$@TNS-T8 >$9-VTN1 G$TQNV,?OT?65!#$% MS.W\ .R
STQS.QSN1P\.P.1PH1 \VN1QP# MT1./H-.SMN1HPNQH.1$Tk.VWQH.1$N1U
WPTNP/.VT-WVNS/NS[TSP+C,,KT1TQH-N$44%$$$#*$,
+#,!
œ¯
$
âC¯
$
´?+
$
C
,
ÛÜÝÞ
?65!#$%
ßàáâáV
Žàáâ£
!>?
1̤¥‡˜©
+C,,
$%$,-.
$
*$)$ )4!#4,
+%,!8TN1 3>.MTSP$ G@TQTS@NWVP,?6?!#$%?($Q6STQS.QSN1P\.P.1P.R
Q./NQ.!B*!#6.7$!1 67!(61)/9HV +C,,KT1./T$*$$$$&)#
((%,
+(,!G..S/1TTR9$ FT.15GV..PQTSZHTVG9$ 8-O+NSQZ8 B$ TQNV,?OT
MT1TQH-N1U /.VT-WVNSUHPPT-QH.1 .RN^P-HPH-N-HU ^H.P61QOTPHPN1U
PHM1NVQSN1PUW-QH.1 H1 =.%C$:#7$7+C,,@VN1Q@O6PH.VDH.-OT/$
44($)##(),
+4,!?ST1UP@VN1Q8-H$444$&#&%*,
+*$,!FH.QT1^TSM8$ ;.SQO B$ 9NSH.15@.V3,9.VT-WVNS^ H.V.M6N1U
STMWVNQH.1 .RN^P-HPH-N1U ^H.P61QOTPHPH1 \VN1QP+C,,@VN1Q@O6PH.V
DH.-OT/$444$)%#)&,
+*,!?N6V.S:D$DWS^HUNMT3$?O./\P.1 3C,<.1QS.V.RN^P-HPH-N-HU
P61QOTPHP+C,,CX7\ D.Q$*$$$$8##),
+**,!B.QO 8$ 9.SMN1QT9$ 8N1-OTZC@,KT1./T5+HUTMT1TT7\STP5
PH.1 \S.RHVH1MH1 =.%C$:#7$7)-%($%1% STkTNVP1T+QNSMTQP.RN^5
P-HPH-N-HU N1U VNSMTV6H/\NHSTU MT1TSTMWVNQH.1 H1 QOTN^HV5V/W5
QN1Q+C,,C+*),!@TNS-T8 >$8QWNSQ5>.MTSP<$G1.79>$TQNV,>N\HU HP.VNQH.1 .R
\VN1Q?6V?!#$% MS.W\ STQS.QSN1P\.P.1 F?>PTbWT1-TPR.S/.VT-W5
VNS/NS[TSPQWUHTP+C,,@VN1QC$444$4!##%,
+*&,!5STQS.QSN1P\.P.1PN1U 9:?XP#
-.1QS.V.RQSN1P\.PHQH.1 N1U H/\N-Q.1 QOTTk.VWQH.1 .R\VN1QMT1TPN1U
MT1./TP+C,,KT1T$*$$)$)#,
+*,!DTMWHSHPQNH1 ?$KSN1U^NPQHT1 93$@WHMU./T1T-O @$TQNV,?OSTT
Q1Q PW^RN/HVHTPPO.+UHRTST1QPQSTPPUHRTST1QPQSTPP5NPP.-HNQTU
\NQTS1 .RT7\STPPH.1 H1 Q.^N--.,<.1PTbT1-TPR.SSTQS.QSN1P\.P.1
-.1QS.VN1U Tk.VWQH.1 H1 \VN1QP+C,,@VN1Q@O6PH.V$*$$$*%#
**,
+*#,!CH1 LG$ DT11TQZT1 CF,8QSW-QWSTN1U -.UH1M\S.\TSQHTP.RD8:$
N/NHZTSTQS.kHSWP5VH[TQSN1P\.P.1+C,,@S.-;NQV3-NU 8-HE83$
4(4$(###*),
+*%,!BW J$ YNP]@$ 9TPPH1MC, T`.15$N/T/^TS.RN1T+/NHZT
STQS.QSN1P\.P.1 RN/HV6+C,,9.VKT1 KT1TQ$44$*&(#&%,
+*(,!CH1 LG$ DT11TQZT1 CF,:1QTMSNQH.1 N1U 1.15SN1U.//WQNQH.1 .R
N\VNP/N/T/^SN1T\S.Q.1 3?@NPTMT1TRSNM/T1Q+HQOH1 QOTD8:
STQS.VT/T1Q.R/NHZT+C,,@VN1Q+*4,!9NSHV.11TQ8$ JTPPVTS8 >,X7QST/TPQSW-QWSNVOTQTS.MT1THQ6
N/.1MQOT/T/^TS.RN/NHZTSTQS.QSN1P\.P.1 RN/HV6+C,,KT1TQ5
H-P$44($$#*&,
+)$,!
ÜÆ
$
ÇÞ­
$
È®¯
$
C
,
LV
?65!#$%
ߏáVŽá
â£1̤¥¦˜©
+C,,
ó$a
$*$$($) !(#&%,
-)4%-
t
)4
Öt
$
v
*$&
o

p
!!!!!! ! ! ! ! ! !!!! ! ! ! !
A.Vc)4$:PPWT!$
9N6$ *$&
*J?RB@J4Q4@ABE-AIRE.KB.@A-/D -RB.P?=D+23)
E?BE@BE-AIK@I@AI.A8%+0(2*)#529))+&%
F:<.1M$ 8:CH15\H1M$ K3]LN15OWH
!
$ B`ELW5bHW$ C:3;KLWN1
!;.).$12 @)%)$1 +.@)%)6A6*B%C.%).*+@C).#$!%(@$(5$!().6$ V-6G$%12 =2.$!().%(%1: &.67).*41$56.7$)*$
V-6G$%12 ,.5$1!$%(@).%)62$!=($%1!6+.<6!-1$!%(T115%)$1 $1 T1:7).*+P61:.C$/+$!$1%(6$ B$1R%1 ))$$$ 8-$1%
+4LIBE-RB,!EPH1MW1HkTSPNV\SH/TS?65!#$% STQS.QSN1P\.P.1 >?$ QOT-.1PTSkTU STkTSPTQSN1P-SH\QNPTU./NH1 .RN^.WQ*#$ ^+NPN/\VHRHTU ^6>?5@<>RS./QOTP61:.C$/+$!$1%(6+OH-O H1UW-TU ^6$$
!
/.V-F
g
N^P-HPH-N-HU !3D3$ H1UH-NQH1MQOTPT
STQS.QSN1P\.P.1PN-QHkNQTU ^6$$
!
/.V-F
g
3D3,?OTN/\VH-.1P+TSTST-.kTSTU N1U -V.1TU$QOT1 PTbWT1-TU N1U N1NV6ZTU ^6STVNQTU
^H.H1R.S/NQH-PP.RQ+NST,I.SQ65Q+.?65!#$% VH[TSTQS.QSN1P\.P.1 >?QSN1P-SH\QH.1NV6N-QHkNQTU +TST.^QNH1TU +HQO OHMO OTQTS.MT1THQ6,
?OTVT1MQO .RQOTPTPTbWT1-TPkNSHTU RS./*&% Q.*## ^\$ N1U +NPSH-O H1 3?N1U O./.V.M6SN1MTU RS./&#,)i Q.4(,4i,?OTPN/T
Q.?6?!#$% VH[TSTQS.QSN1P\.P.1 >?.RMT1./T$ UHRTST1Q-=P5N-QH1MSTMWVNQ.S6TVT/T1QPH1UW-TU ^6PQSTPP-.1UHQH.1PN1U QOTPQNSQH1M
QSN1P-SH\QH.1 PHM1NVP$ -.STP\.1UH1MQ.<33?^.7$ ?3?3^.7-.1PTSkTU PTbWT1-TPN1U P./T.QOTSSTMWVNQ.S6TVT/T1QP,?OT-=P5N-QH1M
STMWVNQ.S6TVT/T1QPH1UW-TU ^6PQSTPP-.1UHQH.1P.RSTkTSPTQSN1P-SH\QNPTQSN1P-SH\QH.1NV6N-QHkNQTU .R?65-.\HNSTQS.QSN1P\.P.1P+TST
PHM1HRH-N1QV6H1-STNPTU QON1 QONQ.R?6?!#$% VH[TSTQS.QSN1P\.P.1 >?.RMT1./T,JOT1 ^TH1MQSN1PVNQTU H1Q.N/H1.N-HUP$ RHRQTT1 PT5
bWT1-TP\STPT1QTU PQ.\ -.U.1 /WQNQH.1$ 1H1TQTT1 PTbWT1-TP\STPT1QTU RSN/TPOHRQ/WQNQH.1$ N1U NVPTbWT1-TP\STPT1QTU -.1PTSkTU PT5
bWT1-Tp8FLKGap/WQNQH.1,IHkT-NQTM.SHTP+TSTHUT1QHRHTU QOS.WMO \O6V.MT1H-N1NV6PHPNRQTSNVHM1/T1QN1NV6PTP.RQOTHSN/H1.N-HU
PTbWT1-TP$ N1U +HQO ?6?!#$% VH[TSTQS.QSN1P\.P.1 >?.RMT1./TONkH1MV.+O./.V.M6$ +OH-O H1UH-NQTU QONQSTkTSPTQSN1P-SH\QNPTQSN15
P-SH\QH.1NV6N-QHkNQTU .R?65!#$% STQS.QSN1P\.P.1P+OH-O H1UW-TU ^63D3ONU 8HM1HRH-N1QV6UHRTST1-TP+HQO ?6?!#$% VH[TSTQS.QSN1P5
\.P.1 >?.RMT1./T,
+T?D V@EFI,!P61:.C$/+$!$1%(6* 3D3* ?65!#$%5VH[TSTQS.QSN1P\.P.1* STkTSPTQSN1P-SH\QNPT!>?* QSN1P-SH\QH.1NV6N-QH5
kNQTU
F@.# )`(abORYR,,()b

xy-0
!
~€
#
-&4%-
t
)4
Öt
$
v
*$&
o

p
!!!!!! ! ! ! ! ! !!!! ! ! ! !
A.Vc)4$:PPWT!$
9N6$ *$&