全 文 ：广 西 植 物 Guihaia　May２０１３,３３(３):３９５－４００　　　　　　　　　　 http://journal．gxzw．gxib．cn　
DOI:１０．３９６９/j．issn．１０００Ｇ３１４２．２０１３．０３．０１９
DoVanTruong,刘晟源,韦毅刚,等．越南苦苣苔科植物四新记录种[J]．广西植物,２０１３,３３(３):３９５－４００
DoVT,LiuSY,WeiYG,etal．FournewlyrecordedGesneraceousspeciesfromVietnam[J]．Guihaia,２０１３,３３(３):３９５－４００
越南苦苣苔科植物四新记录种
DoVanTruong１,刘晟源２,韦毅刚３,温　放３∗
(１．越南国家自然博物馆 维管束植物分类与多样性保育中心,越南 河内;２．广西弄岗国家级自然保护区
管理局,广西 龙州５３２４００;３． 广 西壮族自治区中 国 科 学 院 广西植物研究所,广西 桂林５４１００６)
摘　要:报道了４个苦苣苔科(Gesneriaceae)植物在越南的分布新记录,其中１种为半蒴苣苔属(Hemiboea)的红
苞半蒴苣苔(H．rubribracteata),２种为报春苣苔属(PrimulinaHance)的文采报春苣苔(P．wentsai)和疏花报春
苣苔(P．laxiflora),还有１种为吊石苣苔属(Lysionotus)的桂黔吊石苣苔(L．aeschynanthoides).文中列出了每
个种的标本引证和地理分布情况.
关键词:红苞半蒴苣苔;桂黔吊石苣苔;疏花报春苣苔;文采报春苣苔;新记录;越南
中图分类号:Q９４９．７７８．４　　文献标识码:A　　文章编号:１０００Ｇ３１４２(２０１３)０３Ｇ０３９５Ｇ０６
FournewlyrecordedGesneraceous
speciesfromVietnam
DOVanTruong１,LIUShengＧYuan２,WEIYiＧGang３,WENFang３∗
(１．ClassificationofVascularPlantsandBiodiversityConservationCenter,VietnamNationalMuseumofNature,HaNoi,
Vietnam;２．NonggangNationalNatureReserveAdministrationofGuangxi,Longzhou５３２４００,China;３．GuangxiInstitute
ofBotany,GuangxizhuangAutomousRegionandtheChineseAcademyofSciences,Guilin５４１００６,China)
Abstract:OnespeciesofHemiboea,H．rubribracteatatwospeciesofPrimulina,P．wentsaiandP．laxiflora,and
onespeciesofLysionotusL．aeschynanthoides,arereportedforthefirsttimefromVietnam．ThecitationofspeciＧ
mensandtheirdistributionisgiveninthepresentpaper．
Keywords:Hemiboearubribracteata;Lysionotusaeschynanthoides;Primulinalaxiflora;Primulinawentsai;new
records;Vietnam
　 　 Thegenus,Hemiboea C．B．Clarke,belongsto
GesneriaceaesubfamilyCyrtandroideaetribeDidymoＧ
carpeae．ItsrelativeChineseendemicgenus,MetabriggＧ
sia W．T．Wang (１９８３a),wasmergedwithHemiboea
(Weberetal．,２０１１)．Inaddition,aftertwonewspecies,
H．purpureaYanLiu& W．B．Xu(Xuetal．,２０１０)and
H．angustifoliaF．Wen&Y．G．Wei(Wenetal．,２０１１),
andonevariety,H．folicularisC．B．Clarkevar．retroflＧ
exaYanLiu&Y．S．Huang(Huangetal．,２０１１)were
published,thisgenuscomprisesatleast２８speciesand
sixvarieties．ThisgenusmainlydistributesinSandSW
ChinaandNVietnam (Wangetal．,１９９０;Wangetal．,
１９９８;Li& Wang,２００４;Weietal．,２０１０)．Thereare
about２８speciesofformerChiritaBuch．ＧHam．exD．Don
wererecorded．However,amajorrevisionoftheonce
monotypic,Primulina Hance(１８８３),wasreleasedin
收稿日期:２０１２Ｇ１１Ｇ２３　　修回日期:２０１３Ｇ０１Ｇ２０
基金项目:国家自然科学基金(３１２６００３８);广西自然科学基金(２０１１GXNSFB０１８０５０);广西植物研究所博士课题启动基金(桂植业１１００３);中国
科学院西部之光人才培养计划项目
作者简介:DoVanTruong(１９８４Ｇ),Vietnamese,MasterinBotany,ResearcherinDepartmentofBiology,Vietnam NationalMuseumofNature,
VAST,(EＧmail)dovantruong_bttn＠yahoo．com.
∗通讯作者:温放,博士,副研究员,主要从事植物分类学与相关研究,(EＧmail)wenfang７６０６０８＠１３９．com.
２０１１．ThosespeciesinformersectionGibbosaccusof
ChiritaBuch．ＧHam．exD．Don．,Chiritopsis W．T．Wang
andtwospeciesofWentsaiboeaD．Fang&D．H．Qinwere
revisedtobemembersofPrimulina (Wangetal．,
２０１１;Weberetal．,２０１１)．Thus,somenewrecordsof
originalChiritasect．Gibbosaccusshouldbereportedas
Primulinaspecies．Lastly,thegenus,LysionotusD．Don,
consistsofabout２７species,andisrangingfromNIndiＧ
a,NNepal,NThailand,NVietnamandSChinatoSJaＧ
pan(Li& Wang,２００４;Nongetal．,２０１０)．Beforeour
report,onlyfivespeciesofthisgenuswerereportedin
Vietnam．
DuringourextensiveexpeditiontoVietnaminthe
pasttwoyears,wecolectedsomespecimensofGesneＧ
riaceaespeciesasnewrecordsfromVietnam,whichare
notbementionedinbotanicalorcorrelativemonograph
ofVietnam(Hoetal．,２０００;Nguyenetal．,２０００)．
１．Hemiboearubribracteata　Z．Y．Li&YanLiuin
ActaPhytotax．Sin．４２(６):５３７,fig．１．２００４．(Fig．１)
Specimensexamined:Vietnam:TuyenQuang
Prov．,NaHangDistr．,NaHangNatureReserve,apＧ
proximately２２°２０′４２″N,１０５°２５′２０″E,ca．３２０m．１７．Jun．
２０１２,DoVanTruong & WenFang,VMN_CN２４１
(VMN,IBK)．
Ecology:Growsinthecrevicesonalimestone
hil,undertropicalevergreenbroadleafforestandbuＧ
shes．３２０ma．s．l．
Conservationandexistencestatus:Thisspeciescan
onlybefoundinasunnylimestonehilinNaHang
NatureReserve．However,thesuitablehabitatisbeing
seriouslyinvadedbyainvasionplant,Zebrinapendnla
Schnizl．WeconsideredthatthepopulationofHemiＧ
boearubribracteataisinarecession．
Distribution:SWGuangxi,CHINA,NewtoVietnam．
Discussion:Firstly,thisspecies wasdiscovered
fromJingxiCounty,Guangxi,China,andconsideredas
anendemiconetoGuangxi．Thisareaabutstheborder
withNVietnam．Thus,itisnowonderthatthenew
recordwasdiscoveredandconfirmed．Italsohints
thereisadirectintimateconnectionbetweentheflora
ofSWChinaandNVietnam．
ThepopulationofGuangxi,Chinaislocatedina
largeＧscalelimestonegorge,andalplantsgrowinthe
bottomofthisgorgewithhighhumidityandshade
density．ButthepopulationofNVietnamgrowsinthe
crevicesofthehilsideoflimestonehil withstronger
sunlight,poorsoilandlow humidity．Somesmal
diferencescanbeobservedbetweentwodiferentpopＧ
ulationsfromChinaandVietnam,forexamplecolorof
involucre(vividfuchsiaorpalepurplishＧred),colorof
corola(whiteoryelowishＧwhite),heightofplants(０．４
－１．５morhigher),etc．ThediferenceofhabitatconＧ
ditionsresultthose．
BasedonthebotanicaldiagnosiswhenitwaspubＧ
lished,thisspeciesissimilartoHemiboeacavaleriei
Lévl．(Li& Liu,２００４),buteasilydistinguishedby
thosecharactersshowedinTable１．Thetable１accordＧ
inglyshowsanothersimilarspecies,H．angustifoliaF．
Wen&Y．G．Wei(Wenetal．,２０１１)．
２．Primulinalaxiflora　(W．T．Wang)Y．Z．Wangin
J．Syst．Evol．４９:６１．２０１１≡ChiritalaxifloraW．T．Wangin
Bul．Bot．Res．,Harbin４(１):２１．１９８４．(Fig．２AＧD)
Specimensexamined:Vietnam:HaGiangProv．,
BacMeDistr．,BacMeNatureReserve,approximately
２２°２２′１８″N,１０５°２７′４８″E,ca．１４５．６m．１６．Jun．２０１２,Do
VanTruong& WenFang,VMN_CN２２９(VMN,IBK)．
Ecology:Growsattheentranceofalimestone
caveandcrevicesoflimestonerocknearwaterfal．
１４５．６ma．s．l．
Distribution:NWGuangxi,China,NewtoVietnam．
Conservationandexistencestatus:Thewaterfaland
caveareusedbylocalpeopletoswimandenjoythe
cool．Thuslocalpeople’sactivitiesmighthaveafected
thenormalreproductionofthispopulation．
Discussion:Thetypelocality,LinyunCounty,is
locatedinNW Guangxi．Itisseparatedfrom Vietnam
bysomecountiesofGuangxiandYunnan．However,
wesuccessivelydiscoveredsomedistributionpointsin
diferentCountiesofGuangxiandYunnannearborder
ofChinaandVietnam．Thenewestcolectedrecordin
ChinawasinGaozhouandHuazhouofGuangdongby
aplantenthusiast．ItalsohintsthecloseconnectionbeＧ
tweenfloraofSChinaandNVietnam．UnquestionaＧ
ble,moreandmoredistributionlocalitieswilbefound
inthefutureintwocountries．
Whenthisspecieswaspublished(Wang,１９８４),
６９３ 广　西　植　物　　　 　　　　　　　　　　　　　　３３卷
Fig．１　HemiboearubribracteataZ．Y．Li& YanLiu　A．Habitat,showingtheinvasionplant,ZebrinapendnlaSchnizl;
B．PurplishＧredinvolucre;C．Frontalviewofcyme;D．Sideviewofcorolaandcyme;E．Frontalviewofcorola;F．Capsule．
Wang(１９８４)considereditresembledChiritaswinglei
(Merr．)W．T．Wang(nowPrimulinaswinglei(Merr．)
Mich．Möler&A．Weber),buteasilydistinguishedby
itssmalerleafblades,moreflowersononecyme,
smalerflower,corolanearlytubular．Furthermore,
whenthisspeciesgrowsindarklimestonecaves,they
looksimilartoChiritatenuifolia W．T．Wang (now
Primulinatenuifolia(W．T．Wang)Y．Z．Wang)(Weber
etal．,２０１１)．Table２summarizesthemorphological
diferencesbetweenP．laxifloraandtwosimilarspeＧ
cies．
３．Primulinawentsai　(D．Fang&L．Zeng)Y．Z．
WanginJ．Syst．Evol．４９:６２．２０１１≡ChiritawentsaiD．
Fang&L．ZenginActaPhytotax．Sin．３１(５):４６６．１９９３．
(Fig．２EＧH)
Specimensexamined:Vietnam:LangSonProv．,
Trang DinhDistr．,TriPhuongCommune,approxiＧ
mately２２°３９′２９″N,１０６°０′１０″E,ca．４２４．１m．１６．Jun．
２０１２,DoVanTruong & WenFang,VMN_CN１７３
(VMN,IBK)．
Ecology:Growsinthecrevicesofsunnyvertical
clifsonalimestonehil．TherearealmostnoaccomＧ
panyingplantsnearthoseindividuals．４２４􀆰１６－４６０m
a．s．l．
７９３３期　　　　　　　　　　DoVanTruong等:越南苦苣苔科植物四新记录种
Fig．２　Primulinalaxiflora(W．T．Wang)Y．Z．Wang(AＧD)　A．Habitat;B．Habit;C．Frontalviewofcorola;D．Side
viewofcyme．　P．wentsai (D．Fang&L．Zeng)Y．Z．Wang(EＧH)　E．Habitat;F．Habit;G．Bud;H．Frontalviewofcorola．
　　Distribution:SWGuangxi,China,NewtoVietnam．
Conservationandexistencestatus:Clusteredand
hugeindividualsgrowontheverticallimestoneclifs
fromthreetotwentyorhighermetersaboveground．
Thusgrowth andreproduction ofthis population
shouldnotbeafectedbyartificialinterference．
Discussion:Thisspecieswaspublishedin１９９３
(Fangetal．)．Beforeournewrecordofthisspeciesin
Vietnam,itcanonlybeknowninonelocalityinLongＧ
zhouCounty,Guangxi．ItwasassessedasCriticalyEnＧ
dangered(CRA１a＋C２a(i))basedonredlistcategories
(IUCN,２００１;Weietal．,２０１０)．Thenewlocalityisvery
closetotheborderoftwocountries,butthedistancebeＧ
tweennewandoldlocalitiesisabout５０kilometers．New
localitiesofPrimulinawentsai mightbediscoveredin
thefutureinthiscontiguousarea．Thisspecieslookslike
P．ophiopogoides(D．Fang& W．T．Wang)Y．Z．Wangand
P．spinulosa(D．Fang& W．T．Wang)Y．Z．Wangwithout
flowers,howeveritissoeasilydiferedonbasisofsome
charactersastheTable３belowshow．
８９３ 广　西　植　物　　　 　　　　　　　　　　　　　　３３卷
Table１　MorphologicalcomparisonofHemiboearubribracteata,H．cavalerieiandH．agustifolia
Diferedcharacters H．rubribracteata H．cavaleriei H．agustifolia
Stem
strong,１０—１３mm
indiam．,stuf
slimandsoft,
５—７mmindiam．
strong,５—７mmindiam．
Involucrecolor red greenorbrownishgreen whitishgreenorpaleyelowishgreen
Calyxlobes longer,８—１５mmlonger shorter,５—７mmlong ７．８－８．８mmlong,betweentwoformer
Corola
white,outsideglabrous,
abaxiallip３Ｇlobedtowards
themiddle
Paleyelow,outsidesparely
glandularpubescences,abＧ
axial３Ｇlobed
greyishＧwhiteorwaxyＧwhite,adaxialoutside
glandularＧpuberulent,abaxialoutsideextremeＧ
lyshortpubescenttonearlyglabrous
Table２　MorphologicalcomparisonofP．laxiflora,P．swingleiandP．tenuifolia
Differedcharacters P．laxiflora P．swinglei P．tenuifolia
Leafblade ６—９．６cmlong １３—１７cmlong ２．２—５cmlong
Flowersononecyme ７—１２ １—６ １
Flowersize ca．１．７cmlong ２．８—４．２cmlong ca．２．８cmlong
Corola nearlytubular tubular slenderfunnelform
Table３　MorphologicalcomparisonofP．wentsaii,P．ophiopogoidesandP．spinulosa
Differedcharacters P．wentsaii P．ophiopogoides P．spinulosa
Flowersnumber
ononecyme
２—３ ５—７ ca．９ormore
Bracts ３０—３５mmlong ９—１５mmlong ２—２．５mmlong
Calyxlobes １０—１２mmlong ４—５mmlong ca．１．５mmlong
Corola ４．５—５cmlong;bluishＧpurple;
outsidesparselyglandularＧpubescence
２．５—２．６cmlong;palepurple;
outsidewithoutglandularＧpubescence
ca．１．３cmlong;purple;upperＧoutside
sparselyextremelypubescence
Corolatube campaniform,３—３．５cmlong sliminfundibuliform,ca．１．４cmlong nearlytubular
Staminodes ３,lateralones９mmlong ２,ca．５mmlong ２,ca．１．５mmlong
Pistil ca．２．６cmlong ca．１．５cmlong ca．１．４cmlong
　　４．LysionotusaeschynanthoidesW．T．Wang　in
Guihaia３(４):２６５,pl．４,fig．７Ｇ９．１９８３etFl．Reip．Pop．
Sin．６９:５３６,tab．１５３:７Ｇ９．１９９０;W．T．Wangetal．InZ．Y．
WuetRaven,Fl．China１８:３８８．１９９８．
Specimensexamined:Vietnam:BacKanProv．,Na
RyDistr．,Kim Hy Nature Reserve,approximately
２２°１１′２４″N,１０６°３′４３″E,ca．４５７．１m．１０．Jun．２０１２,Do
VanTruong & WenFang,VMN_CN８３(VMN,IBK)．
Ecology:GrowsonthetrunksofevergreenbroadＧ
leavedtreesorcrevicesoflimestonerocksundereverＧ
greenbroadＧleavedtreesandbushes．４５７．１ma．s．l．
Distribution:Middle,SW Guangxi,SEYunnan,
SWGuizhou,China,NewtoVietnam．
Conservationandexistencestatus:Thepopulations
inornearKim HyNatureReservesurviveingood
state．Therearealotofyoungplantsreproduceby
seeds(sexualreproduction)andrhizomes(asexualreＧ
production)．Theindividualsofpopulationsgrowand
renewverywel．
Discussion:Atfirstthisspecieswasconsideredas
endemiconetoChina(SW Yunnan,MiddleandSW
Guangxi,andSW Guizhou)．Thenewrecordofthe
speciesinVietnamhintsthecloseconnectionbetween
floraofSandSWChinaandNVietnam．Sofar,four
speciesofLysionotusD．Donwererecorded,namelyL．
pauciflorusMaxim．,L．peteloti Pel．,L．serratusD．
Don(Ho,２０００)andthisnewrecordedspecies．This
speciesbelongstoSect．Lysionotusser．HeteroPhylli
W．T．Wang,buteasilydistinguishedotheronesofthis
sectionbyitsanisopleuralleafblades,entireleafblade
margin,smalerflowers,yelow corola,infundibuliＧ
formＧtubularcorolatube．Furthermore,thescientific
namehintsthatthemorphologyofvegetableorganof
thisspeciesissimilartoAeschynanthusJackplants
(Wang,１９８３b)．
Acknowledgments　Theauthors wouldliketo
９９３３期　　　　　　　　　　DoVanTruong等:越南苦苣苔科植物四新记录种
thankstafattheherbariumofVietnamNationalMuＧ
seumofNature,Vietnam(VMN),herbariumofInstiＧ
tuteofEcologyandBiologyResource,Vietnam(HN),
theherbariumoftheInstituteofBotanyGuangxi,ChiＧ
na(IBK);theherbariumoftheSouthChinaBotanical
Garden,ChineseAcademyofSciences,China(IBSC);
theherbariumoftheKunmingInstituteofBotany,
ChineseAcademyofSciences,China(KUN)．Wealso
thankMr．WeiＧJunWuforprovidingcolectedinformaＧ
tionofPrimulinalaxiflorainGuangdong,China,and
Mr．XinＧXinZhuforprovidingcolectedinformationof
Lysionotusaeschynanthoides．
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苔)[J]．ActaPhytotaxSin(植物分类学报),４２(６):５３７－５４０
NguyenTB,VuVC,VuVD,etal．２０００．VietnamForestTrees[M]．
Hanoi:AgriculturePublishingHouse,４６０(inVietnamese)
NongDX,XuWB,WuWH,etal．２０１０．Lysionotusfengshanensis
YanLiu&D．X．Nongsp．Nov．(Gesneriaceae)fromGuangxi,ChiＧ
na[J]．NordJBot,２８:７２０－７２２
WangWT,PanKY,LiZY．１９９８．ChiritaBuch．ＧHam．ex．D．Don
[M]//WuZY,RavenPH．FloraofChina．Vol．１８．Beijing:Science
Press;St．Louis:MissouriBotanicalGardenPress:３１１－３４５
WangYZ,MaoRB,LiuY,etal．２０１１．Phylogeneticreconstruction
ofChiritaandalies(Gesneriaceae)withtaxonomictreatments
[J]．JSystEvol,４９(１):５０－６４
WeberA,MiddletonDJ,ForrestA,etal．２０１１．MolecularsystemＧ
aticsandremodelingofChiritaandassociatedgenera(GesneriＧ
aceae)[J]．Taxon,６０(３):７６７－７９０
WeberA,WeiYG,SontagS,etal．２０１１．InclusionofMetabriggＧ
siaintoHemiboea(Gesneriaceae)[J]．Phytotaxa,２３:３７－４
WenF,Tang WX,WeiYG．２０１１．Hemiboeaangustifolia(GesneriＧ
aceae),anewspeciesendemictoatropicallimestoneareaof
Guangxi,China[J]．Phytotax,３０:５３－５９
XuWB,WuWH,NongDX,etal．２０１０．Hemiboeapurpureasp．
nov．(Gesneriaceae)fromalimestoneareainGuangxi,China[J]．
NordJBot,２８:３１３－３１５
􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓􀤓
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　一新记录种———中越双唇兰)[J]．Guihaia(广西植物),３１(５):
５８９－５９１
HuangYS(黄俞淞),LiuSY(刘晟源),PengRC(彭日成),etal．
２０１２．AnewlyrecordedspeciesofPhaius(Orchidaceae)from
China(中国鹤顶兰属(兰科)一新记录种)[J]．Guihaia(广西植
物),３２(２):１４３－１４５
JiangRH(蒋日红),WuWH(吴望辉),NongDX(农东新),etal．
２０１１．SciaphilastellataAver,anewlyrecordedspeciesofTriuriＧ
daceaefromlimestoneareainChina(中国石灰岩地区霉草科一新
记录种———星状霉草)[J]．Guihaia(广西植物),３１(１):９－１０
LittleDP,SchwarzbachAE,AdamsFP,etal．２００４．ThecircumＧ
scriptionandphylogeneticrelationshipsofCalitropsisandthe
newlydescribedgenusXanthocyparis(Cupressaceae)[J]．AmJ
Bot,９１(１１):１８７２－１７７１
MaoKS,HaoG,LiuJQ,etal．２０１０．DiversificationandbiogeograＧ
phyofJuniperus(Cupressaceae):variablediversificationrates
andmultipleintercontinentaldispersals[J]．New Phytol,１８８
(１):２５４－２７２
MilRR,FarjonA．２００６．ProposaltoconservethenameXanthoＧ
cyparisagainstCallitropsisOerst．(Cupressaceae)[J]．Taxon,
５５(１):２２９－２３１
NongDX(农东新),WuWH(吴望辉),JiangRH(蒋日红),etal．
２０１１．NotesonCalocedrus(Cupressaceae)inGuangxi,China(广
西翠柏属(柏科)植物小志)[J]．Guihaia(广西植物),３１(２):
１５５－１５９
TamuraK,PetersonD,PetersonN,etal．２０１１．MEGA５:molecular
evolutionarygeneticsanalysisusingmaximumlikelihood,evoluＧ
tionarydistance,andmaximumparsimonymethods[J]．MolBiol
Evol,２８(１０):２７３１－２７３９
XiangQP,FarjonA．２００３．CuticlemorphologyofanewlydiscovＧ
eredconifer,Xanthocyparisvietnamensis(Cupressaceae),anda
comparisonwithsomeofitsnearestrelatives[J]．BotJLinn
Soc,１４３:３１５－３２２
XuWB(许为斌),LiangYY(梁永延),ZhangXC(张宪春),etal．
２００８．CaobangiaA．R．Smith&X．C．Zhang(polypodiaceae),anewly
recordedferngenusfromChina(中国水龙骨科一新记录属———
高平蕨属)[J]．JSystEvol(植物分类学报),４６(６):９１６－９１８
XuWB(许为斌),HuangYS(黄俞淞),YeXX(叶晓霞),etal．２０１０．
AspidistraconnataH．J．Tilich,anewlyrecordedspeciesofAspiＧ
distra(Ruscaceae)fromChina(中国蜘蛛抱蛋属一新记录种———
合瓣蜘蛛抱蛋)[J]．Guihaia(广西植物),３０(５):６１３－６１５
００４ 广　西　植　物　　　 　　　　　　　　　　　　　　３３卷