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Corydalis hegangensis, a new species of Fumariaceae from Heilongjiang

鹤岗延胡索, 黑龙江紫堇科一新种



全 文 :  Guihaia  Jun. 2016ꎬ 36 (增刊 1): 233-235
http: / / journal.gxzw.gxib.cn
http: / / www.guihaia-journal.com
【原文刊载】“鹤岗延胡索ꎬ 黑龙江紫堇科一新种” 已于 “中国知网 (CNKI)” 单篇优先数字出版ꎬ «广西植物»待刊ꎮ
鹤岗延胡索ꎬ 黑龙江紫堇科一新种
王文采
( 中国科学院植物研究所 系统与进化植物学国家重点实验室ꎬ 北京 100093 )
摘  要: 描写了自黑龙江省发现的紫堇科紫堇属一新种ꎬ鹤岗延胡索ꎮ 此种与新疆元胡相似ꎬ但本种叶的小
叶多在顶端具 1~5牙齿ꎬ花序苞片有 3~5小裂片ꎬ上花瓣及其辦片和距均较小ꎬ蜜腺呈狭卵球形ꎬ弯曲ꎬ顶端
尖锐而与新疆元胡相区别ꎮ
关键词: 紫堇科ꎬ 紫堇属ꎬ 新种ꎬ 黑龙江
中图分类号: Q949    文献标识码: A    文章编号:1000 ̄3142(2016)增刊 1 ̄0233 ̄03
Corydalis hegangensisꎬ a new species of
Fumariaceae from Heilongjiang
WANG Wen ̄Tsai
( State Key Laboratory of Systematic and Evolutionary Botanyꎬ Institute of Botanyꎬ Chinese Academy of Sciencesꎬ Beijing 100093ꎬ China )
Abstract:A species of the genus Corydalis (Fumariaceae)ꎬ C. hegangensisꎬ is described as new from Heilongjiang
Province. It is similar to C. glaucescens Regelꎬ differing in its usually apically 1 ̄5 ̄dentate leafletsꎬ 3 ̄5 ̄lobutale bractsꎬ
smaller upper petal with shorter limb and shorter spurꎬ and narrowly and curvedly ovoid nectary.
Key words: Fumariaceaeꎬ Corydalisꎬ new speciesꎬ Heilongjiang
    鹤岗延胡索  图 1
Corydalis hegangensis W. T. Wangꎬ sp. nov.
Fig. 1. (sect. Corydalis)
Ob plantam tubere globoso praeditamꎬ caulem supra
basin folio squamiformi unico praeditumꎬ folia biternataꎬ
nectarem brevisissimum 1-1.5 mm longum species nova
haec est similis C. glaucescenti Regelꎬ a qua foliolis
plerumque apice 1 ̄5 ̄dentatisꎬ racemi bracteis 3 ̄5 ̄lobula ̄
tisꎬ petalo supero minore 18 mm longo cum limbo 11 mm
longo et calcari 7 mm longoꎬ nectare anguste et curvate
ovoideo apice acuto differt. In C. glaucescenteꎬ foliola
plerumque 3 ̄fida vel 3 ̄partitaꎬ racemi bracteae indivisaeꎬ
petalum superum majus 20-50 mm longum cum limbo
10-11 mm longo et calcari 10-14 mm longoꎬ et nectar fil ̄
ioforme rectum apice obtusum est.
Herbs prennialꎬ glabrousꎬ tuber ̄bearing.Tuber glo ̄
boseꎬ ca. 1.8 cm in diam. Stem ca. 12 cm tallꎬ below
scale ̄like leaf downwards thinningꎬ with one scale ̄like
leaf and three true leavesꎬ from axil of the scale ̄like
leaf with one flowering branch ca. 8 cm long. Scale ̄like
leaf paperyꎬ narrowly ovateꎬ ca. 2 × 0. 85 cmꎬ apex
slightly acuteꎬ margin entire. Leaves bi ̄bernateꎬ petio ̄
lateꎻ blade broadly rhombic in outlineꎬ 3-6 × 3.2-4.5
cmꎻ leaflets long or shortly petiolulateꎬ rarely sessileꎬ
obovate or cuneate ̄obovateꎬ rarely ellipticꎬ 0.64-2.4 ×
0.25-1.3 cmꎬ apex subtruncate and 1-5 ̄dentateꎬ rarely
收稿日期: 2016 ̄06 ̄25    修回日期: 2016 ̄07 ̄28
作者简介:王文采(1926 ̄)ꎬ男ꎬ研究员ꎬ中国科学院院士ꎬ著名植物分类学家ꎬ长期从事有花植物分类学研究ꎮ
图 1  鹤岗延胡索  A. 植株全形ꎻ B. 苞片ꎻ C. 花ꎻ D. 上花瓣辦片ꎻ E. 下花瓣ꎻ F. 内花瓣ꎻ G. 雄蕊束ꎻ H. 蜜腺ꎻ I. 雌蕊ꎮ
Fig. 1  Corydalis hegangensis  A. Habitꎻ B. Bractꎻ C. Flowerꎻ D. Limb of upper petalꎻ E. Lower petalꎻ
F. Inner petalsꎻ G. Stamen bundleꎻ H. Nectaryꎻ I. Pistil. (from holotype)
acute and entireꎬ base cuneateꎬ with petiolules 0.1-1.8
cm longꎻ petioles 1. 2 - 2 cm long. Racemes terminalꎬ
that of stem 3.5 cm longꎬ densely 8 ̄floweredꎬ that of
branch 1 cm longꎬ densely 4 ̄floweredꎻ bracts cuneate ̄
obovate or broadly rhombicꎬ 6-11 × 3-4.5 mmꎬ apex
with 3-5 narrowly triangular or linear lobules unequal in
sizeꎻ pedicels (2 ̄)4-7 mm long. Flower: Sepals cadu ̄
cousꎬ unknown. Petals purple (?)ꎬ not crestedꎻ upper
petal ca. 18 mm longꎬ with limb obtrapeziformꎬ 11 × 5
mmꎬ at apex subtruncate and retuseꎬ and spur tubularꎬ
ca. 7 mm longꎬ 1. 8 - 2 mm in diam.ꎬ straightꎻ lower
petal similar to upper petal both in shape and sizeꎬ
above base slightly saccateꎻ inner petals ca. 11 mm
longꎬ with limbs coherent apicallyꎬ narrowly elliptic ̄ob ̄
432 广  西  植  物                                      36卷
ovateꎬ 6 × 2. 2 mmꎬ and claws slenderꎬ ca. 5mm
long. Stamens with filaments thinly membranousꎬ lance ̄
olate ̄linear. ca. 8 × 0. 8 mmꎬ longitudinally 1 ̄nerved
and anthers yellowishꎬ narrowly oblongꎬ 0.6 - 0.8 mm
long. Nectary blackꎬ narrowly ovoidꎬ ca. 1.2 mm longꎬ
slightly curvedꎬ at apex acute. Pistil 8-9.5 mm longꎻ o ̄
vary greenꎬ linearꎬ 5-5.5 × 0.5-0.7 mmꎻ style ca. 2.5
mm longꎻ stigma orbicularꎬ ca. 1.2 mm in diam.ꎬ with 8
dense yellowish subglobose tubercles. Capsule
unknown.
黑龙江 (Heilongjiang): 鹤岗市ꎬ鹤北 (Hegang
Shiꎬ Hebei)ꎬ鹤北林业局ꎬ北影场ꎬ河岸草地 (Hebei
Forestry Bureauꎬ Beiyingchangꎬ in grassy place on
river bank)ꎬ 1995 ̄07 ̄15ꎬ东北林学院植物调查队
(Pl. Exped. from N. ̄E Forest. Inst.) 9531023 (holo ̄
typeꎬ PE).
本种的植株具球形块茎ꎬ茎基部之上具一鳞片
状叶ꎬ蜜腺极短ꎬ长在 1 ~ 1.5 mm 之间ꎬ因此与也具
上述形态特征的新疆元胡(Corydalis glaucescens Re ̄
gel)相似ꎬ与后者的区别在于本种叶的小叶多在顶
端具 1~5枚牙齿ꎬ花序苞片顶端有 3 ~ 5 枚小裂片ꎬ
上花瓣较小ꎬ长 18 mmꎬ其辦片长 11 mmꎬ其距长 7
mmꎬ蜜腺狭卵球形ꎬ弯曲ꎬ顶端尖锐ꎮ 在新疆元胡ꎬ
叶的小叶多为三中裂或三深裂ꎬ花序苞片不分裂ꎬ上
花瓣较大ꎬ长 20~25 mmꎬ其辦片长 10~11 mmꎬ其距
长 10~14 mmꎬ蜜腺呈丝形ꎬ直ꎬ顶端钝(吴征镒等ꎬ
1999)ꎮ
致谢  深切感谢孙英宝先生为本文绘图ꎮ
参考文献:
WU CYꎬCHUANG HꎬSU ZYꎬ 1999. Corydalis [M] / / Flora Reipu ̄
blicae Popularis Sinicae. Beijing: Science Press. 32: 96 -
481. [吴征镒ꎬ庄璇ꎬ苏志云ꎬ 1999. Corydalis [M] / /中国植
物志. 北京:科学出版社ꎬ32: 96-481.]
􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠􀤠
( 上接第 173页 Continue from page 173 )
laresꎬ0.6-1 mm longaeꎬglabraeꎬalbidaeꎬmajores superne
brunneolaeꎬvel viridesꎬciliolatae. Flos pistillatus breviter
crasseque pedicellatus: tepala 5ꎬleviter inaequaliaꎬsubae ̄
quilongaꎬovata vel anguste ovataꎬca. 0.38 mm longaꎬbasi
connataꎬ dorso leviter sub apice 1-4 ̄pileataꎬ pileolis 0.1
-0.25 mm longis subhyalinisꎻ pistillum ca. 0􀆰 3 mm lon ̄
gumꎬ ovario ovoideo ca. 0.26 mm longoꎬ stigmate minuto
breviter penicillato.
Habitu Pellionia cephaloidea W. T. Wang est
simillima P. scabrae Benth.(sect. Pellionia)ꎬquae inflo ̄
rescentiis pistillatis dense cymosis receptaculis et
involucris carentibusꎬ floris pistillati tepalis valde inae ̄
qualibus 2-3 majoribus navicularibus dorso leviter sub
apice longe corniculatis ceteris minoribus planis ellipticis
haud corniculatis(Fig. 1: H) praeclare distinguitur. Spe ̄
cies duae hae fortasse ex eodem generis auctore exstincto
oriundae fuerunt.
Additional specimen examined. Hunan: Heng
ShanꎬYubanqiao (玉板桥)ꎬ alt. 350 mꎬ 1996 ̄03 ̄14ꎬ
J. L. Xia(夏江林) 59(PE).
致谢  蒙孙英宝先生为本文绘图ꎬ并协助浸泡
花序等工作ꎬ作者谨表示衷心感谢ꎮ
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段林东ꎬ李明红ꎬ 2002. 蔓赤车的一个新异名 [J]. 植物分类
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ZYꎬRAVEN PH (eds). Flora of China. Beijing:Science Pressꎻ
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532增刊 1                        王文采: 鹤岗延胡索ꎬ 黑龙江紫堇科一新种