The Villin/Gelsolin/Fragmin Superfamily Proteins in Plants An Arabidopsis Class II Formin, AtFH19, Nucleates Actin Assembly, Binds to the Barbed End of Actin Filaments and Antagonizes the Effect of AtFH1 on Actin Dynamics Composition of Microfilament Skeleton and Its Regulatory Effects on Polar Growth of Pollen Tubes