Morphogenesis of Inflorescence and Flower in Ostrya (Betulaceae) Sequences of low-copy nuclear gene support the monophyly of Ostrya and paraphyly of Carpinus (Betulaceae) Chromosome Numbers of Two Threatened Species of Betulaceae Development of Ovules and Embryo sacs in Ostrya virginiana (Betulaceae) and Its Systematic Significance