全 文 :书!"#$%&
,2016,36(7):1366-1375
犃犮狋犪犅狅狋.犅狅狉犲犪犾.犗犮犮犻犱犲狀狋.犛犻狀.
!"#$:10004025(2016)07136610 犱狅犻:10.7606/j.issn.10004025.2016.07.1366
%&(
:20160518;)*&%+(:20160627
,-./
:
()*+%,-
(31370237);./0)*+%,-(152300410176);."0)*+%,-(C2016111016);./012%3
45+678
(15B180007)
0123
:
9:;
(1969-),<,=>,?@A,6BCDEFG#$%。Email:jzbotany@163.com
HIJK:LMN,@A,=>OPQ,RSTUFG#$%6B。Email:zhaojiancheng@hebtu.edu.cn
,456789:;<=>?
@ABCDEFGHI
!"#
1,2,
$%&
2,3,
(2,)*+2
(1VJ12QWX+%3 O$Y,./VJ454001;2."QWZ% O[+%%\,](^050024;3_`%\ O[+%%
,
."_`053000)
J K:EabcdGefgG+、hiG+jYklY,mncdG+jYkopqrcdG+setu,v
6BwrsxjcdG+#$qylez4000{|}~E,j
%6B,r35、40
|j4DNA(狉狆狊4、狋狉狀G、狋狉狀L狋狉狀F、犪狋狆B狉犫犮L)sYks,(NJ)qZ
*
(ML)Ms。 ¡¢£:(1)¤sr,cdGefgG+#$s¥¦¤§¨1©ª«j¬s©
®
,¯
、¯
°2
%±²³¨´Zµ¶
,
cdGe·T¸¹jgG+sº
。(2)¤sr,»*cdGef
hiG+#$¦¤¼s©r
,
½¾
%j¿ÀÁÂ
,
ÃK¬·ÄE¼ÅYez
。(3)cdG+Ƽ)*e
z
,
cdG+ÇÈÉʤËÌÍjYklY
,
cdG+jRSÎ¥±²E
:
#$ÏÐÑ
,
ÒÓsÔ
;¯
q¯°E
ÕÖGÑ
(犘狅犺犾犻犪like),×Ø
ÙÚÛÜ
,r®Ý°ÞÚ,ßÂà
áâã
;OäåæOçOÔè;édE
êOëdì
,
Ó¨¬íîjïðá¼ìédcñ
,
òëìédcñ
。(4)rcdG+óô¨5È,õcdGÈ
(犕犻犲犾犻犮犺犺狅犳犲狉犻犪)、ÕÖGÈ(犘狅犺犾犻犪)、öÕÖGÈ(犘狊犲狌犱狅狆狅犺犾犻犪)、dGÈ(犛狔狀狋犺犲狋狅犱狅狀狋犻狌犿)qЯGÈ
(犈狆犻狆狋犲狉狔犵犻狌犿),8÷Eø¿ù34(ôúseÅo)。
LMN
:
cdG+
;
gG+
;
hiG+
;
ÁÂ
;
s
;
se
@O;P$
:Q949.35 !QRST:A
犘犺狔犾狅犵犲狀狔犪狀犱犆犾犪狊狊犻犳犻犮犪狋犻狅狀狅犳狋犺犲犕狅狊狊犉犪犿犻犾狔犕犻犲犾犻犮犺犺狅犳犲狉犻犪犮犲犪犲
犻狀犆犺犻狀犪犐狀犳犲狉狉犲犱犳狉狅犿犆犺犾狅狉狅狆犾犪狊狋犇犖犃犛犲狇狌犲狀犮犲狊犪狀犱犕狅狉狆犺狅犾狅犵狔
LIUYongying1,2,NIUYulu2,3,CAOZhen2,ZHAOJiancheng2
(1DepartmentofBiology,JiaozuoTeachersColege,Jiaozuo,Henan454001,China;2ColegeofLifeScience,HebeiNormal
University,Shijiazhuang050024,China;3ColegeofLifeScience,HengshuiUniversity,Hengshui,Hebei053000,China)
犃犫狊狋狉犪犮狋:Thepaperaimstodiscusstherelationshipbetween犕犻犲犾犻犮犺犺狅犳犲狉犻犪groupandBryaceae,toidenti
fymorphologicalsynapomorphybetween犕犻犲犾犻犮犺犺狅犳犲狉犻犪groupandMniaceae,tosolvetheproblemofsys
tematicpositionof犕犻犲犾犻犮犺犺狅犳犲狉犻犪group,tofigureouttheclassificationofMielichhoferiaceaeinChina.
Morphologicalcharactersofmorethan4000specimensofMielichhoferiaceaeanditsrelatedgroupsinChi
nawereanalyzed.Andthen,neighborjoining(NJ)andmaximumlikelihood(ML)treewerebuiltbased
oncombined狋狉狀G,狉狆狊4,狋狉狀L狋狉狀F,and犪狋狆B狉犫犮LcpDNAsequencesof40specimensfor35species.
Combiningmorphologicalandmoleculardataintheanalysesshowedthat:(1)notonlywere犕犻犲犾犻犮犺犺狅犳犲
riagroupandBryaceaefamilyinthedifferentcladesinthemoleculartree,butalsomorphologicalfeatures,
suchastheleafshape,theleafcelsofthetwogroupsweredifferent.犕犻犲犾犻犮犺犺狅犳犲狉犻犪groupshouldbesep
aratedfromBryaceae.(2)Nomorphologicalsynapomorphycanbedetectedfor犕犻犲犾犻犮犺犺狅犳犲狉犻犪groupand
Mniaceaefamily,althoughtheywereinthesamecladeinthemoleculartree.Thetwogroupswerenot
monophyletic.(3)Mielichhoferiaceaeisanaturalmonophyleticgroup.InMielichhoferiaceae,theplants
aresmal,stemstendtobemuchbranchedthaninBryaceae;leavesare犘狅犺犾犻犪likeinshapeandareola
tion;gametangiaareoftenproducedonlateralbranchesorthebaseofstems;peristomeisdiplolepideous
alternate,oftenreducedtoonelayerorrarelyabsent.(4)Therearefivegenera,犕犻犲犾犻犮犺犺狅犳犲狉犻犪,犘狅犺犾犻犪,
犘狊犲狌犱狅狆狅犺犾犻犪,犛狔狀狋犺犲狋狅犱狅狀狋犻狌犿and犈狆犻狆狋犲狉狔犵犻狌犿,inMielichhoferiaceaeinChina,total32speciesand2
variaties.
犓犲狔狑狅狉犱狊:Mielichhoferiaceae;Bryaceae;Mniaceae;morphologicalcharacters;moleculartree;classification
cdG+(Mielichhoferiaceae)ûÈGü
(Musci),gGýü(Bryidae),gG8(Bryales)。þ
ÿ!"o%Kjkù
,
v+¿óô#6È290
(
ôú2$
)[12]。
vezjZæsx"%
&
,
Æ(
、
)*
、
+,-./01Gez2rj4
S3NNs
,
§¨4SjO%J
。
cdGÈ
(犕犻犲犾犻犮犺犺狅犳犲狉犻犪)#$4ÓOÚô561j,7
®
,
89E
“
:5Ge
”;
cdGÈqÕÖGÈ
(犘狅犺犾犻犪)j;æ:O<Á,7®,4=,>?#
$@AB,7jCDNsqÁ7§¨4SjEF
J
。
cdG+ÆFG#$rjGHezI¼
,
se
%6BJK´ELM
。
ÚNwO
,
cdG+jYk
opqseYkʤPZsQ
(
¢1)。RS
Funch 1818 TUVW8÷óXvÈj
犕犻犲犾犻犮犺犺狅犳犲狉犻犪犿犻犲犾犻犮犺犺狅犳犲狉犻犪狀犪,½YZ8pÐ
]GÈ
(犠犲犻狊狊犻犪)[[G+(Pottiaceae)]r[16]。Nees
2wÅìédqOçOÔèjOäåE±²MWcdGÈ
(犕犻犲犾犻犮犺犺狅犳犲狉犻犪)[17]。Bruch2]Ec
dGÈjÅìédfÕÖGÈjÇd짨À
Á
,
4^_`gG+
(Bryaceae)r[18]。Schimperw
犕犻犲犾犻犮犺犺狅犳犲狉犻犪EabÈM\WcdG+(Mieli
chhoferiaceae),YZcô1È[19]。dOj;æT
r
,
Zæse%(
[68,1213,2022]
efBrotherus[4]j
g5
,
hªgG+Ƽisjk^elæje
z
,
§¨ménÝ£o
、
édpqrsáïð2m
ϱ²q#tÐÑ
、¯
×Ø
、¯
°Ú
2uÏ
±²j¼?ez
:
cdGÈ
、
ÕÖGÈ
、
ЯGÈ
(犈狆犻狆狋犲狉狔犵犻狌犿)、öÕÖGÈ(犘狊犲狌犱狅狆狅犺犾犻犪)、
dGÈ
(犛狔狀狋犺犲狋狅犱狅狀狋犻狌犿)qvwGÈ(çöx,
犛犮犺犻狕狔犿犲狀犻狌犿)yy8p¸¹gG+jcdGý
+
(Mielichhoferioideae)、gGý+(Bryoideae)áÕ
ÖGý+
(Pohlioideae)r。
z10{TO,fË{|@¸¹gG+se%6
Bj}`
,¨
lcdG+
、
gG+qhiG+
(Mni
aceae)jse%~qYklYºÿWPæ。
,°q¯ÏDNA
6B ¡oFºÕ
ÖGÈ
、
cdGÈqЯGÈ2ezMgG+
,
^fhiG+jlYoÌÍ
[23]。
>?g5dO
³8Oæj ¡
[2426],
¼?
%Kj©ª
。
Æ
,
,
%qsj
þs6B
,
>?çg58Pæ%Káæá(
ÿ!GeseYkr
,
Crosby2
ÕÖGÈ_ÙhiG+
,
cdGÈ2pcdG
+
[1];Hil2[11]Koponen[27]lhiG+js
e%g5
,
Goffinet2[28]jóô¨cdGÈ
jZ+
———
hiG+s
,
sºWcdG+
;
Frey2[14]@Gofinet2jGeseYk[28]¤+$w
úÅoW
,
wÕÖGÈ
、
öÕÖGÈ
、
cdG
È
、
dGÈqvwGÈM\WcdGý+
,
ЯG
È8phiGý+
(Mnioideae)r。
lrcdG+j6B
,
[6]
¤
《
rG
eÈ
》(
®¡
)
rhªWBrotherus[4]jg5,c
dGÈr¢¼j
———
r£cdG
(犕犻犲犾犻犮犺犺狅犳犲
狉犻犪狊犻狀犲狀狊犻狊)pgG+cdGý+Ir,fdº¤
jr(CFG
[2935]
-rFG#$6Bj4
S¥¦
《
rFG
》(
§4¨)[12]q;¥¤(犕狅狊狊
犉犾狅狉犪狅犳犆犺犻狀犪)[13]©ªjGeseY
k
[6],
cdGezpgG+r
,
«¬vez¤
r¨4È33(ôúseÅo)。NÉ,ÕÖG
Èç[xW3:¯ÕÖG(犘狅犺犾犻犪犿犪犮狉狅犮犪狉
狆犪)[36]、£dÕÖG(犘.犺狔犪犾狅狆犲狉犻狊狋狅犿犪)[36]q®¯
ÕÖG
(犘.犺犻狊犪犲)[37];z,9:;2[xWdG
È1ç———°±dG(犛狔狀狋犺犲狋狅犱狅狀狋犻狌犿犽狌狀
犾狌狀犲狀狊犲),vÈÆrj²³«¬[38]。þ®V,c
dGez¤r´«¬#¨5È34(ôúse
Åo
)。
µ¶·2
[39]
,¸ÏDNAITS
j
Ykpqçs]E
,
ÕÖGÈqcdGÈ2
76317N 9:;,2:,
%±²qsjrcdG+¹º6B
Ƽ»\hiG+I¼jez
,
½ Hil2[11]
M\cdG+jg5
。
9:;2
[4041]
H¾¸k6
Bvez}~
,
¿ÀW¼?(mçsxáÁW¼
?j[x
。《
FG#$
,
rO$$x¬
》[15]
r
ÂÃFrey2[14]Ge#$seYkcdG2ez
phiG+r
,
zº¤j
《
ÄÅFG#$
》[42]
r³=Yk
。
~¥Æ¤H¾,
%±²qsj6
B
,
abcdGefgG+jlY-fhiG+¨
¾¿ÀÁÂ
,
^ÇvezjYkopqÈÉr
cdG+jsetu
,
Ê@vezse%j]Î
,
³
@Ge#$gG8jYkse6B§¨4S˹
。
1 qC
1.1 U V
6BrsxjcdG+#$qylez
j}~4000{|。40|,¿35
sO$%
,
rhiG+7,ÕÖGÈ
14,ÌÍGÈ(犅狉犪犮犺狔犿犲狀犻狌犿)7,g
GÈ7,ÎGÈ(犃狀狅犿狅犫狉狔狌犿)、+éGÈ
(犘犾犪犵犻狅犫狉狔狌犿)、Z¯GÈ(犚犺狅犱狅犫狉狔狌犿)qÏÐG
È
(犔犲狆狋狅犫狉狔狌犿)"1,ÑÒGÚÓÔ
(犌狔犿狀狅狊狋狅犿犻犲犾犾犪狏犲狉狀犻犮狅狊犪var.犪犮狌犿犻狀犪狋犪)E¼e
z
。
IÕÖ¢2。
W1 BCDEXYPZ;P[\]^_`W
Table1 TaxonomicsystemofMielichhoferiaceae
È+
Family
ý+×s
Subfamily
ôÈ
Genus
¥¦
References
gG+Bryaceae
cdGý+Mielichhoferioideae cdGÈ犕犻犲犾犻犮犺犺狅犳犲狉犻犪,ØÙGÈ(çöx)犎犪狆犾狅犱狅狀狋犻狌犿
gGý+Bryoideae ÕÖGÈ犘狅犺犾犻犪,¸ÚGÈ犕狀犻犫狉狔狌犿,ЯGÈ犈狆犻狆狋犲狉狔犵犻狌犿
[3]
gG+Bryaceae
cdGý+Mielichhoferioideae dGÈ 犛狔狀狋犺犲狋狅犱狅狀狋犻狌犿
,
ØÙGÈ 犎犪狆犾狅犱狅狀狋犻狌犿,cdG
È犕犻犲犾犻犮犺犺狅犳犲狉犻犪
[4]
gGý+Bryoideae
ЯGÈ犈狆犻狆狋犲狉狔犵犻狌犿,¸ÚGÈ犕狀犻狅犫狉狔狌犿,ÕÖGÈ犘狅犺犾犻犪,
öÕÖGÈ犘狊犲狌犱狅狆狅犺犾犻犪
gG+Bryaceae ÕÖGý+Pohlioideae cdGÈ犕犻犲犾犻犮犺犺狅犳犲狉犻犪,ÕÖGÈ犘狅犺犾犻犪 [5]
gG+Bryaceae
cdGý+Mielichhoferioideae dGÈ 犛狔狀狋犺犲狋狅犱狅狀狋犻狌犿
,
ØÙGÈ 犎犪狆犾狅犱狅狀狋犻狌犿,cdG
È犕犻犲犾犻犮犺犺狅犳犲狉犻犪
[6]
gGý+Bryoideae
ЯGÈ犈狆犻狆狋犲狉狔犵犻狌犿,¸ÚGÈ犕狀犻狅犫狉狔狌犿,ÕÖGÈ犘狅犺犾犻犪,
öÕÖGÈ犘狊犲狌犱狅狆狅犺犾犻犪
gG+Bryaceae ÕÖGý+Pohlioideae
cdGÈ犕犻犲犾犻犮犺犺狅犳犲狉犻犪,ÕÖGÈ犘狅犺犾犻犪,ЯGÈ犈狆犻狆狋犲狉狔犵犻
狌犿,¸ÚGÈ犕狀犻狅犫狉狔狌犿
[7]
gG+Bryaceae
cdGý+Mielichhoferioideae cdGÈ犕犻犲犾犻犮犺犺狅犳犲狉犻犪 [8]
gGý+Bryoideae ÕÖGÈ犘狅犺犾犻犪
cdG+ Mielichhoferiaceae ——— cdGÈ 犕犻犲犾犻犮犺犺狅犳犲狉犻犪
,
öÕÖGÈ 犘狊犲狌犱狅狆狅犺犾犻犪,vwGÈ
犛犮犺犻狕狔犿犲狀犻狌犿,dGÈ犛狔狀狋犺犲狋狅犱狅狀狋犻狌犿
[1]
hiG+ Mniaceae ——— ЯGÈ犈狆犻狆狋犲狉狔犵犻狌犿,ÕÖGÈ犘狅犺犾犻犪
gG+Bryaceae ——— cdGÈ犕犻犲犾犻犮犺犺狅犳犲狉犻犪,ЯGÈ犈狆犻狆狋犲狉狔犵犻狌犿 [9]
hiG+ Mniaceae ——— ÕÖGÈ犘狅犺犾犻犪
hiG+ Mniaceae ÕÖGý+Pohlioideae ÕÖGÈ犘狅犺犾犻犪,cdGÈ犕犻犲犾犻犮犺犺狅犳犲狉犻犪,ЯGÈ犈狆犻狆狋犲狉狔犵犻狌犿 [10]
cdG+ Mielichhoferiaceae ——— ЯGÈ犈狆犻狆狋犲狉狔犵犻狌犿
,
ÕÖGÈ犘狅犺犾犻犪,vwGÈ犛犮犺犻狕狔犿犲狀犻
狌犿,cdGÈ犕犻犲犾犻犮犺犺狅犳犲狉犻犪
[11]
gG+Bryaceae ———
cdGÈ犕犻犲犾犻犮犺犺狅犳犲狉犻犪,ÕÖGÈ犘狅犺犾犻犪,ЯGÈ犈狆犻狆狋犲狉狔犵犻
狌犿,öÕÖGÈ犘狊犲狌犱狅狆狅犺犾犻犪
[12,13]
gG+Bryaceae ——— ØÙGÈ犎犪狆犾狅犱狅狀狋犻狌犿 [14]
hiG+ Mniaceae
cdGý+Mielichhoferioideae cdGÈ 犕犻犲犾犻犮犺犺狅犳犲狉犻犪
,
öÕÖGÈ 犘狊犲狌犱狅狆狅犺犾犻犪,ÕÖGÈ
犘狅犺犾犻犪,vwGÈ犛犮犺犻狕狔犿犲狀犻狌犿,dGÈ犛狔狀狋犺犲狋狅犱狅狀狋犻狌犿
hiGý+ Mnioideae ЯGÈ犈狆犻狆狋犲狉狔犵犻狌犿
cdG+ Mielichhoferiaceae ——— cdGÈ 犕犻犲犾犻犮犺犺狅犳犲狉犻犪
,
öÕÖGÈ 犘狊犲狌犱狅狆狅犺犾犻犪,ÕÖGÈ
犘狅犺犾犻犪,ЯGÈ犈狆犻狆狋犲狉狔犵犻狌犿
[2]
hiG+ Mniaceae ——— cdGÈ 犕犻犲犾犻犮犺犺狅犳犲狉犻犪
,
öÕÖGÈ 犘狊犲狌犱狅狆狅犺犾犻犪,ÕÖGÈ
犘狅犺犾犻犪,ЯGÈ犈狆犻狆狋犲狉狔犵犻狌犿
[15]
8631 ! " # $ % & 36¨
W2 abcdefg犌犲狀犅犪狀犽hi$
Table2 TaxasequencedinthisstudywithGenBankaccessionnumbersandvoucherspecimeninformation
seTaxon
+Family Species
Û}~
Voucherinformation
GenBankܬÝGenBankaccessionnumber
狉狆狊4 狋狉狀G 狋狉狀L狋狉狀F 犪狋狆B狉犫犮L
cdG+
Mielichho
feriaceae
ÞÔÕÖG犘狅犺犾犻犪犾犪狋犻犮狌狊狆犻狊 LMNZhaoJiancheng20072654 KX355967 KX289674 — KX355896
ÕÖG犘狅犺犾犻犪犲犾狅狀犵犪狋犪 ßßh£·àáâSulaymanMamtimin07100 KX355968 KX289675 KX355934 KX355897
¶ãÕÖG犘狅犺犾犻犪犾犲狌犮狅狊狋狅犿犪 µäå WangMeizhi081085 KX355985 KX289676 — KX355898
æÕÖG犘狅犺犾犻犪狀狌狋犪狀狊 ßßh£·àáâSulaymanMamtimin14985 KX355971 KX289677 KX355935 KX355899
öÚéÕÖG犘狅犺犾犻犪犾狅狀犵犻犮狅犾犾犻狊 ßßh£·àáâSulaymanMamtimin200426 KX355972 KX289678 KX355936 KX355900
/ýÕÖG犘狅犺犾犻犪犵犲犱犲犪狀犪 1çGaoQian18376 KX355973 KX289679 KX355937 KX355901
èéÕÖG犘狅犺犾犻犪狆狉狅犾犻犵犲狉犪 éCaoTong93277 KX355986 KX289680 — KX355902
èéÕÖG犘狅犺犾犻犪狆狉狅犾犻犵犲狉犪 êëìLiLiangqian764 KX355987 KX289681 KX355938 KX355903
kOÕÖG犘狅犺犾犻犪犮狉狌犱犪 í HeQiang838 KX355975 KX289682 KX355939 KX355904
öÚéÕÖG犘狅犺犾犻犪犾狅狀犵犻犮狅犾犾犻狊 í HeQiang718 KX355976 KX289683 KX355940 KX355905
kOÕÖG犘狅犺犾犻犪犮狉狌犱犪 í HeQiang698 KX355977 KX289684 JF277360.1 KX355906
ÕÖGæÔ犘狅犺犾犻犪犲犾狅狀犵犪狋犪var.犿犻狀狅狉 îï;PeiLinying192 KX355969 KX289685 KX355941 KX355907
/ýÕÖG犘狅犺犾犻犪犵犲犱犲犪狀犪 êðñLiFenxia1053 KX355970 KX289686 KX355942 KX355908
öÚéÕÖG犘狅犺犾犻犪犾狅狀犵犻犮狅犾犾犻狊 9:;LiuYongying1158 KX355974 — JF277344.1 KX355909
hiG+
Mniaceae
Ó¯òiG犘犾犪犵犻狅犿狀犻狌犿犮狌狊狆犻犱犪狋狌犿 ó£ôXiaoMingxuan0417 KX355980 KX289688 KX355944 KX355911
ÑiG犜狉犪犮犺狔犮狔狊狋犻狊狌狊狊狌狉犻犲狀狊犻狊 ó£ôXiaoMingxuan0087 KX355984 KX289689 KX355945 KX355912
ÑiG犜狉犪犮犺狔犮狔狊狋犻狊狌狊狊狌狉犻犲狀狊犻狊 ó£ôXiaoMingxuan0086 KX355981 KX289690 KX355946 —
Z¯õiG犘犾犪犵犻狅犿狀犻狌犿狊狌犮犮狌犾犲狀狋狌犿 ó£ôXiaoMingxuan0420 KX355978 KX289691 KX355947 KX355913
ö¯hiG犕狀犻狌犿狊狆犻狀狅狊狌犿 ó£ôXiaoMingxuan0098 KX355983 — KX355948 KX355914
÷¯hiG犕狀犻狌犿狋犺狅犿狊狅狀犻犻 ó£ôXiaoMingxuan0415 KX355982 KX289692 KX355949 KX355915
øùõiG犘犾犪犵犻狅犿狀犻狌犿犲犾犾犻狆狋犻犮狌犿 ó£ôXiaoMingxuan0105 KX355979 KX289693 KX355950 KX355916
gG+
Bryaceae
ú¯+éG犘犾犪犵犻狅犫狉狔狌犿犵犻狉犪犾犱犻犻 ûüý WangYoufang4354A KX355958 KX289687 KX355943 KX355910
ÎG犃狀狅犿狅犫狉狔狌犿犼狌犾犪犮犲狌犿 9:;LiuYongying5036 KX355951 AY163052.1 KX355917 KX355880
[OÌÍG犅狉犪犮犺狔犿犲狀犻狌犿狆犲狀犱狌犾狌犿 ßßh£·àáâSulaymanMamtimin07021 KX355952 KX289659 KX355918 KX355881
®ÔÌÍG犅狉犪犮犺狔犿犲狀犻狌犿犲狓犻犾犲 êþÿLiXiuqin99063 KX355961 KX289660 KX355919 KX355882
!ùÌÍG犅狉犪犮犺狔犿犲狀犻狌犿犾狅狀犵犻犱犲狀狊 é"CaoNa20070929 KX355965AY163055.1 KX355920 KX355883
#éÌÍG犅狉犪犮犺狔犿犲狀犻狌犿狆狋狔犮犺狅狋犺犲犮犻狌犿 û$£ WangQinghua751 KX355953AY163058.1 KX355921 KX355884
j¯ÌÍG犅狉犪犮犺狔犿犲狀犻狌犿犮犪狆犻狋狌犾犪狋狌犿 êðñLiFenxia1633 KX355962 KX289661 KX355922 KX355885
r£ÌÍG犅狉犪犮犺狔犿犲狀犻狌犿狊犻狀犲狀狊犲 µäå WangMeizhi48304 KX355955 KX289662 KX355923 KX355886
ÌÍG犅狉犪犮犺狔犿犲狀犻狌犿狀犲狆犪犾犲狀狊犲 é"CaoNa20070948 KX355966 KX289663 KX355924 AF546823.1
æ%gG犅狉狔狌犿狆犪犾犾犲狊犮犲狀狊 ßßh£·àáâSulaymanMamtimin15265 KX355957 KX289666 KX355926 KX355889
¯gG犅狉狔狌犿犮犪狆犻犾犾犪狉犲 &(NiuYulu20100196 KX355956 KX289667 KX355927 KX355890
&égG犅狉狔狌犿狋狌狉犫犻狀犪狋狌犿 ßßh£·àáâSulaymanMamtimin15095 KX355960 KX289668 KX355928 KX355891
)%gG犅狉狔狌犿狆狌狉狆狌狉犪狊犮犲狀狊 &(NiuYulu110045 KX355959 KX289669 KX355929 KX355892
ú
gG犅狉狔狌犿犪犿犫犾狔狅犱狅狀 *¦+PengXuanjun20050189 — KX289670 KX355930 KX355893
¨¯gG犅狉狔狌犿狋犺狅犿狊狅狀犻犻 &(NiuYulu20110900 KX355954 KX289671 KX355931 KX355894
,gG犅狉狔狌犿犫狅狉狀犺狅犾犿犲狀狊犲 9:;LiuYongying415 KX355963 KX289672 KX355932 KX355895
Z¯G犚犺狅犱狅犫狉狔狌犿狉狅狊犲狌犿 ûüý WangYoufang2011-0127 KX355964 KX289673 KX355933 —
-G+
Meesiaceae ÏÐG犔犲狆狋狅犫狉狔狌犿狆狔狉犻犳狅狉犿犲 ê.LiLin030570 KX355988 KX289665 AF023736.1 KX355888
[G+
Pottiaceae
ÑÒGÓ¯Ô犌狔犿狀狅狊狋狅犿犻犲犾犾犪狏犲狉狀犻犮狅
狊犪var.犪犮狌犿犻狀犪狋犪 9:;LiuYongying05433 KX355989 KX289664 KX355925 KX355887
/
:¢F
ú0)GenBank,—¢Fcñ
Note:indicatessequencedownloadfromGenBank,and—indicatesabsentdata
96317N 9:;,2:,
%±²qsjrcdG+¹º6B
1.2 j k
1.2.1 lm;P7HI ÂÃǼyl¥
¦
[2,613,2937],
1È23
、
o43q5U6o4
32@rsxjcdG+#$qylez4000
{|}~j
±²6B
。
Û}~Ê7
."QWZ%#$}~8
(HBNU)。
1.2.2 犇犖犃no:犘犆犚pq ¤È23ú9
,
:;n<=ˤ®j>7
,
)*?.d@
DNAhABCD("EFOð+G¨HIJ)
KÃL£MhANDNA。PCROÊ25μLÏ
Y
(
N DNA20ng,dNTP200mmol·L-1,10×
plusbuffer犜犪狇2.5μL,primer"0.5μmol·L
-1,
MgCl21mmol·L-1,犜犪狇polymerase0.1unit),
$IÕÖ¢3。
"OÊP·íj¹QºR©E95℃S
ÔÁ7min,d72℃TU5min。rÉ35V
W
,
ºRú
:狉狆狊4、狋狉狀L狋狉狀F:95℃(25s),58℃
(5s),72 ℃ (2 min);犪狋狆B狉犫犮L、狋狉狀G:95 ℃
(25s),47℃(5s),72℃(2min)。OÊX$Y®Z
[\O$]%+G¨HIJ^
,
^
h_
GenBank(¢2)。
1.2.3 =>rs ^j
`¼¤ß
(http://blast.ncbi.nlm.nih.gov/Blast.cgi)f
GenBankrja@,wb<¨¼Àcdj
ʤ
。
@ClustalXef[47]@gsj
)hqihb
qa@
,
Í<
jkjalq
¬rsm
。
1 MEGA5.05[48],nDNA
MYk
,
¤
o¨opqrj÷hús
æ
tYks
。
~6Brjse
zóXWhiG+
、
gG+
、
cdG+q-G+
(Meeseaceae)2;SrÝs}~T!LMu
vsYks
,
Z狉狆狊4、狋狉狀G、狋狉狀L
狋狉狀Fq犪狋狆B狉犫犮L24
jw¨40|。@
W3 tufg:vw
Table3 Informationofprimersusedtoamplify
thefourregions
DNA
Marker
$
Primer
ÂÁ¥¦
Reference
狉狆狊4 R
:TACCGAGGGTTCGAATC
F:ATGTCCCGTTATCGAGGACCT
[43]
犪狋狆B狉犫犮L R
:ACATCKARTACKGGACCAATAA
F:AACACCAGCTTTRAATCCAA
[44]
狋狉狀L狋狉狀F R
:CGAAATCGGTAGACGCTACG
F:ATTTGAACTGGTGACACGAG
[45]
狋狉狀G R
:ACCCGCATCGTTAGCTTG
F:GCGGGTATAGTTTAGTGG
[46]
sez8´
、
Mx
js@
(NJ)qZ*(ML)y´z({|“ÚÔ}”
~jXO
,
T^MºyjYkpq
[49]。
2 ¡fs
NJq MLjsC@4DNA
js ¡,~¼
,
¤= NJO
Fs ¡
(
1),wÑÒGÚÓÔ,s
ysEA、Bj)©ª«©E100%jRs©。
s©ASÕÖGÈ、hiGÈ(犕狀犻狌犿)、òi
GÈ
(犘犾犪犵犻狅犿狀犻狌犿)qÑiGÈ(犜狉犪犮犺狔犮狔狊狋犻狊)3
N
,
óôW3´Zj³$s©。²,2|kOÕ
ÖG
(犘狅犺犾犻犪犮狉狌犱犪)、3|öÚéÕÖG(犘.犾狅狀犵犻
犮狅犾犾犻狊)、ÞÔÕÖG(犘.犾犪狋犻犮狌狊狆犻狊)、2|/ýÕÖ
G
(犘.犵犲犱犲犪狀犪)、ÕÖG(犘.犲犾狅狀犵犪狋犪)qæÕÖG
(犘.狀狌狋犪狀狊)2¦Na©,»*©ª«¬
50%,½
¨£oj
%¿ÀÁÂ
:
#$Ï
;
¾¯Oã
m
;
méÚ
,
Ü
,
nݣo
,
fÒÝ2Ú
,
mé¼
°Ú
,
oÑ
;
Çdìd¨ÞÙjj
,¨
d¯
,
ÚáÌ
,¨
=
,
áïð
。
vs©f Hil
2
[11]
jÕÖG3
(犘狅犺犾犻犪)óôjRS$,
~¼
。
³
,2|èéÕÖG(犘.狆狉狅犾犻犵犲狉犪)
q¶ãÖG
(犘.犾犲狌犮狅狊狋狅犿犪)¦Nb©,©ª«E
100%。vs©f Hil 2[11]jéG3(犆犪犮
狅犱狅狀)óôjRS$,~¼,£oj±²Æ©
¨¯Oã°
。
>ezɤO
、
#tZÐ
、¯
jî
、¯
、¯
°ZÐq
Â2Cisy
,
m¥RS¤ã°jZÐ
、
Â
、
8q¯,,j
sðíî
,
¡¤¾ã°Z
,
>?Pms
。
%®opj¼Á£|YklYÌÍ
[50]。
s
³
,
hiGÈ
、
òiGÈqÑiGÈj¦Ec©,
)©ª«E100%,©ªW,se%j6
B ¡
———
hiG+ƼÅYez
。2|
Ñ
iGj(}~s¥¦¤¬j³$s©r
,
yv
ÆS
“
ÚÔ}
”
~
。
s©BóXgGÈ、ÌÍGÈ、Z¯GÈ、+é
GÈqÎGÈ
,
¿¨j
%ÁÂE
:
#$ÏÐÑ
,
¯×Ø
Ùè
,
r®Ý°´Ì
,
Úà
Ùù
;
mÏOÒè
;
édpqÓrs
。
fCrosby2[1]、
Hil2[11]qShaw2[2]jgG+W¼。
3 b
3.1 xyDz{|DE?L[
¤,ssºjsr
,
ó
0731 ! " # $ % & 36¨
s©®j¢F)©ª«≥50%。A.ÕÖGÈhiG+;B.gG+;a.ÕÖG3;b.éG3;c.hiG+
1 狉狆狊4、狋狉狀G、狋狉狀L狋狉狀Fq犪狋狆B狉犫犮L
Numbersabovebranchesarebootstrapvalues≥50%from1000replicates.A.犘狅犺犾犻犪Mniaceae;B.Bryaceae;
a.Section犘狅犺犾犻犪;b.Section犆犪犮狅犱狅狀;c.Mniaceae
Fig.1 Consensustreeofneighborjoining(NJ)treederivedfrom狉狆狊4+狋狉狀G+狋狉狀L狋狉狀F+犪狋狆B狉犫犮Ldata
ôÕÖGÈqhiG+ezjs© AfóôgG
+ezjs©B©¨P1j©ª«,fzTOjs
¡
[2326]
¼
,
¼º£WÕÖGȬÈ
gG+
,^
fhiG+jlYÌÍ
。
¤s
erRS,mÏjyÁÕÖGÈgG
+
,
½jKjuϨ´æµ¶
(
2):ÕÖGÈ#
$¯´ÞÚ
,
Ú×Ø
Ù×Ø
,¯
x¾sðù
,
r
-èÙñ¯Óú
,¯
°ÞÚ
,
âã
;
gG
+#$¯´j
,
è
、
ÛÜ
、
á×Ø
,¯
xÓ
¨£osðù
,
r¡
,
¢è
、
rº¯ÓÙ
NÚ
¯Ó
,¯
°´j
,
ù
ÙÚà
。
w
,
þ
%ÁÂqs
s
,
ÕÖGÈ·_ºgG+
,
>¼g5£¤´¥N¿Î
,
8¸k·zNp¢
j¥¦r
[12,911,1415,28,4142,51]。
3.2 xyDz{n}DE?L[
¦§s ¡r
,
ÕÖGÈ#$fhiG+
17317N 9:;,2:,
%±²qsjrcdG+¹º6B
A~D.¨ÓgG;E~H.öÚéÕÖG;I~L.æéõiG.A、E、L.#t;B、F、K.¯;C、H、J.¯ÓÝ°;D、G、I.¯rÝ°
2 cdG+、gG+qhiG+#$
±²a´
A-D.犅狉狔狌犿狀犲狅犱犪犿犲狀狊犲;E-H.犘狅犺犾犻犪犾狅狀犵犻犮狅犾犾犻狊;I-L.犘犾犪犵犻狅犿狀犻狌犿犿犲犱犻狌犿.A,E,L.Plant;B,F,K.Leaves;
C,H,J.Distalleafcels;D,G,I.Medialleafcels
Fig.2 ComparisonofmorphologicalcharactersforMielichhoferiaceae,BryaceaeandMniaceae
+ôez¦¤¼¨
,
¨P1j©ª«
,
½S¤
Yk©ð¾íryv¨ª«¬á®æÁp
O
,
wsu¯f$¼
[52],
|IÉj
YkpqlY¨gæj©ª
。
T
ÁÂa
´
(
2):ÕÖGÈ#t¼°´Ð,¯
q¯°Þ
Ú
,
OäåÓORÒ,ÝjçOÔè
,
Çìéd¨
¬íîjïð
;
hiG+#t¼°ZÑ
,¯
ÓÌ
^ø
,¯
°£¤2±
,
mÏÓORÒè
,
éd
pqrs
。
@w
%ÁÂE,²jse%
¡fwsE,²jÿ!se% ¡jqr
,
yvjȳÆ>jezQ¿®
,
¤´´N
pOsQ
,
Sµáµ?(7JKUfj,4
,
Ê
¤cñez
,
¶ªezIÉ·xlY¸M
,
wo
¨y
%ÁÂ
。
SÃK
%®¨´Zµ
¶
,¨
{hºwseM\¨Ojjsez¤
£¤¹¬y
ÁÂjº»ú
,
¬·vÄE¼
ez
,
ÕÖGezyvÆRNT¸¹jhiG+
sº+©ð^Oj
[2]。
,ÕÖGÈfhiG
+Iɾ
%¿ÀÁÂ
,
¼½|¾¬·8ÄE
¼ÅYez
,
»\cdG+¿¤¯
。Hil 2[11]
ÂÃKoponen[27]jÐ+~Goffinet2[28]hº
jhiG+s
,
ÕÖGÈ
、
cdGÈ
、
Я
2731 ! " # $ % & 36¨
GÈqvwGÈ2ez»\EcdG+
,Crosby
2
[1]、Shaw[2]qShaw2[51]¼?%K©ªW
>g5
。Cao2[53]1À¼ÁsabÕÖ
GÈfhiG+jYklY
,
©ªWÕÖGÈ
cdG+jg5
。
3.3 xyDz{BCDz、~DzPZ?L[
SЯGÈjéd fÕÖGÈj£¤¾
µ¥
,Andrews[54]q Ochi[55]ÂJEÕÖGÈ
j¼3
。
¤Pedersen2[2526],
%ÁÂq
cpDNA
þsºjYkpqrЯG
ÈfÕÖGÈlYÌÍ
。Guerra2[56]H¾cpDNA
s³Wyj ¡
,
]EЯGÈf
Hil2[11]¹jÕÖGȸÚG3(犃狆犪犾狅犱犻犮
狋狔狅狀)NlYÌÍjÃÄz。T
%±²s
¬pÿ|j¿ÀÁÂÆøà
j¯°
。
cd
GÈfÕÖGÈjÕÖG3¨yjuÏ
,
ÃK
jm¥RSÆéd
:
ÕÖG3jéd£ojì
,
¨pqëzj¼dìqpqëzá¼íîïðj
Çdì
;
cdGÈÓc¨ïðj¼dì
,¨
Z³¨,
ÅᬣojÇdì
[57]。
cdGÈfÕÖGÈE
Yezs©ª
[2324]。
ÕÖGÈf
dGÈ
、
öÕÖGÈjYklY¦§Æ¾s
,
½Ç¾mÏ
、
cwuÏ£¤¾ms>ÈÈ#
$
,
É
ÁÂjyÁyw]E|ÆYkl
YÌÍjez
,
¼cdG+
[1,2]。
4
4.1 BCDE_PZ
,
%ÁÂqs
jþs
,
cd
Gez·_ºgG+
;
,fhiG+cÊ
%
ÁÂj¿ÀÁ
,
cdGez³¬·phiG+
。
SË>ÌðjO$©ðÍßÎ^
,
§¨pqy@rjëìéd jdGÈyv
fÕÖGÈ´,QjezYklYÌÍ
;
§¨Ïî
ïðjéd jcdGÈÐÈѼº»
,
õ
fÕÖGÈ´ðjezYklYÌÍ
。
wyj
¯
q¯°ÎÒЯGÈyvfÕÖGÈj¸Ú
G3YklYÌÍ
。
<éd ¼
,
£¤¾¹
öÕÖGÈfÕÖGÈ
%ÁÂjµ¶
,
w|yv¨Àyáy
。
NI
,
cdG+·Æ
¼)*ez
,
cdG+ÇÈÉʤËÌÍjYk
pqlY
。
cdG+RSÎ¥±²E
:
#$ÏÐÑ
,
ÒÓs
Ô
;¯
q¯°EÕÖGÑ
(犘狅犺犾犻犪like),×Ø
ÙÚÛÜ
,
r®Ý°ÞÚ
,
ßÂà
áâã
;
OäåæOçOÔè
;
édEêOëdì
,
Ó¨¬
íîjïðá¼ìédcñ
,
òëìédcñ
。
4.2 @ABCDE;P
rcdG+8÷¨5È,34(ôúse
Åo
)。
@ABCDE;z
W
1.#tÓÔ+,¨ 1~3
aÕ¯ÐjÖ¯;¯ ÛÜ
,¯ ù¨1~4
ÞÚ°Njsðù
~Dz犈狆犻狆狋犲狉狔犵犻狌犿
××××××××
××××××××××××××××××××××××××××××××××××
1.#t¬Ô+,¾Ö¯qÕ¯Is;¯ ×Ø
ÙÚ×Ø
,¯ ù¾£oÞÚ°Njsðù 2×××××××
2.édÓïð,ëìáÅì,òcñ BCDz犕犻犲犾犻犮犺犺狅犳犲狉犻犪
2.éd£oëì 3××××××××××××××××××××××××××××××××××××××××
3.ÇdìØÙ¼dì®,¬\s,yjd¯yêÙ,fd¬\ms CDz犛狔狀狋犺犲狋狅犱狅狀狋犻狌犿
3.Çdìf¼dìs,Ǩd¯\fdms 4××××××××××××××××××××××××××
4.mé\,édN@,Çdìf¼dì2Ú xyDz犘狊犲狌犱狅狆狅犺犾犻犪
4.méÚ\ÙúÛ,éds¬,Çdì´¼dìÌ xyDz犘狅犺犾犻犪
!Q
:
[1] CROSBYMR,MAGILLRE,ALLENB,犲狋犪犾.AChecklist
oftheMosses[M].St.Louis:MissouriBotanicalGarden,
1999:1210.
[2] SHAW AJ.BryophyteFloraofNorthAmerica[DB/OL].
MissouriBotanicalGarden,(2009323)[20160515].ht
tp://www.mobot.org/plantscience/BFNA/V2/BryaMieli
chhoferiaceae_family.htm.
[3] BROTHERUSVF.Musci(Bryaceae)[M]//ENGLERA,
PRANTLK.DienatürlichenPflanzenfamilien1ed.Hlfte.
37317N 9:;,2:,
%±²qsjrcdG+¹º6B
Leipzig:WilhenlmEngelmann,1909:532601.
[4] BROTHERUSVF.Musci(Bryaceae)[M]//ENGLERA,
PRANTLK.DienatürlichenPflanzenfamilien2ed.Hlfte.
Leipzig:WilhenlmEngelmann,1925:347404.
[5] ANDREWSA L.StudyintheBryaceae[J].犅狉狔狅犾狅犵犻狊狋,
1950,53:115118.
[6] ,ÜÝÞ,1 ç,2.rGe#$È (®¡)[M].
"E
:
+%º¤ß
,1963:251270.
[7] GANGULEEHC.MossessofEasternIndiaandAdjacentRe
gions.Vol.2[M].Calcutta:EastendPrinters,1974:888
1022.
[8] NYHOLM E.Ilustrated MossFloraofFennoscandia.2.
Musci.Fasc.3[M].Lund:CWKGleerup,1958:189288.
[9] IWATSUKIZ.NewcatalogofthemossesofJapan[J].犑狅狌狉
狀犪犾狅犳狋犺犲犎犪狋狋狅狉犻犅狅狋犪狀犻犮犪犾犔犪犫狅狉犪狋狅狉狔,2004,96:1182.
[10] SMITHAJE.TheMossFloraofBritainandIreland[M].
Cambridge:CambridgeUniversityPress,2004:528636.
[11] HILLMO,BELLN,BRUGGEMANNANNENGA M A,
犲狋犪犾.AannotatedchecklistofthemossesofEuropeand
Macaronesia[J].犑狅狌狉狀犪犾狅犳 犅狉狔狅犾狅犵狔,2006,28(3):
198267.
[12] àáâ.rFG (§4¨)[M]."E:+%º¤ß,2006:
1111.
[13] ZHANGDC,LIXJ,HES.Bryaceae[M]//LIXJ,CROS
BY M R,HES.MossFloraofChina.EnglishVersion.
Vol.4.Beijing:SciencePress& StLouis,MO:Missouri
BotanicalGardenPress,2007:392
[14] FREYW,STECHM,FISCHERE.SylabusofPlantFami
lies[M].Berlin:Gebr.BorntraegerVerlagsbuchhandlung,
2009:193198.
[15] ã ä,í å.FG#$,rO$$x¬ §¼¨[M].
"E
:
+%º¤ß
,2013:125147.
[16] FUNCHHC.CryptogamischeGewchsedesFichtelgebirgs
[M].Leipzig:Grau,1818.
[17] NEESvon ESENBECK C G D,HOMSCHUCH C F,
STURMJ.BryologiaGermanica.Vol.2[M].Nürnberg:
NabuPress,1831.
[18] BRUCHP,SCHIMPERWP,GüMBELWT.BrologiaEu
ropaea.Vol.4[M].E.Stuttgart:Schweizerbart,1836
1855.
[19] SCHIMPER W P.CorolariumBryologiacEuropaea[M].
Stuttgartiae:E.Schweizerbart,1856.
[20] SMITHAJE.TheMossFloraofBritainandIreland[M].
Cambridge:CambridgeUniversityPress,1978.
[21] CRUM HA,ANDERSONLE.MossesofEasternNorthA
merica[M].NewYork:ColumbiaUniversityPress,1981.
[22] DIXONHN.TheStudentsHandbookofBritishMosses3
[M].Eastbourne:V.T.Sumfield,1954:325376.
[23] COXCJ,HEDDERSONTAJ.Phylogeneticrelationships
amongtheciliatearthrodontousmosses:evidencefromchlo
roplastandnuclearDNAsequences[J].犘犾犪狀狋犛狔狊狋犲犿犪狋犻犮狊
犪狀犱犈狏狅犾狌狋犻狅狀,1999,215:119139.
[24] COXCJ,GOFFINETB,NEWTONAE,犲狋犪犾.Phyloge
neticrelationshipsamongthediplolepideousalternatemosses
(Bryidae)inferredfrom nuclearandchloroplastDNAse
quences[J].犅狉狔狅犾狅犵犻狊狋,2000,103(2):224241.
[25] PEDERSENN,COXCJ,HEDENSL.Phylogenyofthe
mossfamilyBryaceaeinferredfrom chloroplastDNA se
quencesandmorphology[J].犛狔狊狋犲犿犪狋犻犮犅狅狋犪狀狔,2003,28
(3):471482.
[26] PEDERSENN,HEDENSL.Taxonomicandnomenclatural
implicationsofphylogeneticstudiesoftheBryaceaebasedon
moleculardataandmorphology[J].犅狉狔狅犾狅犵犻狊狋,2005,108
(1):123128.
[27] KOPONENT.ThephylogenyandclassificationofMniaceae
andRhizogoniaceae(Musci)[J].犑狅狌狉狀犪犾狅犳狋犺犲犎犪狋狋狅狉犻犅狅
狋犪狀犻犮犪犾犔犪犫狅狉犪狋狅狉狔,1988,64:3746.
[28] GOFFINETB,BUCK W R.Systematicsofthebryophyta
(mosses):from moleculestoarevisedclassification[M]//
GOFFINETB,HOLLOWELLVC,MAGILLRE.Molecu
larSystematicsofBryophytes.St.Louis:MissouriBotanical
GardenPress,2004:205239.
[29] æ%ë.ÇçGFG#$ [M].èqé±:ÇçGZ%º¤
ß
,1997:263293.
[30] êëí,µäå.ìÒmFG [M]."E:+%º¤ß,
2000:388411.
[31] íîï.ðñ#$ (§3¨,§¼¡)[M]."E:+%º¤
ß
,1978:99114.
[32] Lòó,é .ôFG#$ [M].õ/:ô+%Göº
¤ß
,1998:165185.
[33] 1 ç.ô"÷e#$ [M]."E:+%º¤ß,1977:
142159.
[34] àáâ,ø ù.!úFG#$ [M]."E:+%º¤ß,
198:161194.
[35] àáâ.û/#$ (§18¨)[M]."E:+%º¤ß,2002:
382448.
[36] ZHANGDC,LIXJ,HIGUCHIM.Twonewspeciewsof
thegenus犘狅犺犾犻犪 (Bryaceae,Musci)formChina[J].犃犮狋犪
犘犺狔狋狅狆犺狔狊犻狅犾狅犵犻犮犪犛犻狀犻犮犪,2002,40(2):176182.
[37] KOPONENT,LOUJS.犘狅犺犾犻犪犺犻狊犪犲,speciesnova(Mus
ci,Bryaceae)fromWolongNatureReserve,Sichuan,China
[J].犎犻犽狅犫犻犪,1986,9:315317.
[38] LIU YY,ZHAOJC,MAMTIMINS.犛狔狀狋犺犲狋狅犱狅狀狋犻狌犿
犽狌狀犾狌狀犲狀狊犲(Mielichhoferiaceae,Musci),anewmossspecies
fromtheKunlunMountainRange,China[J].犖狅狏狅狀,2013,
22(4):428433.
4731 ! " # $ % & 36¨
[39] µ¶·,LMN.ÕÖGÈ(犘狅犺犾犻犪Hedw.)jYkop-È
ÇÉlYab
[J].#$6B,2010,30(5):517526.
WANGCY,ZHAOJC.Phylogeneticpositionofthegenus
犘狅犺犾犻犪 Hedw.:evidencefromthenuclearribosomalDNA
ITSregion[J].犅狌犾犾犲狋犻狀犅狅狋犪狀犻犮犪犚犲狊犲犪狉犮犺.2010,30(5):
517526.
[40] 9:;,ßßh£·àáâ,LMN.çüÕÖGÈ(犘狅犺犾犻犪)
jse%ýþ
[J]../m6B,2013,30(3):505511.
LIUYY,MAMTIMINS,ZHAOJC,犲狋犪犾.Arevisionof
犘狅犺犾犻犪Hedw.(Mielichhoferiaceae,Musci)inXinjiang,Chi
na[J].犃狉犻犱犣狅狀犲犚犲狊犲犪狉犮犺,2013,30(3):505511.
[41] LIUYY,CAOZ,LIL,犲狋犪犾.Threepropaguliferousspecies
of犘狅犺犾犻犪(Mielichhoferiaceae,Musci)inChina[J].犑狅狌狉狀犪犾
狅犳犅狉狔狅犾狅犵狔,2015,37(2):120127.
[42] ÿÀç.ÄÅFG#$ §1¨[M].Ä!:ÄÅ+Gº¤
ß
,2014:359461.
[43] NADOTS,BITTARG,CARTERL,犲狋犪犾.Aphylogenetic
analysisofmonocotyledonsbasedonthechloroplastgene
狉狆狊4,usingparsimonyandanewnumericalpheneticsmethod
[J].犕狅犾犲犮狌犾犪狉犘犺狔犾狅犵犲狀犲狋犻犮狊犪狀犱犈狏狅犾狌狋犻狅狀,1995,(4):
257282.
[44] CHIANGTY,SCHAALBA,PENGCI.Universalprim
ersforamplificationandsequencinganoncodingspacerbe
tweenthe犪狋狆Band狉犫犮LgenesofchloroplastDNA[J].犅狅
狋犪狀犻犮犪犾犅狌犾犾犲狋犻狀狅犳犃犮犪犱犲犿犻犪犛犻狀犻犮犪 (Taipei),1998,39:
245250.
[45] TABERLETP,GIELLYL,PAUTOUG,犲狋犪犾.Universal
primersforamplificationofthreenoncodingregionsofthe
chloroplastDNA[J].犘犾犪狀狋犕狅犾犲犮狌犾犪狉犅犻狅犾狅犵狔,1991,17
(5):11051109.
[46] SHAW AJ,COXJC,BOLESSB.PolarityofPeatmoss
(Spbagnum)evolution:whosaysbryophyteshavenoroots?
[J].犃犿犲狉犻犮犪狀犑狅狌狉狀犪犾of犅狅狋犪狀狔,2003,90 (12):1777
1787.
[47] THOMPSONJD,GIBSONTJ,PLEWNIAKF,犲狋犪犾.The
ClustalXwindowsinterface:flexiblestrategiesformultiple
sequencealignmentaidedbyqualityanalysistools[J].犖狌犮犾犲
犻犮犃犮犻犱狊犚犲狊犲犪狉犮犺,1997,24:48764882.
[48] TAMURAK,PETERSOND,PETERSONN,犲狋犪犾.MEGA5:
molecularevolutionarygeneticsanalysisusingmaximumlikeli
hood,evolutionarydistance,andmaximumparsimonymethods
[J].犕狅犾犲犮狌犾犪狉犅犻狅犾狅犵狔犪狀犱犈狏狅犾狌狋犻狅狀,2011,28:27312739.
[49] à¼", à,íý+.Ykpq6Br“ÚÔ}”~~
V
[J].ª«,2007,29(6):659667.
LIYW,YUL,ZHANGYP.“LongbranchAttraction”ar
tifactinphylogeneticreconstruction[J].犎犲狉犲犱犻狋犪狊,2007,29
(6):659667.
[50] SHAWAJ.Ataxonomicrevisionofthepropaguliferousspe
ciesof犘狅犺犾犻犪Hedw.(Musci)inNorthAmerica[J].犑狅狌狉狀犪犾
狅犳狋犺犲犎犪狋狋狅狉犻犅狅狋犪狀犻犮犪犾犔犪犫狅狉犪狋狅狉狔,1981,50:181.
[51] SHAW AJ,RAMSAYHP.2013.AustralianMossesOn
line.69.Mielichhoferiaceae:犘狅犺犾犻犪[DB/OL].(201352)
[2016515].http://www.anbg.gov.au/abrs/Mosses_
online/Mielichhoferiaceae_Pohlia.pdf.
[52] KNOWLESLL.Estimatingspeciestrees:methodsofphylo
geneticanalysiswhenthereisincongruenceacrossgenes[J].
犛狔狊狋犲犿犪狋犻犮犅犻狅犾狅犵狔,2009,58(5):463467.
[53] CAOZ,LIUYY,ZHAOJC.Efficientdiscriminationof
somemossspeciesbyFourierTransformInfraredSpectrosco
pyandchemometrics[J],犑狅狌狉狀犪犾狅犳犛狆犲犮狋狉狅狊犮狅狆狔,2014:1
9.
[54] ANDREWSAL.Bryaceae[M].//GROUTAJ.MossFlo
raofNorthAmerica,NorthofMexicoVol.2[M].Ver
mont:Newfane,1935,184242.
[55] OCHIH.ARevisionofMossesoftheBryaceaeinJapanand
theAdjacentRegions[M].Tottori:TheBiologicalInstitute,
FacultyofLiberalArts,TottoriUniversity,1959,1124.
[56] GUERRAJ,JIMéNEZMARTíNEZJF,CANOMJ,犲狋犪犾.
AcontributiontothephylogeneticstudyofMielichhoferiace
aeMniaceae(Bryophyta)basedonmolecularsequencedata
[J].犖狅狏犪犎犲犱狑犻犵犻犪,2011,93(1):4756.
[57] SHAWAJ,CRUMH.Peristomehomologyin犕犻犲犾犻犮犺犺狅犳犲
狉犻犪andataxonomicaccountofNorthAmericanspecies[J].
犑狅狌狉狀犪犾狅犳狋犺犲 犎犪狋狋狅狉犻犅狅狋犪狀犻犮犪犾犔犪犫狅狉犪狋狅狉狔,1984,57:
363381.
(
!"
:
#$%
)
57317N 9:;,2:,
%±²qsjrcdG+¹º6B