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Morphological Characters and New Geographical Distribution of Pohlia lutescens (Bryophyta, Mielichhoferiaceae) in China

念珠丝瓜藓(缺齿藓科)形态特征及其在中国的新分布



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,2016,36(6):1250-1256
犃犮狋犪犅狅狋.犅狅狉犲犪犾.犗犮犮犻犱犲狀狋.犛犻狀.
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I J:K\]^_`aJb犘狅犺犾犻犪(cdJ+ Mielichhoferiaceae)#$efghi%34,]jbklmno
Bpqrstuvwux
———
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Š

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犕狅狉狆犺狅犾狅犵犻犮犪犾犆犺犪狉犪犮狋犲狉狊犪狀犱犖犲狑犌犲狅犵狉犪狆犺犻犮犪犾犇犻狊狋狉犻犫狌狋犻狅狀狅犳犘狅犺犾犻犪
犾狌狋犲狊犮犲狀狊(犅狉狔狅狆犺狔狋犪,犕犻犲犾犻犮犺犺狅犳犲狉犻犪犮犲犪犲)犻狀犆犺犻狀犪
WANGXiaorui1,2,LILin1,LIMin1,ZHAOJiancheng1
(1ColegeofLifeScience,HebeiNormalUniversity,Shijiazhuang050024,China;2HuihuaColegeofHebeiNormalUniversi
ty,Shijiazhuang050091,China)
犃犫狊狋狉犪犮狋:Basedonthesystematicalytaxonomicstudyofthegenus犘狅犺犾犻犪Hedw.inChina,thispaperre
visesaspeciesofthegenuswithtoruloidgemmaegrowingontherhizoids,犘狅犺犾犻犪犾狌狋犲狊犮犲狀狊.Morphologi
calcharacters,geographicaldistributionandhabitatofthisspecieswerediscussed.Thecomparisonamong
犘.犾狌狋犲狊犮犲狀狊anditsrelatedspecies,犘.犮狉狌犱犪,犘.犺狔犪犾狅狆犲狉犻狊狋狅犿犪and犘.狀狌狋犪狀狊,werealsoconducted.
TheresultofidentificationofspecimensshowsthatthedistributionrangeinChinaof犘.犾狌狋犲狊犮犲狀狊isex
pandedfromfivetotenprovinces.Meanwhile,thedistributionmapofthespeciesinChinaisprovided.
犓犲狔狑狅狉犱狊:Bryaceae;犘狅犺犾犻犪犮狉狌犱犪;gemma;newdistribution
  `aJb(犘狅犺犾犻犪Hedw.)©ª«p¬¬­
®¯°SJ.E.Pohl±V²,Ž^³²´µ¶Ž·
¸y¹`a
[1]。
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,`
aJb»¼½
w¾¿y

·¸m¿uÀÁ

·¸ÂˌlÄŸ
d±ÆÇȧJ+
(Bryaceae)^ [13]。ÉŸ,,È
hÊefË1h–ÌÍ]§J+ŒŽÎ#$ue
fhi%34ÏÐÑÒÓ`aJbºfÔÕÖ×
ØJ+
(Mniaceae)ubÙeÚÛÜ[45],»Ý,Áh
IJ%NÞ`aJbßà×ØJ+
[68]。
±`aJ
b#$œyzÏá}~ԍ×ØJ+âã•U

ä
åyz}~hÊefË1uæ—

çNèµ
Hil[9]Shaw[1011]ué€,Þ`aJbŽb,
¹cdJb
(犕犻犲犾犻犮犺犺狅犳犲狉犻犪 Nees)、ê`a J b
(犘狊犲狌犱狅狆狅犺犾犻犪Wiliams)ë¼Jb(犈狆犻狆狋犲狉狔犵犻狌犿
Lindberg)8,ÇÈcdJ+(Mielichhoferiaceae)ì。
pq`aJ
[犘狅犺犾犻犪犾狌狋犲狊犮犲狀狊 (Limpr.)H.
Lindb.]»Ž#$í^îÁ¼ïð,ÁunoÔl
m}ñupqrstuvw± òÈ`aJbŽó
xi

jx6ôõhÈ^Uö÷ø

çNœ
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ò

è"

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犱犪)、’d`aJ(犘.犺狔犪犾狅狆犲狉犻狊狋狅犿犪)“`aJ
(犘.狀狌狋犪狀狊)8Jiu4Êí5ª6。œ7m·
Êí8œu9:î

pq`aJûûÆ;<=›ª
‘B`aJ8ŽóJi

»Ý

o—³>?@

pq
`aJœ^uhõAÈìBC

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EF

G
H!I5/ [23,8]。
jJK]pq`aJuyz}~‚ƒ„L½u
MN

OP‡£¤„QR¦

o—˜jL™£¤„p
q`aJœ^uh¦

Ž6x‘B`aJ8
Jiu#$í}~‚ƒ„”•

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q`aJu34Œ˜jL™uS€

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¦]Ⅰ,Ⅱ,Ⅲ,Ⅳ)
犘狅犺犾犻犪犾狌狋犲狊犮犲狀狊 (Limpr.)H.Lindb.,Acta
Soc.Sc.Fenn.1899;Crosby犲狋犪犾.,ChecklistMosses
191.1999;Smith,MossFl.Britain&Ireland.610-
611.2004;Li,Fl.Bryo.Sin.4:26.2006;Li犲狋犪犾.,
MossFl.China4:82.2007.———犠犲犫犲狉犪犾狌狋犲狊犮犲狀狊
Limpr.,Laubm.Deutschl.2:270.272.1892.———
犕狀犻狅犫狉狔狌犿犾狌狋犲狊犮犲狀狊(Limpr.)Loesk.,Stud.Morph.
Syst.Laubm.124.1910.
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Bgf•h

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m¦n

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T1 45678U3VWXV<=>?YZ
Table1 Thecomparisonofmorphologicalcharactersamongthe犘.犾狌狋犲狊犮犲狀狊andthreerelatedspecies
}~
Character
pq`aJ
犘.犾狌狋犲狊犮犲狀狊
‘B`aJ
犘.犮狉狌犱犪
’d`aJ
犘.犺狔犪犾狅狆犲狉犻狊狋狅犿犪
“`aJ
犘.狀狌狋犪狀狊
#$íPlants ldeGlossy ldeGlossy ldeGlossy ydeDul
¼^Á½wyr
Shapeofmedianleafcels
Ryð™šy
Linearrhomboidalor
narrowlyvermicular
Ryð™šy
Linearrhomboidalor
narrowlyvermicular
Ryð™šy
Linearrhomboidalor
narrowlyvermicular
¿ZŽyð[y
Elongatehexagonal
orrhombic
½wsCelwal xsThinwaled xsThinwaled xsThinwaled %s Thickwaled
·ÊíSporophyte ©ªAbsent ûªCommonlypresent ûªCommonlypresent ûªCommonlypresent
¸dPeristome ©ªAbsent l}Papilose \Smooth l}Papilose
y vw Asexualgem
mae
noB

pqr
Toruloidonrhizoids yAbsent yAbsent yAbsent
[:
!"#$%&()*
(HIMC)+*,-./01234,567*89:;<=>?@!A(KUN)BCD3
4/EFGHIJ=>?
(IFP)KL34M@!=>NOPQRS,MTUVWXYZ。
]^!P

[1] ÔÕÖ.^Ji#$bÈ(Ô])[M]."^:+%Ó]Ð,
1963:173227.
[2] ÛÜÝ.^IJÈ(,E-)[M]."^:+%Ó]Ð,2006:
835.
[3] LIXJ,CROSBYMR,HES.MossFloraofChina(Vol.4)
[M].SciencePress,Bejing & MissouriBotanicalGarden
Press,SaintLouis,2007:6788.
[4] COXCJ,HEDDERSONTAJ.Phylogeneticrelationshipsa
mongtheciliatearthrodontousmosses:evidencefromchloro
plastandnuclearDNAsequences[J].犘犾犪狀狋犛狔狊狋犲犿犪狋犻犮狊and
犈狏狅犾狌狋犻狅狀,1999,215:119139.
[5] GOFFINETB,COXCJ,犲狋犪犾.Thebryophyta(mosses):
systematicandevolutionaryinferencesfroman狉狆狊4gene(cp
DNA)phylogeny[J].犃狀狀犪犾狊狅犳犅狅狋犪狀狔.2001,87:191208.
[6] CROSBYMR,MAGILLRE,ALLENB,犲狋犪犾.AChecklist
oftheMosses[EB/OL].St.Louis:MissouriBotanicalGar
den.(19991220)[20131120].http://www.mobot.org/
MOBOT/tropicos/most/checklist.shtml
[7] FREY W,STECH M,犲狋犪犾.SylabusofPlantFamilies[M].
Berlin:Gebr.BorntraegerVerlagsbuchhandlung,2009.195198.
[8] _ `,a b.IJ#$[M]//ª^B$$x²³ (,k
-
)."^:+%Ó]Ð,2013:134144.
[9] HILLMO,BELLN,BRUGGEMANNANNENGAMA,犲狋
犪犾.AannotatedchecklistofthemossesofEuropeandMacaro
nesia[J].犑狅狌狉狀犪犾狅犳犅狉狔狅犾狅犵狔,2006,28:198267.
[10] SHAWAJ.Mielichhoferiaceae:BryophytefloraofNorthA
merica.Version1[EB/OL].(20090323)[20131120].ht
tp://www. mobot. org/plantscience/bfna/V2/MielMieli
chhoferiaceae.htm
2521 ! " # $ % &                   36-
[11] SHAWAJ,RAMSAYHP.AustralianMossesOnline.69.
Mielichhoferiaceae:Pohlia[EB/OL].(20130502)[2013
1120].http://www.anbg.gov.au/abrs/Mosses_online/
Mielichhoferiaceae_Pohlia.pdf
[12] SMITHAJE.TheMossFloraofBritainandIreland[M].
Cambridge:CambridgeUniversityPress,2004:593611.
[13] 3~4,cde,8.78xÊ#$+uh iref
[J].GH#$34,2003,25(3):245247.
WUZY,ZHOUZK,犲狋犪犾.Thearealtypesoftheworldfamilies
ofseedplants[J].犃犮狋犪犅狅狋犲狀犻犮犪犢狌狀狀犪狀犻犮犪.2003,25(3):
245247.
  N_Ⅰ pq`aJ
1.21^(‚f");2.Bg^(~I");3.Bg^(‚f");4.ikÜl;5.noBvw。(1.TanBC952081;2.Ç
ȏ6665;3~5.>?@201307102)
犘犾犪狋犲Ⅰ 犘狅犺犾犻犪犾狌狋犲狊犮犲狀狊(Limpr.)H.Lindb.
Fig.1.Vegetativeplant;Fig.2.Generativeplantwhendry;Fig.3.Generativeplantwhenmoist;Fig.4.Transverse
sectionofstem;Fig.5.Gemmaegrowingontherhizoid.(1.TanBC952081;2.WEIZhiping6665;35.WANGXiaorui
201307102)
35216+            >?@,8:pq`aJ(cdJ+)yz}~ŒŽœ^u¨h
  N_Ⅱ pq`aJ
1~7.¼;8~11.œž¼;12.¼½w;13.¼^Á½w;14.¼,Á½w;15.¼^ÁkÜl。(>?@201307102)
犘犾犪狋犲Ⅱ 犘狅犺犾犻犪犾狌狋犲狊犮犲狀狊(Limpr.)H.Lindb.
Fig.1-7.Leaves;Fig.8-11.perichaetialleaves;Fig.12.Apicalleafcels;Fig.13.Middleleafcels;Fig.14.Basalleafcels;Fig.
15.Transversesectionofmiddleleaf.(WANGXiaorui201307102)
4521 ! " # $ % &                   36-
  N_ Ⅲ pq`aJ
1.21^;2.¼½w;3.¼^Á½w;4.¼,Á½w;5.ikÜl;6.¼^ÁkÜl;7.no;8~9.noBvw;10.¼

>?@201307102)
犘犾犪狋犲Ⅲ 犘狅犺犾犻犪犾狌狋犲狊犮犲狀狊(Limpr.)H.Lindb.
Fig.1.Vegetativeplant;Fig.2.Apicalleafcels;Fig.3.Middleleafcels;Fig.4.Basalleafcels;Fig.5.Transversesec
tionofstem;Fig.6.Transversesectionofmiddleleaf;Fig.7.Rhizoid;Fig.8-9.Gemmaegrowingontherhizoid;Fig.10.
Leaves(WANGXiaorui201307102)
55216+            >?@,8:pq`aJ(cdJ+)yz}~ŒŽœ^u¨h
  N_ Ⅳ pq`aJœ^uh¦
犘犾犪狋犲Ⅳ Thedistributionmapof犘狅犺犾犻犪犾狌狋犲狊犮犲狀狊inChina

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6521 ! " # $ % &                   36-