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Natural Succession and Characteristics of Pine Community after the Invasion of Bursaphelenchus xylophilus in Zhoushan Islands of Zhejiang Province

浙江舟山岛松材线虫入侵后松林群落的自然演替和特征


对舟山岛21个松材线虫病入侵前的马尾松林、黑松林进行典型样地调查,根据更新层树种的重要值采用模糊聚类结果划分为7种演替类群,与17年后对不同演替类群具代表性样地的复查结果比较分析,并综合群落的组成结构、动态特征、树种生态生物学特性、更新树种植株结构以及立地条件、海拔等因子,确定舟山岛松林近阶段的演替结果分别为白栎、枫香占优势的落叶阔叶林,枫香、白栎、苦槠或石栎、四川山矾、栓皮栎占优势的常绿落叶阔叶混交林,马尾松、黄檀、山合欢占优势的针阔混交林,枫香或朴树占优势的落叶阔叶林(疏林),白栎占优势的萌生灌丛和柃木占优势的常绿灌丛。研究结果说明松材线虫病发生迹地的植物群落自然演替具有一定的方向性和确定性。可针对松材线虫病为害后林分不同的演替趋势、立地条件和培育目的,应用本文的预测和林相修复技术及早采取封育、人工造林等措施恢复植被; 同时,依据本文总结的演替规律,通过前期的调查和分类,可以较快推测和诊断出演替的过程和结果,从而及早在松材线虫病大面积发生后,根据不同的演替方向进行人工干预和规划。

Based on an investigation conducted between 1991-1992 in 21 Masson (Pinus massoniana) and Black pine (Pinus thunbergii) plots unattacked by pine wood nematode (PWN) in Zhoushan islands of Zhejiang province, the 21 plots were divided into 7 types of succession by SPSS fuzzy clustering with the importance values of regeneration tree species. A comparative analysis was conducted 17 years later between the previous investigated data and a set new survey data collected between 2007-2008 in representative plots of 7 types of succession, integrated with tree species composition, dynamic characteristics, eco-biological characteristics of pine, plant structure, soil, elevation and other factors. Thus, recent pine forests were developed into 8 vegetation types, including deciduous broadleaved forest dominated by Quercus fabric, Liquidambar formosana; evergreen mixed with deciduous broadleaved forest dominated by L. formosana, Q. fabric and Castanopsis sclerophylla or Lithocarpus glaber, Symplocos setchuensis and Q. variabilis; conifer mixed with broadleaved forest dominated by Pinus massoniana, Dalbergia hupeana and Albizia kalkora; deciduous broadleaved forest with sparse density which is dominated by L. formosana or Celtis sinensis; sprouted shrub community dominated by Q. fabric and evergreen shrub community occupied by Eurya japonica etc.. The results showed that the natural succession direction of PWN slash could be defined and judged to some extent. Thus, according to different succession trends, site conditions and cultivation purposes, restoring measures, such as closing forest, artificial afforestation and some other ways, should be adopted as soon as possible. Simultaneously, in the future forestry management, through the papers finding and the former investigation and classification, the succession process and sequence could be deduced and diagnosed quickly. Consequently human intervening and programming could be applied based on different succession directions as soon as possible after the pine wilt disease occurred.


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