Abstract:There are few rough endoplasmic reticulum (RER) pocket-like units which have large amount of coated vesicles (100-150nm) in the hydration pollen grain and pollen tube of buckwheat (Fagopyrum esculentum Monch.).In the newly -formed pollen tube, the coated vesicles mainly come from the pollen grain. The coated vesicles in the growing pollen tube are secreted from golgi body. And the breakage of RER is another resource of coated vesicles in the growing pollen tube. In the pollen tube of buckwheat, the function of coated vesicles which comtain the precursor of pollen tube wall is to form pollen tube besides decomposing lipid bodies and amyloplast. The decomposing sustance of stigma and stye cells play an important role in the growth of pollen tube of buckwheat.