全 文 ：生物多样性 2016, 24 (6): 714–718 doi: 10.17520/biods.2016071
Biodiversity Science http: //www.biodiversity-science.net

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收稿日期: 2016-03-14; 接受日期: 2016-06-06
 通讯作者 Author for correspondence. E-mail: nhxia@scib.ac.cn
中国蔷薇科、荨麻科和壳斗科植物新组合
童毅华 夏念和*
(中国科学院华南植物园, 广州 510650)
New combinations of Rosaceae, Urticaceae and Fagaceae from China
Yihua Tong, Nianhe Xia*
South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650

笔者在编研《中国生物物种名录》第一卷第五
分册(蔷薇科至叶下珠科)的过程中, 根据最近的研
究结果, 对《中国植物志》及Flora of China中一些
属的概念做了调整, 有22个名称需要进行重新组
合, 现报道如下。
1 蔷薇科(Rosaceae)
广 义 的 李 属 (Prunus s.l.) 包 括 了 桃 属
(Amygdalus)、杏属(Armeniaca)、樱属(Cerasus)、桂
樱属(Lauro-cerasus)、稠李属(Padus)及狭义的李属
(Prunus s.s.)等多个属。最近的分子系统学研究支持
广义李属的单系性质, 但樱属、桂樱属、稠李属等
属却都不是单系类群, 各属之间的关系还不十分清
楚(Potter et al, 2007; Chin et al, 2010, 2014; Shi et al,
2013)。在广义李属内, 传统分类使用的形态性状并
不能很好地区分各属或者各亚属, 给鉴定造成了困
扰, 以致两三百年来不同学者对于这一类群的系统
发育关系有着不同的看法, 分类意见一直不能统一
(Kalkman, 1965; Bortiri et al, 2001, 2006)。但相对于
分为桃属、杏属、樱属、桂樱属、稠李属和狭义李
属等多个属的处理来说, 有更多证据支持广义的李
属概念, 因此Flora Europaea (Webb, 1978)、Flora of
North America (Phipps, 2014)以及国内很多地方志
如《西藏植物志》(俞德浚等, 1985)、《贵州植物志》
(陈谦海, 1989)、《福建植物志》(张永田, 1985)、《广
东植物志》(陈伟球, 2000)、Flora of Hong Kong (Xia
& Deng, 2008)等志书也都使用广义李属。有鉴于此,
我们在编研《中国生物物种名录》的时候也沿用广
义李属的概念, 故有下列20个名称需组合至李属。
(1)陕梅杏
Prunus armeniaca L. var. meixianensis (J. Y.
Zhang, T. Z. Li, X. J. Li & Y. He) Y. H. Tong & N.
H. Xia, comb. nov.
Basionym: Armeniaca vulgaris Lam. var.
meixianensis J. Y. Zhang, T. Z. Li, X. J. Li & Y. He,
Bull. Bot. Res., Harbin 9(3): 66, f. 3 (1989). Type:
China, Shaanxi Province, Meixian, alt. 700 m, 15 Jun.
1982, J. Y. Zhang et al. B21104 (holotype, Herbarium
of Liaoning Institute of Pomology).
(2)熊岳大扁杏(熊岳杏)
Prunus armeniaca L. var. xiongyueensis (T. Z.
Li, J. Y. Zhang, X. J. Li & Y. He) Y. H. Tong & N. H.
Xia, comb. nov.
Basionym: Armeniaca vulgaris Lam. var.
xiongyueensis T. Z. Li, J. Y. Zhang, X. J. Li & Y. He,
Bull. Bot. Res., Harbin 9(3): 65, f. 2 (1989). Type:
China, Liaoning Province, Xiongyue, alt. 22 m, 25
Jul. 1985, T. Z. Li et al. C21201 (holotype, Herbarium
of Liaoning Institute of Pomology).
(3)志丹杏
Prunus armeniaca L. var. zhidanensis (C. Z.
Qiao & Y. P. Zhu) Y. H. Tong & N. H. Xia, comb.
nov.
Basionym: Armeniaca zhidanensis C. Z. Qiao &
·生物编目·
第 6期 童毅华和夏念和: 中国蔷薇科、荨麻科和壳斗科植物新组合 715
Y. P. Zhu, Acta Phytotax. Sin. 31(2): 188 (1993).
–Armeniaca vulgaris Lam. var. zhidanensis (C. Z.
Qiao & Y. P. Zhu) L. T. Lu, Acta Phytotax. Sin.
38(3): 281 (2000). Type: China, Shaanxi Province,
Zhidan County, Taipingshan, C. Z. Qiao & Y. P. Zhu
890176 (holotype, Herbarium of the Second Military
Medical University).
(4)武夷红樱
Prunus campanulata Maxim. var. wuyiensis (X.
R. Wang, X. G. Yi & C. P. Xie) Y. H. Tong & N. H.
Xia, comb. nov.
Basionym: Cerasus campanulata (Maxim.) A.
N. Vassiljeva var. wuyiensis X. R. Wang, X. G. Yi &
C. P. Xie, Acta Bot. Yunnan. 29(6): 616, f. 1 (2007).
Type: China, Fujian Province, Wuyishan, alt. 900 m,
on forests evergreen, 6 Mar. 2006, X. G. Yi 30604
(holotype, NF).
(5)华仁杏
Prunus cathayana (D. L. Fu, B. R. Li & J. Hong
Li) Y. H. Tong & N. H. Xia, comb. nov.
Basionym: Armeniaca cathayana D. L. Fu, B. R.
Li & J. Hong Li, Bull. Bot. Res., Harbin 30: 1 (2010).
Type: China, Hebei Province, Zhuolu County, 11 Jul.
2008, D. L. Fu 2008071101 (holotype, CAF).
(6)长叶桂樱
Prunus dolichophylla (T. T. Yu & L. T. Lu) Y. H.
Tong & N. H. Xia, comb. nov.
Basionym: Lauro-cerasus dolichophylla T. T.
Yu & L. T. Lu, Bull. Bot. Res., Harbin 4(4): 50, f. 2
(1984). Type: China, Yunnan Province, Xichou
County, alt. 1,300–1,500 m, 24 Sep. 1947, K. M. Feng
11997 (holotype, PE barcode no. 00004582!).
(7)新疆桃(吕宛桃, 大宛桃)
Prunus ferganensis (Kostina & Rjabov) Y. Y.
Yao ex Y. H. Tong & N. H. Xia, comb. nov.
Basionym: Prunus persica (L.) Batsch subsp.
ferganensis Kostina & Rjabov, Trudy Prikl. Bot., Ser.
8, Polodovye Jagodnye Kul’t. 1: 323, f. 27 II (1932).
–Persica ferganensis (Kostina & Rjabov) Kovalev &
Kostina, Trudy Prikl. Bot., Ser. 8, Polodovye Jagodnye
Kul’t. 4: 75 (1935). –Amygdalus ferganensis (Kostina
& Rjabov) T. T. Yu & L. T. Lu, Fl. Reipubl. Popularis
Sin. 38: 20 (1986). –Prunus ferganensis (Kostina &
Rjabov) Y. Y. Yao, Fl. Desert. Reipubl. Popul. Sin. 2:
158 (1987), nom. inval. Type: unknown.
姚育英(1987)在发表新组合Prunus ferganensis
(Kostina & Rjabov) Y. Y. Yao时未引证基名Prunus
persica subsp. ferganensis Kostina & Rjabov的文献
出处, 而只引证了基于该名称的另一组合Persica
ferganensis (Kostina & Rjabov) Kovalev & Kostina的
文献出处。根据International Code of Nomenclature
for Algae, Fungi, and Plants (Melbourne Code)条款
41.5的规定(McNeill et al, 2012), 姚育英(1987)没有
清楚地指出其基名, 因此其组合为不合格发表。
(8)鹤峰樱
Prunus hefengensis (X. R. Wang & C. B. Shang)
Y. H. Tong & N. H. Xia, comb. nov.
Basionym: Cerasus hefengensis X. R. Wang &
C. B. Shang, Ann. Bot. Fennici 44: 151 (2007). Type:
China, Hubei Province, Hefeng County, alt.
800–1,000 m, roadside on edge of forest, 11 Apr.
1996, X. R. Wang 96069 (holotype, NF).
(9)洪平杏
Prunus hongpingensis (T. T. Yu & C. L. Li) Y. H.
Tong & N. H. Xia, comb. nov.
Basionym: Armeniaca hongpingensis T. T. Yu
& C. L. Li, Acta Phytotax. Sin. 23(3): 209, pl. 1, f. 1
(1985). Type: China, Hubei Province, Hongping
County, alt. 1,800 m, 31 Jul. 1977, Shennongjia Ex-
ped. 34031 (holotype, HIB barcode no. 0096980!).
(10)钝核甘肃桃
Prunus kansuensis Rehder var. obtusinucleata
(Y. F. Qu, X. L. Chen & Y. S. Lian) Y. H. Tong & N.
H. Xia, comb. nov.
Basionym: Amygdalus kansuensis (Rehder)
Skeels var. obtusinucleata Y. F. Qu, X. L. Chen & Y.
S. Lian, Acta Bot. Boreal.-Occident. Sin., 29(6): 1281
(2009). Type: China, Gansu Province, Kangxian, alt.
1,500 m, 1 Aug. 2008, Y. S. Lian et al. 080024 (holo-
type, NWTC).
(11)李梅杏
Prunus limeixing (J. Y. Zhang & Z. M. Wang) Y.
H. Tong & N. H. Xia, comb. nov.
Basionym: Armeniaca limeixing J. Y. Zhang &
716 生 物 多 样 性 Biodiversity Science 第 24卷
Z. M. Wang, Acta Phytotax. Sin. 37(1): 107, f. 2
(1999). Type: China, Liaoning Province, Xiongyue,
in cultivation, 15 Aug. 1996, J. Y. Zhang et al. 96-2
(holotype, Herbarium of Liaoning Institute of Pomol-
ogy; isotype, PE barcode no. 01790027!).
(12)北亚稠李
Prunus padus L. var. asiatica (Kom.) Y. H.
Tong & N. H. Xia, comb. nov.
Basionym: Padus asiatica Kom., Fl. URSS 10:
578 (1941). –Padus racemosa (Lam.) Gilib. var. asi-
atica (Kom.) T. T. Yu & T. C. Ku, Fl. Reipubl. Popu-
laris Sin. 38: 98 (1986). –Padus avium var. asiatica
(Kom.) T. C. Ku & B. M. Barthol., Fl. China 9: 423
(2003). Type: unknown.
(13)泰山野樱
Prunus serrulata Lindl. var. taishanensis (Yi
Zhang & C. D. Shi) Y. H. Tong & N. H. Xia, comb.
nov.
Basionym: Cerasus serrulata (Lindl.) Loudon
var. taishanensis Yi Zhang & C. D. Shi, Acta Phyto-
tax. Sin. 37(1): 87 (1999). Type: China, Shandong
Province, Tai’an City, Mt. Taishan, alt. 1,000 m, 29
May 1998, Y. Zhang & C. D. Shi 98001 (holotype,
Herbarium of Shandong Institute of Pomology; iso-
type, PE barcode no. 01438567!).
(14)辽海杏
Prunus sibirica L. var. pleniflora (J. Y. Zhang,
T. Z. Li, X. J. Li & Y. He) Y. H. Tong & N. H. Xia,
comb. nov.
Basionym: Armeniaca sibirica (L.) Lam. var.
pleniflora J. Y. Zhang, T. Z. Li, X. J. Li & Y. He,
Bull. Bot. Res., Harbin 9(3): 65, f. 1 (1989). Type:
China, Liaoning Province, Beipiao, Daheishan, alt.
800 m, 10 Apr. 1981, J. Y. Zhang et al. B10401 (hol-
otype, Herbarium of Liaoning Institute of Pomology).
(15)仙居杏(杏梅)
Prunus xianjuxing (J. Y. Zhang & X. Z. Wu) Y.
H. Tong & N. H. Xia, comb. nov.
Basionym: Armeniaca xianjuxing J. Y. Zhang &
X. Z. Wu, Bull. Bot. Res., Harbin 29(1): 1 (2009).
Type: China, Zhejiang Province, Xianju County,
Baita, Mt. Kuocangshan, alt. 50–500 m, 16 May 2008,
J. Y. Zhang et al. 2008-1 (holotype, Herbarium of
Liaoning Institute of Pomology).
(16)雪落樱
Prunus xueluoensis (C. H. Nan & X. R. Wang)
Y. H. Tong & N. H. Xia, comb. nov.
Basionym: Cerasus xueluoensis C. H. Nan & X.
R. Wang, Ann. Bot. Fennici 50: 79 (2013). Type:
China, Hubei Province, Enshi Tujia and Miao Au-
tonomous Prefecture, Xuan’en County, Xueluozhai
Mt., in alpine shrubbery beside road, alt. 1,430 m, 3
Apr. 2009, C. H. Nan 040301 (holotype, NF).
(17)姚氏樱(西藏樱桃)
Prunus yaoiana (W. L. Zheng) Y. H. Tong & N.
H. Xia, comb. nov.
Basionym: Cerasus yaoiana W. L. Zheng, Acta
Phytotax. Sin. 38(2): 195, pl. 1 (2000). Type: China,
Xizang Province, Nyingchi City, alt. 2,950 m, in for-
ests, 4 Jun. 1989, G. Yao et al. 1152 (holotype, Her-
barium of Xizang Institute of Plateau Ecology; iso-
types, PE barcode no. 01821794! & no. 01821795!).
(18)政和杏
Prunus zhengheensis (J. Y. Zhang & M. N. Lu)
Y. H. Tong & N. H. Xia, comb. nov.
Basionym: Armeniaca zhengheensis J. Y. Zhang
& M. N. Lu, Acta Phytotax. Sin. 37(1): 105 (1999).
Type: China, Fujian Province, Zhenghe County,
Waitun, Mt. Chouling, alt. 780–940 m, 17 Jul. 1996,
J. Y. Zhang et al. 96-1 (holotype, Herbarium of
Liaoning Institute of Pomology; isotype, PE barcode
no. 01790028!).
2 荨麻科(Urticaceae)
陈家瑞(1982)发表了疣果冷水花(Pilea verru-
cosa Hand.-Mazz.)的两个亚种闽北冷水花(P. ver-
rucosa Hand.-Mazz. subsp. fujianensis C. J. Chen)和
离基脉冷水花(P. verrucosa Hand.-Mazz. subsp. sub-
triplinervia C. J. Chen)。Chen和Monro (2003)将纤细
冷水花(P. gracilis Hand.-Mazz.)并入P. verrucosa
Hand.-Mazz., 而且将闽北冷水花和离基脉冷水花
作为变种处理, 并引证了它们的原始文献。根据
International Code of Nomenclature for Algae, Fungi,
and Plants (Melbourne Code)条款 41.6的规定
第 6期 童毅华和夏念和: 中国蔷薇科、荨麻科和壳斗科植物新组合 717
(McNeill et al, 2012), Chen和Monro (2003)无意中合
格发表了两个新等级中的名称(name at new rank),
即Pilea verrucosa Hand.-Mazz. var. fujianensis (C. J.
Chen) C. J. Chen & Monro和 Pilea verrucosa
Hand.-Mazz. var. subtriplinervia (C. J. Chen) C. J.
Chen & Monro。但随后Chen等(2007)发现P. verru-
cosa Hand.-Mazz.是P. verrucosa Killip (1925)的晚出
同 名 , 于 是 接 受 纤 细 冷 水 花 (P. gracilis
Hand.-Mazz.)为这个实体的正确名称, 将P. verru-
cosa Hand.-Mazz. (1929)处理为纤细冷水花的异名。
通过文献查阅和标本研究, 我们也认同这两个名称
实为同种, 接受P. gracilis Hand.-Mazz.为正确名称,
但由于Chen等(2007)未对原先发表在P. verrucosa之
下的两个亚种进行处理, 现因编辑名录所需, 在此
重新组合。
(19)闽北冷水花
Pilea gracilis Hand.-Mazz. subsp. fujianensis (C.
J. Chen) Y. H. Tong & N. H. Xia, comb. nov.
Basionym: Pilea verrucosa Hand.-Mazz. subsp.
fujianensis C. J. Chen, Bull. Bot. Res., Harbin 2(3): 55
(1982). –Pilea verrucosa Hand.-Mazz. var. fujianensis
(C. J. Chen) C. J. Chen & Monro, Fl. China 5: 101
(2003). Type: China, Fujian Province, Jianning
County, Wangpingdong, alt. 1,000 m, 3 Jun. 1978, Z.
Y. Li 10880 (holotype, PE barcode no. 00023976!).
(20)离基脉冷水花
Pilea gracilis Hand.-Mazz. subsp. subtripliner-
via (C. J. Chen) Y. H. Tong & N. H. Xia, comb. nov.
Basionym: Pilea verrucosa Hand.-Mazz. subsp.
subtriplinervia C. J. Chen, Bull. Bot. Res., Harbin
2(3): 56 (1982). –Pilea verrucosa Hand.-Mazz. var.
subtriplinervia (C. J. Chen) C. J. Chen & Monro, Fl.
China 5: 101 (2003). Type: China, Hainan Province,
Changjiang County, Qicha Xiang, Qichaling, alt.
400–600 m, 27 Nov. 1956, S. H. Chen 11391 (holo-
type, IBSC barcode no. 0001393!).
3 壳斗科(Fagaceae)
广 义 栎 属 (Quercus s.l.) 常 包 括 青 冈 属
(Cyclobalanopsis)和狭义栎属(Quercus s.s.)。最近大
多数的系统学证据表明, 广义的栎属是一个自然的
单系类群, 青冈属也是单系类群, 但去除了青冈属
的栎属是一个并系类群(Manos et al, 2001; Oh &
Manos, 2008; Hubert et al, 2014)。Hubert等(2014)还
对近年来广义栎属的系统发育研究进行了总结, 发
现不同外类群的选取会影响青冈类群的系统位置,
但这些研究均支持广义栎属的分类处理。有鉴于此,
在编研《中国生物物种名录》时, 我们采用广义栎
属的概念将以下2个仅在青冈属中发表的名称组合
至栎属。
(21)昌化岭青冈
Quercus changhualingensis (G. A. Fu & X. J.
Hong) N. H. Xia & Y. H. Tong, comb. nov.
Basionym: Cyclobalanopsis changhualingensis
G. A. Fu & X. J. Hong, Guihaia, 27(1): 29 (2007), as
‘changhuaglingensis’. Type: China, Hainan Province,
Changjiang County, Changhualing, alt. 180–220 m, 4
Oct. 2004, G. A. Fu 11283 (holotype, HFB).
(22)燕千青冈
Quercus yanqianii (G. A. Fu) N. H. Xia & Y. H.
Tong, comb. nov.
Basionym: Cyclobalanopsis yanqianii G. A. Fu,
Bull. Bot. Res., Harbin, 27(1): 1 (2007), as
‘yin-qianii’. Type: China, Hainan Province, Changji-
ang County, Changhualing, alt. 160–200 m, 5 Oct.
2004, G. A. Fu 11287 (holotype, HFB).
符国瑷(2007)在文中明确指出本种的种加词是
为了纪念我国林学家华南农业大学的徐燕千教授,
按照现代汉语的拼音规则“燕千”的正确拼法是
“yanqian”, “yinqian”属于印刷错误, 需要更正, 且
根据International Code of Nomenclature for Algae,
Fungi, and Plants (Melbourne Code)条款60.9的规定
(McNeill et al, 2012), 连字符仅在组成加词的单词
通常独立存在或连字符前后的字母相同时使用, 因
此这里的连字符是应改正的错误, 故予以删除。
致谢: 感谢中国科学院华南植物园邓云飞研究员在
论文写作过程中提供的修改意见, 感谢几位审稿专
家的建议, 中国科学院植物研究所的杨拓博士帮助
查阅了部分种类的模式标本信息, 在此一并感谢。
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