Abstract:The Aquilapolles are found in Northeast and Eastern part of China. 183 speices are selected from the world. These 183 speices include Aquilapollenites, Mancicorpus, Fibulapollis and Translucentipollis. It is showed that they appeared from Albian of Early Cretaceous to Early Oligocene(?) of Tertiary. They mainly appeared in Senonian. The Aquilapolles are the most flourishing in Cretaceous during Maestrichtian. Tile six speices of which appeared in Tertiary. They are Aquilapollenites granobaculus, A. punctatus, A. quadrilobus, A. spinulosus, Funk., A. spinulosus (Mtched.) n. corab. and Fibulapollis mirificus. There are two phytogeoprovinces of the sporo-pollen in the Northern Hemisphere during Late Cretaceous. One is "Aquilapolles District "-moist subtropical climatic, Circum-Pacific and Siberia. Another is “Normapolles District”-arid (or semiarid) sub- tropical climatic, Circum-Atlantic, Europ and India. The author related the process to form the two phytogeopovinees by distribution of continent and sea in that time. In one country, Tertiary Aquilapollenites is wider than Late Cretaceous Aquilapollenites in palaeogeographic range. The Tertiary speices of Aquilapollenites are Aquila-pollenites spinulosus Funk. and A. granobaculus Song in China. Their kinds, amount and distribution fully show an evidence of relict member. They are relict Aquilapollenires and are not recycled fossils .from Upper Cretaceous strata. On Aquilapollenites, the respective characters of the growth and distribution of each Tertiary Epochs are indicated. The author also explores the relations between the growth, distribution and evolution of paleoenvironment in this paper.