全 文 ：植 物 分 类 学 报 44 (4): 454–456(2006) doi:10.1360/aps050140
Acta Phytotaxonomica Sinica http://www.plantsystematics.com
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2005-09-16收稿, 2006-03-06收修改稿。
基金项目: 贵州省中药现代化资助项目(200308)(Supported by the project of the Modernization of Materia Medica of Guizhou
Province, Grant No. 200308)。
贵州金盏苣苔属(苦苣苔科)一新种——万山金盏苣苔
何顺志
(贵阳中医学院药学系 贵阳 550002)
Isometrum wanshanense, a new species of the
Gesneriaceae from Guizhou, China
HE Shun-Zhi
(Department of Pharmacy, Guiyang College of Traditional Chinese Medicine, Guiyang 550002, China)
Abstract Isometrum wanshanense S. Z. He, a new species of the Gesneriaceae from
Wanshan County, Guizhou, China, is described and illustrated. This species is related to I.
villosum K. Y. Pan in having peduncle and pedicel subglabrous to brown villous, corolla
thinly cylindric, purple, not contracted at the throat, adaxial lip longer than abaxial one, and
stamens and pistil glabrous, but differs by having leaves petiolate, petiole 0.4–1.4 cm long,
leaf blade oblanceolate, 1.5–4×0.5–1.6 cm, cymes 1–4-flowered, calyx segments glabrous
outside or sparsely brown villous near the apex, and thecae confluent. The genus is endemic
to China, and is recorded in Guizhou Province for the first time.
Key words Isometrum, Isometrum wanshanense S. Z. He, Gesneriaceae, new species,
Guizhou, China.
摘要 报道了贵州金盏苣苔属Isometrum一新种, 即万山金盏苣苔I. wanshanense S. Z. He。该种以花序
梗、花梗被褐色长柔毛至近无毛, 花冠细筒状, 紫色, 喉部不缢缩, 上唇稍长于下唇, 雄蕊及雌蕊均无
毛而与柔毛金盏苣苔I. villosum K. Y. Pan 相近缘, 但叶具柄, 柄长0.4–1.4 cm, 叶片较小, 倒披针形, 长
1.5–4×0.5–1.6 cm, 聚伞花序具1–4朵花, 花萼裂片外面近无毛或近顶端疏被褐色长柔毛, 花柱与子房
近等长而不同。
关键词 金盏苣苔属; 万山金盏苣苔; 苦苣苔科; 新种; 贵州; 中国
作者在承担贵州省中药现代化重大项目“贵州中药资源种类的订正与增补研究”的
过程中, 于2003年4月对万山县的中药资源进行了考察, 在万山县城城东十余华里处石灰
岩山的岩石上发现数十株开紫色花、十分美丽的苦苣苔科Gesneriaceae金盏苣苔属
Isometrum Craib植物。该植物与金盏苣苔属已知种类(Pan, 1986, 1990; Pan & Liu 1995;
Wang et al., 1998; Li & Wang, 2004)不同, 是一个新种。该属在贵州也是首次记录。
万山金盏苣苔 新种 图1
Isometrum wanshanense S. Z. He, sp. nov. Fig. 1
Affine I. villoso K. Y. Pan, a quo petiolis 0.4–1.4 cm longis, foliis oblanceolatis, 1.5–4 cm
longis, 0.5–1.6 cm latis, cymis 1–4-floris, calycis segmentis extus subglabris vel apice sparse
brunneo-villosis, loculis antherarum confluentibus, stylo ovario aequilongo differt.
Herba perennis. Rhizoma circ. 0.7 cm longum, circ. 0.4 cm diam. Folia omnia basalia,
4期 何顺志: 贵州金盏苣苔属(苦苣苔科)一新种——万山金盏苣苔 455
图1 万山金盏苣苔 A, 体态; B, 花萼; C, 花冠展开; D, 花盘与雌蕊。(孙英宝、李爱莉根据何顺志030413号标本绘)
Fig. 1. Isometrum wanshanense S. Z. He. A, habit; B, calyx; C, corolla opened; D, disc and pistil. Drawn by Y. B. Sun
and A. L. Li from the type gathering, S. Z. He 030413 (PE).

petiolata; laminae oblanceolatae, 1.5–4 cm longae, 0.5–1.6 cm latae, apice rotundatae, basi
cuneatae, margine crenatae, utrinque brunneo-villosae, nervis lateralibus utrinsecus circ. 3,
supra obscuris, subtus prominulis, brunneo-villosis; petioli 0.4–1.4 cm longi, brunneo-villosi.
Cymae 1–4, axillares, 1–4-florae; pedunculi 4–7 cm longi, brunneo-villosi; bracteae 2,
lineares, 2.5–4 mm longae, brunneo-villosae; pedicelli 1.5–2.5 cm longi, sparse
brunneo-villosi. Calyx ad basin 5-sectus, segmentis lanceolatis, circ. 2.5 mm longis, circ. 0.5
mm latis, apice obtusis, margine integris, extus subglabris vel apice sparse brunneo-villosis,
intus glabris. Corolla tenuiter cylindrica, purpurea, circ. 2 cm longa, utrinque glabra, tubo circ.
1.5 cm longo, 2.5-plo longiore quam limbo, circ. 4 mm diam., labio postico circ. 6 mm longo,
ad medium 2-fido, lobis rotundatis circ. 4 mm longis, labio antico circ. 5 mm longo, prope
basin 3-secto, lobis omnis subrotundatis circ. 4 mm longis. Stamina 4, glabra, ea postica circ.
4 mm longa, ad circ. 7.5 mm supra corollae basin inserta, ea antica circ. 5 mm longa, ad circ. 8
mm supra corollae basin inserta, filamentis apice cum antheris sub angulo recto abrupte
植 物 分 类 学 报 44卷 456
curvatis, antheris per paria cohaerentibus 1 mm longis, loculis confluentibus. Staminodium 1,
circ. 1.2 mm longum, ad 3 mm supra corollae basin insertum. Discus annularis 2 mm altus.
Pistillum glabrum, 1.2–1.4 cm longum, ovario anguste oblongo, 6–7 mm longo, 1.2–1.5 mm
diam., stylo ovario aequilongo, stigmatibus 2, circ. 0.6 mm longis. Capsulae oblanceolatae,
glabrae. Fl. Apr.
China. Guizhou (贵州): Wanshan (万山), alt. 970 m, Karst hills, 2003-04-13, S. Z. He
(何顺志) 030413 (holotype, PE; isotype, HGCM).
多年生草本。根状茎长约0.7 cm, 直径约0.4 cm。叶全部基生, 具柄; 叶片倒披针形,
长1.5–4 cm, 宽0.5–1.6 cm, 顶端圆形, 基部楔形, 边缘具圆齿, 两面被褐色长柔毛, 侧脉
每边约3条, 上面不明显, 下面隆起, 被褐色长柔毛; 叶柄长0.4–1.4 cm, 被褐色长柔毛。
聚伞花序1–4条, 每花序具1–4朵花; 花序梗长4–7 cm, 被褐色长柔毛; 苞片2, 条形, 长
2.5–4 mm, 被褐色长柔毛; 花梗长1.5–2.5 cm, 疏被褐色长柔毛。花萼5裂至基部, 裂片披
针形, 长约2.5 mm, 宽约0.5 mm, 顶端钝, 全缘, 外面近无毛或仅顶端疏被褐色长柔毛,
内面无毛。花冠细筒状, 紫色, 长约2 cm, 内外面均无毛; 筒长约1.5 cm, 为檐部的2.5倍,
直径约4 mm, 上唇长约6 mm, 裂片圆形, 长约4 mm, 下唇长约5 mm, 3裂近基部, 裂片近
圆形, 长约4 mm。 雄蕊无毛, 上雄蕊长约4 mm, 着生于距花冠基部约7.5 mm处, 下雄蕊
长约5 mm, 着生于距花冠基部约8 mm处, 花药成对连着, 长1 mm, 药室汇合; 退化雄蕊
长约1.2 mm, 着生于距花冠基部约3 mm处。花盘高约2 mm。雌蕊无毛, 长1.2–1.4 cm, 子
房长6–7 mm, 直径1.2–1.5 mm, 花柱与子房等长, 柱头2, 长约0.6 mm。蒴果倒披针形, 无
毛。花期4月。
本种以花序梗及花梗被褐色长柔毛, 花冠细筒状, 紫色, 喉部不缢缩, 上唇稍长于下
唇, 雄蕊及雌蕊无毛而近似于柔毛金盏苣苔I. villosum K. Y. Pan, 但叶柄长0.4–1.4 cm, 叶
片倒披针形, 长1.5–4 cm, 宽0.5–1.6 cm, 聚伞花序具1–4朵花, 花萼外面近无毛或近顶端
疏被褐色长柔毛, 药室汇合, 花柱与子房等长而易于区别。在柔毛金盏苣苔, 叶无柄, 叶
片倒卵状长圆形, 长3–8 cm, 宽1.5–3.5 cm, 聚伞花序具8–10朵花, 花萼外面被褐色长柔
毛, 药室不汇合, 花柱短于子房。
本种目前仅在贵州万山县杉木垌发现, 生于海拔970 m的石灰岩山的岩石上。其生境
的土层较薄, 干燥, 伴生植物较少。分布的个体数量不多, 因此该种需要保护。
致谢 潘开玉研究员审阅全文并提出指导性意见, 李振宇和王印政研究员审阅并提出参
考意见, 孙英宝、李爱莉绘制插图, 在此一并致谢！
参 考 文 献
LI Z-Y (李振宇), Wang Y-Z (王印政)．2004. Plants of Gesneriaceae in China. Zhengzhou: Henan Science
and Technology Press. 51–59.
Pan K-Y (潘开玉)．1986. The second revision of the genus Isometrum (Gesneriaceae). Acta Botanica
Yunnanica (云南植物研究) 8: 23–36.
Pan K-Y (潘开玉)．1990. Isometrum Craib. In: Flora Reipublicae Popularis Sinicae (中国植物志)．Beijing:
Science Press. 69: 177–190.
Pan K-Y (潘开玉), Liu Z-Y (刘振宇)．1995. A new species of Isometrum Craib. Acta Phytotaxonomica
Sinica (植物分类学报) 33: 100–102.
Wang W-T (王文采), Pan K-Y (潘开玉), LI Z-Y (李振宇), Weitzman A L, Skog L E. 1998. Isometrum Craib.
In: Wu Z-Y (吴征镒), Raven P H eds. Flora of China. Beijing: Science Press; St. Louis: Missouri
Botanical Garden Press. 18: 263–267.