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The Sub cellular Chromium (Cr6+) Distribution and Tolerance Mechanism to Chromium Stress in Different Tolerant Brassica chinensis L. Cultivars

不同耐铬性青菜中Cr6+的亚细胞分布及其耐性机理



全 文 :书!"#$%&
,2016,36(5):0954-0963
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。Email:sunjianyun@
njau.edu.cn
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abcde% ;f+%%g

ab210095)
F G:KhijkLCr6+QRS^_`[]<,lm2nop`q(Cr)[jkrs‘tuv’(Cr`[rs)w

xyz
’(Cr3{rs)|}~€,‚ƒ0、10、50w100μmol·L
-1Cr6+„<…jk†‡ˆw‰ŠCr‹X、Cr
QŒŽ

‘’“”,
(NPT)、#$•–—(PCs)w˜™š›œ[Qž。Ÿ ¡i:(1)2njkrs†‡ˆ
Cr‹X¢£¤¥‰ŠQ‹X,¦‘tuv’‰ŠCr‹X、CrQ‰Š§¨©、†‡ˆw‰ŠŽª«Cr‹X¬­Cr
‹XQ‚®¯¢£e¥

xyz
’,
°_±²[Cr‹X¬­Cr‹XQ‚®¢£¤¥‘xyz’。(2)Cr6+„<¢£
³u´jk†‡ˆNPTwPCsQ‹X,¦‘tuv’³uQµ¶¢£·¥‘xyz’。(3)¸ ¹¶QCr6+(10μmol·
L-1)º»¼·jkSOD、PODwCATQœ[,¦‘tuv’Q³µe¥‘xyz’;°·¹¶QCr6+(100μmol·
L-1)¢£½¸´SOD、PODwCATQœ[,¦‘tuv’Q½µ¤¥‘xyz’。=>¾K,¿Cr6+ÀÁÂÅ,¬
jkrs

xyz

Ărs

tuv

ºÅÆÇQCr¥‰wŽª«,ÈÉ´ÊË|̍ŽÍQCr,ΦÏM
ƒ·Q˜™š›œ[wPCs‹X,9°¡ÐÑÆÒQ`Cr6+[。
HIJ

jk
;Cr6+;ŒŽ;‘’“”,;˜™š›
;K@L$
:Q945.78 !MNOP:A
犜犺犲犛狌犫犮犲犾狌犾犪狉犆犺狉狅犿犻狌犿(犆狉6+)犇犻狊狋狉犻犫狌狋犻狅狀犪狀犱犜狅犾犲狉犪狀犮犲犕犲犮犺犪狀犻狊犿
狋狅犆犺狉狅犿犻狌犿犛狋狉犲狊狊犻狀犇犻犳犳犲狉犲狀狋犜狅犾犲狉犪狀狋犅狉犪狊狊犻犮犪犮犺犻狀犲狀狊犻狊
犔.犆狌犾狋犻狏犪狉狊
ZHANGHaimin,YANGWenjia,WANGQingya,SUNJianyun
(ColegeofLifeScience,NanjingAgriculturalUniversity,Nanjing210095,China)
犃犫狊狋狉犪犮狋:Inordertoelucidateabsorption,distributionandtolerancemechanismsof犅狉犪狊狊犻犮犪犮犺犻狀犲狀狊犻狊L.
tochromium (Cr6+)stress,wegrewtwoBrassicacultivars;‘Meijiahua’(Cr6+stresstolerant)and
‘Sanyueman’(Cr6+stresssensitive)hydroponicalyinHoagland0,10,50and100μmol·L
-1Cr6+.The
differencesinCrabsorbedcontentsandsubcelulardistribution,nonproteinthiol,phytochelatins(PCs)and
antioxidantenzymeactivitiesinshootsandrootswerestudied.Theresultswereasfolows:(1)thecon
tentsofCrinshootsweresignificantlylowerthanthoseintherootsinbothcultivars.TheCrrootcon
tents,CrretentionrateandtheratioofwalboundedCrtototalCrinshootsandrootsof‘Meijiahua’
weresignificantlyhigherthanthosein‘Sanyueman’,buttheratioofwatersolubleCrtototalCrwasop
posite.(2)Cr6+treatmentsatalconcentrationssignificantlyincreasedtheconcentrationsofnonprotein
thiolandphytochelatins(PCs)inshootsofbothcultivars,buttheincrementin‘Meijiahua’wassignifi
cantlyhigherthanthatin‘Sanyueman’.(3)UnderlowconcentrationofCr6+(10μmol·L
-1),theactivi
tiesofSOD,PODandCATweresignificantlyincreased,buttheextentofincreasewassignificantlyhigher
in‘Meijiahua’thanthatin‘Sanyueman’.TheactivitiesofSOD,PODandCATweresignificantlydown
regulatedunderhighconcentrationofCr6+ (100μmol·L
-1),butthemagnitudeofdownregulationin
‘Meijiahua’wassignificantlylowerthanthatin‘Sanyueman’.TheseresultsindicatedthatunderCr6+
stress,ascomparedto‘Sanyueman’,moreCrwasdistributedincel walsinshootandrootin‘Meiji
ahua’toinhibitthetransmembraneCrtointracelularflow.Moreover,ithashigherantioxidantenzyme
activitiesandPCscontent,soitshowedastrongertolerancetoCr6+.
犓犲狔狑狅狉犱狊:犅狉犪狊狊犻犮犪犮犺犻狀犲狀狊犻狊L.;Cr6+;subcelulardistribution;nonproteinthiol;antioxidantenzyme.
  ÓÔÕÖ、×Ø、ÙښÛ、ÚÜwÝÞßÛd
Qàáâã

äå‹MeXæ-çq
(Cr)Qè~、
èéêÑ

åëìí

îï

~ðw;$ñòopó
¶QCrôØ[1]。Cr¿)*ÂÅMÇsõö,_
«Cr3+wCr6+÷Køùw÷Kúû。Cr6+N[‚
Cr3+·100ü,ýþsÿÒQå!、å"#$,%&
()*¾K3så+-ç$,þ[2]。¿)*-

,Cr6+ú/Cr2O72-w CrO42-012¿。WX
Crº½¸#$L~w34OPQRS,56#7;
8

9æ:±äå#7;<

Φ Cr±/?W=$
>?@ABCD
[34]。
jk
(犅狉犪狊狊犻犮犪犮犺犻狀犲狀狊犻狊L.)
EF¤“k

ý«eGšQHk,þ

¿HkIJ
KZ«LMæ8†M

%&

=>jkLCrQRS
/^`[]6ÏMæ8NO

Cr|Ì#$ðÍP,78QR¿#$‰ˆ,(
S6TUò#$VW

Xý#$LCrQRSwYZ
¿$swrs‡2¿¢£ž
[5]。
#$L CrQ[
\]^ù¥ŽªL CrQ_ùw`aL CrQb
š
[6]。
Žªýæ-ç|̍ŽÍQcþdef

æ-çgWŽª:

¬Žª‡QhiP

jhi
Pw kßel$mâ;š%\Z°(_ù…
n
[7]。
oæ-ç¿#$Žª‡pòqwP


ŽÍQæ-纬eXQM]rMð

•–$

’
“mß

Ÿ–0søtQ•–$

Î(Tuv`a«
wx

½¸ŽÍQ)yæ-çzlQ¹¶

9°½
¸_N[
[8]。
{|-ç}–ð78~M]€


,€^_‚;$

-烒“
(MTs)w#$•–—
(PCs)ß[9]。op#$sBVrsLæ-çQ`[
Mÿež

_Læ-çQ`[]6„M…op

†‡
,Cr¿_#$«QRS^_`[]<=>ƒ
Ç

°ˆ¥jkLCrQRS/^`[]oÇû

‰=>/`Cr[opQ2njkrsK
€Š

=>Cr¿jkðÍQŒŽwCr6+L
‘’“”,‹X

˜™š›œ[Q‹Œ

¿iŽ2
njkrsLCrQRS^;<ŒZQž,h#
$`Cr6+]<,K*@HkQl‘^’¼K<
“”•

1 ŠÞwI–
1.1 QRST
?W‡—˜‚ƒ´Cr6+L41njkrs™
$mæ56Qž

šlÑ2n`Cr6+[opQj
krs

tuv
’(Cr6+`[rs)w‘xyz’(Cr6+
3{rs
),
Î/{2nrsK€Š。jksl¯›
)./0cd+%g

1.2 QRUV
œl e ¤ þ å Q s l  5% NaClO ž N
5min,Ÿ ™¡PÅsl¯¢£¤¿™¡¥¦Q§
‡
,¨
Ì©ª4«¬©­â

®¯°¶±K
24℃w20℃。²“PÅsl³Ì´Mµ¶QI³
««·¸;8
,10dPœl8¹þåQº»¼#¿
‹5L34`Q4½«4,¾4½¼»20
¿

 4 ` K Hoagland 3 4 `。 À Á 1/
2Hoagland34`43d,ÂÁÃ Hoagland34
`4

ÄÅ?í

¾2dÆÆ1Ç34`,¾ÈÁ
HClÉÊpH ¿5.1ËÌ[10]。ÍWþÎ:lm8
¹þåQº»|}op¹¶q„<
。Cr„<¹¶

/Cr6+Ï]ÐÑ0、10、50、100μmol·L
-1
ß4n


¾n„<¹¶ÐÑ3næÒ。Cr6+/æq€
Ó
(K2Cr2O7)01Ôuò34`«,)*©ª。„
<10dPSÕ,Öù×ØÙÚ。
1.3 WX./YZ[
1.3.1 \]^ ¾„<±m47,Å_‰ŠÑ¥
10%EDTANa2Q²`«Ûa30min,mÑ,Á)
n~wÜzl~ÝÞß 

ņ‡ˆw‰Šà

105℃áj,70℃â««â™vãæ,Fæ。
1.3.2 犆狉_^ řäåsæ,ÂÁç€wè™~
(4∶1)žé,êÁëìílRS©î–ÖùCr‹X。
1.3.3 犆狉<=>?@A ïŽFmðñòä1g,
u10mL¼m`[0.25mol/Lóô+50mmol/L
5595—            123,ß:op`q[jk«Cr6+QŒŽ^_`[]<
TrisHClõŸ`(pH7.5)+lmmol/Löòô÷],
øù=åv¢ú

*PÁázΖ
[11]
ÅärK
Žª
(F1)、Žû(F2)wŽ±²ˆ(F3),/
ç€wè™~
(4∶1)žé,êÁëìílRS©î
–ÖùCr‹X。
1.3.4 `abc,(NPT)_^ m0.5g#$ò
ä

u5.0mLüðQ0.15%(W/V)˜ýþ€ÿ²
`

øù=åv¢ú

øù30min,*PzÎ(20000
×g,4℃)20min,m‡ß`Kä`。!ª Kumari
Bhoomika[12]I–Öù‘’“”,(NPT)‹X。
1.3.5 defg(GSH)hi]jkg(PCs)_^
 m1g#$òä,u5.0mLüðQ10% (W/V)
x"#€

øù=åv¢ú

*PzÎ
(15000×g,
4℃)15min,m‡ß`K$Öä`。!ª Hamid
Hassanpour[13]I–Öù。
PCs‹X=NPT‹X-GSH ‹X[14]
1.3.6 lmn_^ %&(MDA)‹XêÁƒ
()‚*\Z–
[15]
Öù

1.3.7 opqrs8 Fmjkòä0.5g,uÌ
5mL0.1mmol·L-1Q+€õŸ`(pH7.0,͋
0.1mmol·L-1EDTA、1% PVP),øù=åv¢
ú

*PzÎ
(15000×g,4℃)15minm,›-`,
!ª./0
[16]
I–ÖùSOD、POD、CATœ[。
1.3.8 犆狉6t8Vu ‰ŠLæ-çQ§¨
©
[17](Retentionrate,%)=(†…ˆæ-ç‹
X

†‡ˆæ-ç‹X
)/
†…ˆæ-ç‹X
×100%
³¼©
(Translocationratio,%)=(†‡ˆ
Cr‹X/‰ŠCr‹X)×100%
1.4 vwxEh0K
êÁSPSS13.0wExcel1ÃL˜2•|}
I3w¢£[ց

êÁ Sigmaplot10.0
A4

2 Ÿ ¬3
2.1 犆狉6+xEy9z{:\]^<|}
y¡1±û,Cr6+„<¢£½¸´2njkr
sº»†‡ˆw‰ŠQ™$mæ

¦ÓÔCr6+„<
¹¶Q³u

½¸ó¶5n5i¢
(犘<0.05)。¬
LªÄ‚
,10~100μmol·L
-1Cr6+„<…,†‡ˆ
w‰Š×„<£¯56©
,‘
tuv

±K32.6%
w35.3%,‘xyz’±K56.5%w52.5%,Cr6+
ÀÁL

xyz

;8Q56ÆKi¢

rs6ž
¢£
(犘<0.05)。p:,97‰‚(†‡ˆ/‰Š)
8

¬LªÄ‚
,‘
tuv

¿100μmol·L
-1Cr6+À
Á…¢£³u

¡iCr6+L‘tuv’‰ŠQ56ó
¶9W†‡ˆ

°

xyz

:¿50μmol·L
-1Cr6+
ÀÁ…¢£½¸

¡i

xyz

‰ŠLCr6+Q`[
e¥†‡ˆ

2.2 9:z{y犆狉6+<~%h€‚
y¡2±û,2njkrs†‡ˆw‰ŠCr‹
X¯ÓÔ Cr6+ „<¹¶Q³u°¢£³u(犘<
0.05)。10~100μmol·L
-1Cr6+„<…,†‡ˆw
‰Š«QCr‹Xׄ<£¯®,‘tuv’±K
115.5w2791.3mg·kg-1,‘xyz’±K145.2
w2261.0mg·kg-1,¿Äp¹¶QCr6+„<…,
ƒ1 犆狉6+xEy9:\]^<|}
Table1 EffectofCr6+treatmentsonthebiomassof犅狉犪狊狊犻犮犪犮犺犻狀犲狀狊犻狊
rs
Variety
q¹¶
Crconcentration
/(μmol·L-1)
†‡ˆShoot ‰ŠRoot
;$XBiomass
/(g·plant-1)
56©
Inhibition
percentage/%
;$XBiomass
/(g·plant-1)
56©
Inhibition
percentage/%
†‡ˆ

‰Š
Shoot/Root
tuv
Meijiahua
0 6.89±0.20b 0.61±0.01c 11.4±0.5b
10 5.89±0.17d 14.6 0.50±0.01d 18.1 11.8±0.4b
50 4.81±0.14e 30.2 0.43±0.02e 29.9 11.3±0.6b
100 3.23±0.21g 53.1 0.26±0.07f 57.9 13.7±1.2a
xyz
Sanyueman
0 9.95±0.09a 1.05±0.07a 9.5±0.7c
10 6.27±0.35c 37.0 0.68±0.02b 35.3 9.2±0.5c
50 4.06±0.15f 59.2 0.54±0.02d 48.8 7.6±0.5d
100 2.62±0.16h 73.7 0.28±0.02f 73.3 9.4±0.9c
;

p<«op=>¡?žp¢£~£
(犘<0.05);;$X/™æÏ。
Note:Differentlettersinthesamecolumnindicatesignificantdifferenceat0.05level;Thebiomassisdryweight.
659 ! " # $ % &                   36@
ƒ2 9:犆狉<_^、„…†、€^h‡ˆ†
Table2 Thecontent,retentionrate,accumulationandtranslocationratioofCrof犅.犮犺犻狀犲狀狊犻狊
rs
Variety
q¹¶
Crconcentration
/(μmol·L-1)
q‹X
Crconcentration/(mg·kg-1)
†‡ˆShoot ‰ŠRoot
§¨©
Retention
rate/%
qZYX
Accumulation/(μg·plant-1)
†‡ˆShoot ‰ŠRoot
³¼©
Translocation
ratio/%
tuv
Meijiahua
0 nd nd nd nd
10 10.0±0.3e 669.9±8.0d 98.5±0.1a 58.9±3.6c 333.2±6.0b 17.7±0.8e
50 26.1±0.7d 874.2±10.7c 97.0±0.1b 125.6±7.1b 371.9±21.3a 32.2±1.9d
100 79.4±2.9b 1247.2±87.6a 93.6±0.3d 257.3±18.2a 320.0±29.7b 80.5±2.3b
xyz
Sanyueman
0 nd nd nd nd
10 9.2±0.4e 463.5±17.8e 98.0±0.1a 57.6±6.1c 314.1±14.5b 18.3±1.1e
50 34.3±1.6c 693.4±5.7d 95.1±0.2c 139.1±12.0b 371.5±14.9a 38.5±2.2c
100 101.7±1.9a 1104.1±69.1b 90.8±0.8e 266.1±10.0a 309.6±27.2b 86.3±7.5a
;
:nd¡?AMBÖò。p<«op=>¡?žp¢£~£(犘<0.05);q‹X/™æÏ。
Note:ndindicatesnotdetectable.Differentlettersinthesamecolumnindicatesignificantdifferenceat0.05level;TheCrcontentcalcu
latesbydryweight.

tuv

‰Š«QCr‹X¯¢£·¥‘xyz’,°
†‡ˆ«QCr‹X¢£¸¥‘xyz’(10μmol·
L-1CD;犘<0.05);2njkrs‰ŠCr‹XE
E·¥†‡ˆ
(犘<0.01),¡i‰ŠLCrMƒÒQ
§¨AÁ

p:
,2njkrsCrQ§¨©ÓÔ
Cr6+„<¹¶Q³u°¢£½¸(犘<0.05)。10~
100μmol·L
-1Cr6+„<…,‘tuv’w‘xyz’
§¨©×„<£¯®±K96.4%w94.6%,¦¿
50w100μmol·L
-1Cr6+„<…,‘tuv’§¨©
¢£·¥

xyz
’(犘<0.05)。FD,2njkrs
CrQ³¼©w†‡ˆCrZYXGÓÔCr6+„<¹
¶Q³u°¢£³u
(犘<0.05)。10~100μmol·
L-1Cr6+„<…,CrQ³¼©×„<£¯®¿‘t
uv

w

xyz

«±K43.5%w46.8%,¦¿
50w100μmol·L
-1Cr6+„<…,‘tuv’Cr³
¼©¢£¤¥

xyz
’(犘<0.05),°HrsCrQ
ZYX¿Äp¹¶Cr6+ „<…žo¢£(犘>
0.05);Hrsjk‰ŠCrZYX„i¢·¥†‡
ˆ

/‡Ÿ IiCr(jkRSP,78§¨¿‰
Š

¦

tuv

ÏMÆÒQJKCrJ†‡ˆ³TQ
º]

9°ÆLQÄMjkoNVÈOCrN@。
2.3 犆狉‰9:=>?Š@;<@A‚
y¡3±û,jkŒŽQŽª(F1)、Ž
û
(F2)wŽ±²[(F3)P«Cr‹X(­q‹
X
)¯
ÓCr6+„<¹¶QQ·°¢£³u。¿50
w100μmol·L
-1Cr6+„<…,‘tuv’†‡ˆw
‰ŠŽªP
(F1)«QCr‹X±ýÄpCr6+
„<¹¶…

xyz

Q1.56ü、1.44üw1.48
ü
、1.44ü,°_Ž±²[P(F3)«QCr‹X
±ýÄpCr6+„<¹¶…‘xyz’Q0.37ü、
0.83üw0.36ü、0.82ü,rs6ž¢£(犘<
0.05),ŽûP(F2)«QCr‹X,‘tuv’†
‡ˆ±ýÄpCr6+„<¹¶…‘xyz’Q0.40
üw0.60ü,¢£¸¥‘xyz’,°_‰Š:ýÄ
pCr6+„<¹¶…‘xyz’Q1.68üw1.86ü,
¢£·¥

xyz
’。Cr¿jk׌ŽP«
Q‚R‡
,2nrs2¿¢£Qž(犘<0.05)。


†‡ˆ‚R¿

tuv

¡ÐKF1>F3>
F2,°¿‘xyz’:ýF3>F1>F2;¿2nrs‰
Š«‚RSTÄU
,¯
ýF1«÷·,F2«÷
¸

¿ÄpQCr6+„<¹¶…,Cr¿‘tuv’†
‡ˆw‰ŠŽªP
(F1)«(50μmol·L
-1
¹¶
CD

Qr‚R¢£·¥

xyz
’,
°¿±²[P

(F3)«Qr‚R¢£¸¥‘xyz’(犘<
0.05);Cr¿Žû«…LQ‚R,‘tuv’†‡ˆ
¢£¸¥

xyz
’,
°_‰Š
(100μmol·L
-1Cr6+
„<…

«¢£·¥

xyz
’(犘<0.05)。/‡Ÿ
 Ii

tuv

QŽªÏMÆÒQYZCrQº
]

ÈÉ´|̍ŽÍQCr,9°¡ÐÑÆÒQ`
Cr6+[。
2.4 犆狉6+xEy9:`abc,_^<|}
V41…?,ÓÔCr6+„<¹¶Q³u,2nj
krs†‡ˆ‘’“”,‹X¯WÀ³·P½¸Q


ο50μmol·L
-1Cr6+„<…ÑÐ÷e®,
ÓP…½

XY¢£·¥Lª
(犘<0.05);°Z‰
Š«‘’“”,‹X¿Cr6+„<‡P¯AM¢£
7595—            123,ß:op`q[jk«Cr6+QŒŽ^_`[]<
ƒ3 犆狉6+xE‹2Œ9:Ž=>?Š@;犆狉_^BC@‘
Table3 ConcentrationofCrinsubcelularfractionsandtheirpercentagesoftwovarietiesof犅.犮犺犻狀犲狀狊犻狊
underCr6+treatments
#7ˆM
Plantpart
rs
Variety
q¹¶
Crconcentration
/(μmol·L-1)
q‹X
Crconcentration/(mg·kg-1)
F1 F2 F3
­q‹X
TotalCr
concentration
/(mg·kg-1)
r©
Percentages/%
F1 F2 F3
†‡ˆ
Shoot
tuv
Meijiahua
10 1.29±0.26e 0.48±0.07d 0.54±0.04e 2.30±0.31e 55.6b 21.0b 23.4b
50 4.02±0.20c 0.62±0.03d 1.26±0.25d 5.90±0.43d 68.3a 10.5d 21.2b
100 7.34±0.54a 1.30±0.07c 2.17±0.28c 10.81±0.83b 67.9a 12.0d 20.1b
xyz
Sanyueman
10 0.83±0.10e 0.65±0.03d 1.04±0.06d 2.52±0.10e 32.8c 25.8a 41.4a
50 2.58±0.19d 1.54±0.09b 3.45±0.34b 7.58±0.14c 34.1c 20.4b 45.5a
100 4.97±0.59b 2.15±0.27a 5.98±0.27a 13.09±0.98a 37.9c 16.4c 45.7a
‰Š
Root
tuv
Meijiahua
10 32.86±5.85cd 5.46±0.54c 9.13±0.57d 47.45±5.89e 68.9b 11.6b 19.5c
50 56.65±3.87b 9.44±0.90b 11.57±1.30cd 77.66±2.97c 72.9ab 12.2b 14.9d
100 81.69±9.09a 20.35±3.65a 20.30±1.89b 104.21±7.19a 78.3a 19.7a 19.5c
xyz
Sanyueman
10 24.05±4.44d 3.47±0.96c 11.26±1.19cd 38.78±5.75e 61.8c 9.0b 29.2a
50 39.37±2.90c 5.63±0.93c 14.01±1.53c 59.01±2.06d 66.7bc 9.5b 23.8b
100 56.73±5.75b 10.94±1.51b 24.81±2.19a 92.48±5.05b 61.3c 11.9b 26.8ab
;

p<«pþ#7ˆMop=>¡?žp¢£~£
(犘<0.05)。q‹X/òæÏ。F1KŽªP;F2KŽûP;F3KŽ±
²[P

Note:Differentlettersofthesameplantsiteinthesamecolumnindicateasignificantdifferenceat0.05level.TheCrcontentcalculatesby
freshweight.Celswereseparatedintodifferentfractions:celwal (F1),organele(F2),solublefraction(F3)bygradientcentrifugation
techniqueat4℃.

(犘>0.05)。¬LªÄ‚,10~100μmol·L
-1
Cr6+„<…,†‡ˆ‘’“”,‹X³µ¿‘tu
v

«K51.7%~114.9%,£¯K85.9%,°¿‘x
yz

«³µ[\K44.6%~97.8%,£¯K66.
4%,‘tuv’³µ¢£·¥‘xyz’,rs6ž
¢£
(犘<0.05)。Äp¹¶Cr6+„<…,2rs‰
Š«‘’“”,‹Xžo¢£
(犘>0.05)。
  p:,2njkrs†‡ˆw‰Š«]^_—
(GSH)Q‹XÓCr6+„<¹¶Q³u°`a½¸,
¦„<6QžeÇpò¢£~£
(犘<0.05)。¬
LªÄ‚
,10~100μmol·L
-1Cr6+„<…,†‡ˆ
w‰Š«GSH‹X,‘tuv’½µ[\K16.9%
~46.7%w22.9%~63.8%,£¯K28.2%w
46.6%,‘xyz’½µ[\K34.4%~77.4%w
37.1%~78.8%,£¯K56.3%w59.4%,‘tu
v

½µ¢£¸¥

xyz
’,
¦rs6ž¢£
(犘
<0.05)。
FD

¬LªÄ‚
,10~100μmol·L
-1Cr6+„
<¢£³u´2njkrs†‡ˆw‰ŠÍ#$•
–—
(PCs)Q‹X,_³µ[\±K115.9%~
267.2%w52.1%~100%。†‡ˆQ³µ¢£e
¥‰Š
(犘<0.05)。Hrs†‡ˆ«PCsQ‹X¿
50μmol·L
-1 Cr6+ „<…pò÷e®。50 w
100μmol·L
-1Cr6+„<…,‘tuv’†‡ˆPCs
Q‹X±‚Lª³u242.2%w219.9%,‘xy
z

±³u267.1%w203.5%,‘tuv’Q‹X
¢£·¥ÄpCr6+„<¹¶…‘xyz’Q‹X(犘
<0.05);2nrs‰ŠÍPCsQ‹X¿Äp¹¶
Cr6+„<…¢£ž(犘>0.05)。
b–/‡3±cCr6+dä´jkðÍ PCs
w NPTQ–s,¦‘tuv’†‡ˆº–sÆNj
”,$m¬ Cr•–,{„eý_ÏMƒÒfNº
]Qí%,þ

2.5 犆狉6+xEy9: 犕犇犃_^hopqrs8
<|}
  gÀ,ÓÔCr6+„<¹¶Q³u,2njkrs
†‡ˆw‰Š MDA‹XGW‡QX¹(42)。H
rs†‡ˆQ MDA‹X¿¸¹¶(10μmol·L
-1)
Cr6+„<…¬Lª¯¢£ž,°¿«、·¹¶
(50、100μmol·L
-1)Cr6+„<…¯‚Lª¢£³
u
,‘
tuv

w

xyz

³µ[\±K12.1%~
24.1%w21.9%~43.3%,‘tuv’Q³µ¢£¸
859 ! " # $ % &                   36@
41 Cr6+„Fig.1 EffectsofCrtreatmentsonnonproteinthiols,GSHandPCscontentsinshoots(A,C,E)androots
(B,D,F)of犅.犮犺犻狀犲狀狊犻狊
¥p„<¹¶…

xyz

Q³µ
(犘<0.05)。2r
s‰Š« MDA‹X¿Cr6+„<…¯‚Lª¢£
³u
,‘
tuv

w

xyz

¿10~100μmol·L
-1
Cr6+„<…Q³µ[\±K6.3%~47.4%w
11.9%~64.8%。¿Äp¹¶Cr6+„<…,‘xy
z

#7Í

†‡ˆw‰Š
)MDA‹X¯¢£·¥

tuv

Q‹X
(犘<0.05)。


¬LªÄ‚

ÓÔCr6+„<¹¶Q³u,
2njkrs†‡ˆw‰ŠSODœ[WÀ³·P
½¸QX¹

ο 50μmol·L
-1(
†‡ˆ

w
10μmol·L
-1(
‰Š
)Cr6+„<…pò÷e®,&:

tuv

³µ±ý35.7%(†‡ˆ)w7.9%(‰
Š
),‘
xyz

³µ±ý21.2%(†‡ˆ)w4.6%

‰Š
)。
ÓP
,2njkrs†‡ˆw‰ŠSODœ
[¯…½

Xv100μmol·L
-1Cr6+„<:,†‡ˆ
QSOD œ[Y¢£·¥Lª(‘tuv’³µK
28.0%,‘xyz’³µK9.0%),°‰Š:¢£¸
¥Lª
(‘
tuv

½µK8.8%,‘xyz’½µK
67.8%)。¿10~100μmol·L
-1Cr6+ÀÁ„<…,

tuv

†‡ˆw‰Š

C10μmol·L
-1Cr6+D)
ÍSODœ[¯¢£·¥Äp¹¶…‘xyz’Qœ
[
(犘<0.05)。
ÂÇ

¬LªÄ‚
,2njkrs†‡ˆw‰Š
«PODœ[ÓÔCr6+„<¹¶Q³u¯WÀ³·
P½¸QX¹

¦

xyz

w

tuv

†‡ˆ›œ
[±¿10w50μmol·L
-1Cr6+„<…pò÷e
®

°Z‰Š«›œ[¯¿10μmol·L
-1Cr6+:
pò÷e®
(‘
xyz

³µK42.1%,‘tuv’³
µK56.4%);ÓP…½,Xv100μmol·L
-1Cr6+
„<:
,2nrs†‡ˆw‰Š«›œ[Y¢£·
¥Lª
(‘
tuv

†‡ˆw‰Š³µ±ý33.3%
w33.9%,‘xyz’ÄZ³µ:±ý13.2%w
9595—            123,ß:op`q[jk«Cr6+QŒŽ^_`[]<
12.4%)。¿10~100μmol·L
-1Cr6+„<…,‘t
uv

#7Í

†‡ˆw‰Š
)PODœ[¯¢£·¥
Äp¹¶…

xyz

Qœ[
(犘<0.05)。
FD

¬LªÄ‚
,2njkrs†‡ˆw‰Š
«CATœ[¯ÓÔCr6+„<¹¶Q³u„WÀ³
·P½¸QX¹

ο50μmol·L
-1(
†‡ˆ

w
10μmol·L
-1(
‰Š
)Cr6+„<…ÑÐ÷e®(‘t
uv

†‡ˆw‰Š³µ±ý27.4%w138%,

xyz

ÄZ³µ±ý17.7%w87.4%),ÓP
…½

v100μmol·L
-1Cr6+:,‘xyz’†‡ˆw
‰Š«›œ[h¢£¸¥Lª

°

tuv

†‡ˆ«
CATœ[Y¢£·¥Lª。¿10~100μmol·L
-1
Cr6+„<…,‘tuv’†‡ˆw‰ŠÍCAT œ[¯
¢£·¥Äp¹¶…

xyz

Qœ[
(犘<0.05)。
/‡Ÿ Ii

tuv

#7¿Cr6+ÀÁÂÃ
…ÏMÆÒQßC ROSQº],/½¸CrL#$
isQ™šÀÁ

{„ý_` Cr6+ ƒÒQ]<


3 Ÿ“¬“
Crý#$;8Q‘jkOP。|ÌðÍQ
Cr6+L…M;$GMN@[1]。Cr6+ º»56#$
‰ŠŽQl

Jm~RS

9°56‰Q;
8

|°äå#7Wnoµ

#$Q©–AÁ

pR
42 Cr6+„Fig.2 EffectsofCr6+treatmentsontheMDAcontentandtheactivitiesofSOD,PODandCATinshoots
(A,C,E,D)androots(B,D,F,H)of犅.犮犺犻狀犲狀狊犻狊
069 ! " # $ % &                   36@
AÁß×s;<(qâ;rs

÷täå;$X½
¸
[4]。
‰=>¡i

¬LªÄ‚
,Cr6+„<¢£½¸
´jk†‡ˆw‰ŠQ;$X

¡iCr6+Ljk#
7u;N@AÁ

¿Äp¹¶ Cr6+ „<…,‘tu
v

†‡ˆw‰Š;8QN5󶯢£¤¥

xy
z
’,
¡i

tuv

L Cr6+ ÀÁÏMƒÒQ˜[
]6

#$L×sæ-çQ˜[]6ovÄp


ð‡8

þw?WHÂ,‰xy˜Ð

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êz]6

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Ž

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Fþxy
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VŸ–òŽª‡

7}TU
|Ì`a

V¬~|M]€w’“mŸ–ß
[4]。
‰
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,Cr(jkRSP,78§¨¿‰Š,J†
‡ˆTUƒÉ

祀QDˆêz]6

¬

xy
z

Ă
,`
[rs

tuv

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·

¡i_ÏMÆÒQJKCrJ†‡ˆ³TQº
]

9°ƒL†ÄMjkoNVÈOCrN@。
Cêz]6D

#$ŽÍQ`N]6¿#$
˜æ-çÀÁ«„æ8AÁ

_«ÅMNzl‚
ƒšýŽÍæ8QfNxy,þ

°Žªw`
aý‚ƒšQHn78Mt
[11]。
‰˜«

¿Cr
ŒŽ‡
,10~100μmol·L
-1Cr6+„‰Šw†‡ˆ
(‘
xyz

CD

ŽªP…LQ‚
R£¯¿60%/‡(¡3),¡iŽª¿jk`
Cr6+N«æ8AÁ。{¬„……wZhuß[1819]
=>Ÿ þå

p:
,`
[rs

tuv
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“‰Š
w†‡ˆ

¿ÄpQCr6+„<¹¶…,_Žª‡
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Cr6+N@Qí%,þ。*°,¿‰˜jk­ Cr
‹X«

‡M60%(‘xyz’†‡ˆ)w30%(‘tu
v

†‡ˆw‰Š^

xyz

‰Š

QCr?WŽ
ª|Ìí;mÍ

¡3)。_«,‡20%(‘tuv’†
‡ˆw‰Š^

xyz

‰Š

w40%Q(‘xyz’
†‡ˆ
)Cr¿±²ö«。±²ö78ýÙ`
a
[20]。
#$?WōŽm«QCr³TÎwx¿`


½¸Cr¿ŽÍQzlœ¶,¬|CrL_
Žû;<ˆºQ™‰
[21]。
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`a«

ŽÍQM]rð⊴æ8AÁ

{|
M]rð~,€

M]€wQ‹j^€QÇ
—

Z±¬-çzl•–

*PÅ_³Tv`aÍ
w2

‘’“”,
(NPT)QMj^€‹,
(SH),_SH ±¬-çzl•–。#$•–—
(phytochelatins,PCs)ç¥ NPT,_¬æ-ç}–
(Œo¾Ký#$ŽÍfCæ-çN@Qæ8]
6
[22]。
M=>¡i
,Cr6+³u´犘.狊狋狉犪狋犻狅狋犲狊‰«
j^€
(PCsQPss)Q‹X[23]。¿‰=>
«
,10~100μmol·L
-1Cr6+¹¶…,jk†‡ˆ
NPT‹XwPCs‹X¯¢£·¥LªP,¦‘tu
v

³uQµ¶Æe

¡i Cr6+ dä´jkðÍ
PCsw NPTQ–s,¦‘tuv’†‡ˆº–sÆ
Nj”,$m¬ Cr•–,{„eý_ÏMƒÒf
Nº]QFþí%
。GSH ý PCs–sQÞ$,y
PC–s›šu;[24]。Nòæ-çÀÁP,GSH
‹Xy¥PCsQ–s°…½¿Çs#$«GM&
d
[25]。
‰˜«

Ž*Z‘ò2njkrs‰
ŠwWn«GSHQ‹Xy¥PCsQ–s°`a…
½

X³uQPCsQ‹XE·¥ÈÉQ GSH,Ii
Cr6+º’“”Q GSH Q–s。¿Cr6+ÀÁ„<
…

¬LªÄ‚
,2njkrs GSH Q‹X¯W…
½QX¹

X

xyz

…½Qµ¶ÆKi¢

{±/
ˆf•

xyz

w

tuv

Q` Cr6+[ž。
GSH C´±/•–#$ðÍQæ-çD,„ýþ
sÿæ8Q¸l˜™š$

¬SOD、PODwCAT
ߘ™š›þ–s#$Q˜™šŠ—

˜™œ[
™Qš@

iōŽQ›ú;fœ}
[26]。
o|̍ŽÍQ-çzl9W´ŽÍM]r
ðL_Q•–º]:

œzQ-çzlžŽ
ÍQ™šÀÁ
。Cr±/däu;WXQœ[™)
y,
(ROS),Z¬Ÿ€、 Ë、’“m、,€ßâ
;\Z

žo±¡Qš@^Ž;<
[27]。MDA
ýË«oqw ¢€fQu$,þ

(¾Ký£#$¿¡.-.…ŽËW™šó¶Qæ8Ù
Ú
。SOD、CAT、PODý#$ðÍßCœ[™æ8
QŽÄM›B

_œ[·¸±/\£#$L¡.
ÀÁQYZº]
[28]。
‰˜«

¬LªÄ‚
,¸
¹
¶Cr6+ (10μmol·L
-1)
„<…

jk†‡ˆ
SOD、POD、CATœ[¢£³·,¦ MDA‹XAM
¢£#š

Ii¿&¹¶…

jk†‡ˆu;Q
ROSº(˜™šŠ—ßC;ÓÔCr„<¹¶Q|
þ¤¼·

†‡ˆ˜™š›

C

xyz
’CATD)œ
[Y¢£·¥Lª

*° MDA‹X¥¢£³u,I
i#$LCrÀÁQ¦™ýMþùS¶Q,9W&S
¶#$§Nòš@

{¬Kale[29]wAdreesß[30]
1695—            123,ß:op`q[jk«Cr6+QŒŽ^_`[]<
Q=>Ÿ ÄU

FD

¿¸¹¶ Cr6+(10μmol·
L-1)„<…,jk‰Š MDA ‹X¢£Q·,Ii
‰ŠNò‚†‡ˆÆeQ™šÀÁ

¿Äp¹¶Q
Cr„<(CKCD)…,` [rs‘tuv’#7ÍQ
MDA‹X¸¥‘xyz’,°SOD、POD、CATœ[
¢£·¥

xyz
’,
†?´tuvÏMÆÒQßC
ROSQº],/½¸CrL#$isQ™šÀÁ,{
„ý_`Cr6+ƒÒQ]<,þ。
b‡…¨

¿Cr6+ÀÁ…,jk;8Nò¢£
56

jkgÀº?W‰Š_ùCr,JKCrJ†‡
ˆQTU

/½¸#7†‡ˆCr‹X;_Ǻ?W
ŽªŸ–Cr,/ÈÉ|Ìí;mðQCr;ÂDŽ
e?WPCs•–Crzl,ÎÅ_³¼ò`aÍ,ë
_;$œ[bš

FD

©?W³Ò#7ð͘™š
›œ[

/½¸œ[™ßLmËQš@

9°ÆL†
iōŽmËQøù[wª«[

¬

xyz

Ä
‚

jkrs

tuv

ÏMÆÒQ_ù

•–Crw
˜™šQº]

{±ºý_˜ Cr6+ [ƒÒQæ8
í%

’“!M

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