免费文献传递   相关文献

Changes in Antioxidative Enzymes Parameters of Populus ussuriensis Seedlings after Spaceflight of the Seeds

大青杨航天诱变植株早期抗氧化酶生化指标测定


Dry seeds of Populus ussuriensis collected from Heilongjiang area were carried by the recoverable satellite for mutagenesis. Then the growth traits and antioxidant enzymes activities of seedlings from the spaceflight-treated seeds and controls were analyzed. The results showed that the growth traits of the seedlings after spaceflight varied to some degree, but most of the variation was not remarkable. There were no significant changes in soluble protein content,malondialdehyde (MDA) content and superoxide dismutase (SOD) activity in seedlings with response to spaceflight treatment. However,activities of ascorbate peroxidase (APX) and guaiacol peroxidase (POD) in the poplar seedlings after spaceflight were significantly higher than those from the ground control. It suggested that space condition altered the poplar traits in some degree,and the higher antioxidant enzymes might failitate themselves to avoid some damage from active oxygen.

 


全 文 :! !"" ! ##
$ % % &$ # %
& ’ ( )
’()*+,)- ’)./-* ’)+)(-*
/012!"*+02#
341 5*
!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!
$ % % &
;|/LM’_v,NI#O;P@;h.CD"
}~!6’$ K "7 ;l#6’$ N$%6 }A&6 F’_6
662 )>&’F)&eyzHvU/0[9$HsoYKLU ?@A 6"%%!%@
$2 )>&’F)/6())D ?@A 6"%%!%@72 ?@Az’F)0HKezg)D ?@A 6"%%%6X
$%&t F_%@2qY)@/M33@{N*g
’()*+#’#$"26 ,-./0#- ,12+#6%%6 8 #!993$%%&4%# 8 %6!" 8 %"
PQR#t$%%9 8 %: 8 $%o
/STUt4wACVWXYZ([GSTU6;-%:<7%6 8 $ 8 7X*)>&’F)_$(V/Sx4wAC‘>\/S6.=< 8 $&&Xo
"m8huYklmo
!"#$%&’ ($ )$*(+,(-#*(.& /$012&’ 3#4#2&*&4’ +5 !"#$%$& $&&$’()*&(&
6&&-7($%’ #5*&4 68#9&57(%"* +5 *"& 6&&-’
’0>? @A>?BC4>6*$ D4 EA7 .A4 @4FGFA6*$ .A HIA10>?6 ’0>? J4>I>6 KI>? (C4I>LA>?6
662 !"# $%&’(%)’(# ’* +’(",) -("" ."/")01 234(’5"3"/) %/6 70’)"18/’9’:# ’* ;0/0,)(# ’* <6=1%)0’/*>’()8"%,) +’(",)(# ?/05"(,0)# @%(&0/ 6"%%!%@
$2 $0*" A10"/1" B’99":"*>’()8"%,) +’(",)(# ?/05"(,0)# @%(&0/ 6"%%!%@
72 A18’’9 ’* ;=/0104%9 %/6 ):’*4#9*t EMN BFFOB 0P C’4=9=, =,,=(0"/,0, Q011FQRFO PM0G SFA10>?TAI>? IMFI UFMF QIMMAFO VN RCF MFQ0WFMIV1F BIRF11ARF P0M
G4RI?F>FBAB 5 ,CF> RCF ?M0URC RMIARB I>O I>RA0XAOI>R F>YNGFB IQRAWARAFB 0P BFFO1A>?B PM0G RCF BLIQFP1A?CRZRMFIRFO BFFOB I>O Q0>RM01B
UFMF I>I1NYFO5 ,CF MFB41RB BC0UFO RCIR RCF ?M0URC RMIARB 0P RCF BFFO1A>?B IPRFM BLIQFP1A?CR WIMAFO R0 B0GF OF?MFF*V4R G0BR 0P RCF
WIMAIRA0> UIB >0R MFGIM[IV1F 5 ,CFMF UFMF >0 BA?>APAQI>R QCI>?FB A> B014V1F LM0RFA> Q0>RF>R*GI10>OAI1OFCNOF6\E-XQ0>RF>R I>O
B4LFM0XAOF OABG4RIBF6’]EXIQRAWARN A> BFFO1A>?B UARC MFBL0>BF R0 BLIQFP1A?CR RMFIRGF>R 5 S0UFWFM*IQRAWARAFB 0P IBQ0MVIRF
LFM0XAOIBF6-^@XI>O ?4IAIQ01 LFM0XAOIBF6^]EXA> RCF L0L1IM BFFO1A>?B IPRFM BLIQFP1A?CR UFMF BA?>APAQI>R1N CA?CFM RCI> RC0BF
PM0G RCF ?M04>O Q0>RM01 5 )R B4??FBRFO RCIR BLIQF Q0>OARA0> I1RFMFO RCF L0L1IM RMIARB A> B0GF OF?MFF*I>O RCF CA?CFM I>RA0XAOI>R
F>YNGFB GA?CR PIA1ARIRF RCFGBF1WFB R0 IW0AO B0GF OIGI?F PM0G IQRAWF 0XN?F>5
;&1 <+4-’t C’4=9=, =,,=(0"/,0,@BLIQF G4RIRA0>@?M0URC RMIAR@I>RA0XAOI>R F>YNGF
u$-*7?SVW>stk5#_‘IJ2bcGW%#_vTi#j5lD*b/0/MG%HG/
MGyz2~!FlqQ%@u*’#VWou0
vsfZ_‘2n\.h%S*46/F1#/6
3)9Nyz3)96=EW*6&&:@;]:F*
6&&:XoG 6&9# $[-*Z7ur1hgD.To
ac6\>PPH*6-(0)01=3 %",)05=3XGw+6D(#E%
,%)05%XG&36F"% 3%#,XGn26.’,,#40=3 BLL5XGF
i6.9#10/" 3%GXG,-6 A’9%/=3 9#1’4"(,01=3XFop
El0vs*}LHPXZ1t}O6=EW*
6&&:X*u_‘[9$E’1#gDLlP6#c
H*4U]$&elqV%)*oA$_Y)Hv
[9_&8&’VW<*bLH&e6}v|Tu
Xv>bk:#6’(o
0VWA$_2nz6#F_%6 C’4=9=,
=,,=(0"/,0,Xvs.hHA\*[3GHAu~3*b
@A#/M2{N*g/*%zmnOU*[#u
2q[9gD{&e6}vdHz{$mo
6 DE{lM
626 [wCDVIG6’( F_%>PCvh
Yu\T&’yFB>?&Y*{ $%%7 $ 66 % !
R 6"t%%>P! 69 @.Toac*2n 69 q\.
Thqo{ $6R 6%t%!$dW2n<4tuhY $%% [G*HhY 7"% [Go$%%!$
"%A2qY)vsxbMvsHvmnHUH
A*}~L*x 7$/F_%2qY)AxbMA#evC
HI*37IHxlu 6IN3OmnGHo0VWG+L
_GH\/M*##M7;B#F_%2qY)A
"IN3O2bMA " IN3O*b3$/M*#
mn/016GH\ 6I%x :I%X2{N*%z
6GH\ !I%X#UKo
!"# ]JuKeu&WNOIJ +N*0]*
g6$%&Xw3D@cdAaUK6’()**+,+-(.(/ !"
#$ 0*!122X@{,DhZN*0g6345Xw3#U
K*CD# 36)7+F6!118X#lamn@VNej
ZN*0g64%&Xw3x 9&3 m1#UK$M
:)76)7F6!11#X#la@473u0#z_2UK
CD ;<)*=>+<=a6nQHF*#???Xo
9 # F_%2qY)A2bMA
@I%v/M33?@
A(B0 # CDE F+6,)<(/+* +* .DE B<+G.D .<)(./ HE.GEE* /,)FE>-(BD.
.9# !J!?oCDE /EE=-(*B DE(BD. G)/ +*E .E*.D +>
)F.K)- L)-KE
9 ! F_%2qY)A2bMA
!I%v/M33?@
A(B0 ! CDE F+6,)<(/+* +* .DE B<+G.D .<)(./ HE.GEE* /,)FE>-(BD.
.# =){)2
#"! iE+L{2T16rM1Nef^O=P
b|H/Mwx6GH\ !I%2 @I%X#F_%
2qY)A2hqbM#cF MT/M33mn?@69 ! N MXopq%:G
H\ !I%v*F_%2qY)A MT/M33;
><{hqbM*4nUhqbM89’s8&7
w:6+ >z{u ?"?!*#"M#M*?"!8O*% P ?"?QXo_
!*F_%2qY)A2hqbMA @I%v*hq
bMA# MT/M33;4:<{Hv#2qY)
A#67*4l8zl4589’s&7w:6+ >
z{u #"2@2*#"MQ@*#"!?M*% P ?"?QXo=42rs
f2qY)#F_%@A$/M*#XKg41/
M,4@P*4nGH/M*17DE&)o=Ui
.5F6#??#X%:#B8C 82 S2qY)\c<
9:Phq6*Y)Naw##+
%z8*=42U:L~3|H%So
9 M F_%2qY)A2bMA !I%
2 @I%/M33j5?@
A(B0 M CDE BE*E<)- F+6,)<(/+* +* .DE B<+G.D .<)(./ HE.GEE* /,)FE>-(BD.
.(* +*E 6+*.D )*= /(I 6+*.D/
9# !J!?oCDE /EE=-(*B DE(BD. G)/ +*E .E*.D +>
)F.K)- L)-KE
#"# L{2TrQ2]JuKvwPQ !X{,
DhZN*0g6345Xw3zl {,DhZN*
0g6)/F+gAKDbO*@X’(#opqzbX6mowF*#??@Xo0
VW<*# OMSS*f2q>P#F_%Ae[q
< 345w3;<{hqbM69 8X@3<*2qA
[q 345w3u ?"Q2 N M"8O!6+-v6(*
R ! BR ! AT*!
hqbM#F_%[q< 345w3u ?"@# N !"?8
!6+-v6(*
R ! BR !ATol8zl45tf2q>P#F
_%Ae[q< 345w3UhqbM#w389
s8&7w:6% U ?"?QX*_‘Ke#JKv*345b_‘?eJKw3N#
Fr6/Wg*Cbw315*m!b-$}8I#
lDo
#XZN*0g64%&Xw3zl 4%& nu0
bKe!1O!Xo0VW<*f2q>PF_%Ae[q<
4%&w3"bB{hqbM69 QX@3<*2qA<
@8! & ’ ( ) 8Q"
[q !"#w3u $%&$ ’ $%(&!)*+v),-
. & /. & 01*!
hqbM#F_%[q< !"#w3u $%&2 ’ $%32
!)*+v),-
. & /. &01ol8zl45tf2q>PF_
%Ae[q< !"#w3UhqbMw389s8
&7w:6! 4 $%$3X*#JK? !"#gw32,?@*:;?bM@BP
&&2%(35*=42nF_%Ae$XKg41O8
S$_?eJK#fgo
9 6 F_%2qY)A2bMA &I%v 7!8w3?@
0,/9 6 :;< =*)>?@,A*- *- B;< 7!8 ?=B,C,BD E?=B@9 3 F_%2qY)A2bMA
&I%v !"#w3?@
0,/9 3 :;< =*)>?@,A*- *- B;< !"# ?=B,C,BD E?=B@(X473u0,m1zl u0,n-$p
0<;op#rz*u0b?473u0,m1U
{n3%XK#SOo0:LVW%:69 IX*f
2q>PF_%AeUhqbM[q<473u0
,#m1L?)*|X*z{L{ $%J3 ’ $%KL x
$%JI ’ $%2I!)*+v/
. &01b‘*425{|HAe#
Ez4|H*TXO_‘KeVW\F_%473
u0,#m1)*|Xofl8zl45tJ m‘
473u0,#m189’s&73w:6 ! M
$%$3Xo42q>PF_%#473u0,#m1
;>^><{hqbMo=42nTuO8_‘K
eVW#F_%b?473u0m1XKg4#R
S*%g{z3#/6kYo
9 I F_%2qY)A2bMA &I%v
473u0,m1?@
0,/9 I :;< =*)>?@,A*- *- B;< A*+NE+< >@*B<,- =*-B<-B EA>?=A<6X+N*0]*g6O"#Xw3zl +N*0
]*g2KDw3NN3IZN*0G5/b-$
MOP#Ko*n%+bKD+l< &Isop#
I#rz*TX*3w3Uu0{3#/LSo0V
W<*2q>PF_% &I%/Ae[q< O"#w
3*# L(SS,b><{hqbM69 2Xo3<*
2q>P#F_% & I%/Ae[q< O"#w3
L{ $%JL ’ I%LL Pv/. &01b‘*hqbML{ (%I&
’ I%L& Pv/. & 01b‘ol8zl45t2q>P
#F_% & I%/Ae[q< O"#w3Uhqb
M89’s&73w:6! M $%$3X*P
F_%[q O"# w3UhqbM89sHo%V
Wm$b2q>P0!>Y J $/Ae O"#w3
#VWP0!# O"#w3Uhq
bM#89)H*U! &$L?4:1567*~4
:9<{hqbM#676Ga*J$$IXo
9 2 F_%2qY)A2bMA
&I%v O"#w3?@
0,/9 2 :;< =*)>?@,A*- *- B;< O"# ?=B,C,BD E?=B@26&! 2# REEFtF_%2qY)uc8#{N*g/*%zUK
!X% & ’6"#$Xm 1 z l % & ’
6"%&’()*%&)+,-)+*"#$XnG5/b-$MiZN*
Ko#;B60bX*3m1L{ ./01 2 ./34
!5’&v6
7 89:b‘*!hqbMAe[q#"#$m1
L{ ./08 2 ./01!5’&v6
7 8 9:b‘*#2qA 30S
S*3|2q>P\F_%>Y 0$/A[q "#$
m1;<{hqbM*42n5{F_%Aeb_
‘Ke#VW@uEz*O8_‘KeXKg4#
JKlD*Cb2q>P\#F_%6QOO8|
Hg4#JKol8zl45t2q>P>YF_
% 8I%/Ae[q< "#$m1UhqbM89
’s&73w:6! ; ./.!X*%Ae-$MVW}|!oo
9 < F_%2qY)A2bMA
8I%v "#$w3?@
9*6= < >,+ ?’5@%A*B’( ’( C,+ "#$ ?’(C+(C D+CE++( B@%?+F&*6,C
CA+%C5+(C %() ?’(CA’& *( ! = "##"$%&’#%# B++)&*(6B *( ’(+ 5’(C,
0/3 \ZPQ YZb2q>PF_%Ae[2
hqbMAe#/012{N*g/*%z^o#
LS3zl*5r 845*0VW<4U^oszb
‘s8&7NmI&7#LSSOo3<*8 %"
uc#cM2;F[d‘;s8&7NmI&7#=LSS
O*! 8%"#c%z‘4&7NI&7#$LSSO*3XT42
n$GH 8I%v*ucO_‘Ke#VW@F*U
hqbML?/M@u2,@! 4I%v=v17
|]s&*3 T/01%z>B{HXv‘#hq
bM*TX*42v~ 8%"F_%@AU 4%"@
A4$LSSOo${N*%z<*GH#U 8%"
F_% 3T/01%z4&7#=LSSO*=%
g{KDw3Nb-$MOP#Koo
3 "#
2qHvUzP#bcY)Hv?@5%)9
WO$$_Ke?C/00K#yz0,Pb222p_
b6/\TGojG~=GtMF!_}XOVyz
xY/’(1#/T")*=v")S,P/0t
$G_LLyo?^p%{$1G$/j,Mv‘
P26/#yz)9#v‘mgo5{3-$")
WOp<{b-$#")WO*TX*Y%")vf
+CD3-$lu~3*[z_VWvu$_Y)4Cuc/HmgFop33
%/)9*~bvs2uc#JKH*uLH6}v
z{P423o
0VWP#F_%vsdH9#
J@8YM9# 8%"x 4%"/@Ac2;F[dF 3T/M33UhqbMb‘89’
s&7w:*=UFE5F60..4Xb$_Y)\F
cL9$>Y6JG8XTU833)9#VWuUbML?f%)*p7LX~of2q>P#
F_% 8%"/@A>B<{Hv##hqbM@
!/M8 4I%#vA*hqbM#@A 3T/M
33><{2q>P#@A*=42y50VW<
:L~3OhqKeVW\_‘b3\r#17>
%?@o$_Y)#p4[v~)9#bLH"+3&e6}v|TuXv6’(o
_‘Ke<#BooGz2cZGz2NYFI
Jb{hI1=+/M#u0-J*=v?e42_}u0I#gw3#E)o0
VW<*F_%[q< $GK*JH#*GH# F 3 v{N
*gf_‘Yv\*UhqbML?w3;%|H
g4#E)*# JH#S3|gw389;s8&7
w:*=Ur~GF6811-@Ab?{N*gw3%|Hg4z<#szp
7LX~o=42y5_‘#juKebF_%rXK#VW*!{N*gw3#z<%g{KD
w3Nbuc-$#Koo{N*gw30F1m
UP[q/M8 8I%v#>4I%v*5{F_%#_‘[qs,**AN>
<*[j{g#w3^o\]4D3*$"\#VW
z#)*Lbg*m!2O@up,hosgw3
U/M%zb2qY)#Wgo
U]2_Y)#Bq^_q$O7oG<*i
.5F60..0XbB8C LI S_‘Y)#33)9
VWvu_‘Y)b3{n3FEG33r:uX
K#=17VWo!gBiF60..IXb 0IH*}
v6OX_‘Y)\vs#EG33VW%: 0I}
5 ! LM’_X7yy7L;|/@Q<9X#O;@:h.Ld$<)2!
"#$%! "&’ ()**’+#,’- #.#+/010 )2 ,&’ 3*)4,& ,*#1,0 #.- ,&’ 0’5’*#+ #.,1)61-#., ’.7/8’0 #(,151,1’0
$’,4’’. 09#(’2+13&, ,*’#,8’., #.- ().,*)+ 1. ! % "##"$%&’#%# 0’’-+1.30
!%"uc
"##$%&’() *+,#- .’# /.’,0
1%"uc
"##$%&’() *+,#- )&2 /.’,0)
!%"uc
"##$%&’() *+,#- .’# /.’,0
c<
3#&(0, .+
4%*’,)
;F[M
5*2&/*%
%#*+
%#’(,0
;F[d
5*2&/*%
%#*+
6-#*$,0
c<
3#&(0, .+
4%*’,)
;F[M
5*2&/*%
%#*+
%#’(,0
;F[d
5*2&/*%
%#*+
6-#*$,0
789 8:;
473u0
".%<6%#
4-.,#&’
":;
!%"uc
"##$%&’() *+,#-
.’# /.’,0
;F[M
5*2&/*%
%#*+ %#’(,0
=>???!!
;F[d
5*2&/*% %#*+
6-#*$,0
=>1!@ =>AB1!!
1%"uc
"##$%&’() *+,#-
)&2 /.’,0)
c<
3#&(0, .+
4%*’,)
C =>?=@! C =>1BB C =>DEB
;F[M
5*2&/*% %#*+
%#’(,0
C =>?1E!! C =>?!=! C =>?=A! =>?BF!
;F[d
5*2&/*% %#*+
6-#*$,0
C =>?D!! C =>A!D!! C =>EE! =>??E!! =>AD1!!
!%"uc
"##$%&’() *+,#-
.’# /.’,0
789 C =>=@B C =>=A! =>!?B =>B1A C =>!== C =>=!=
8:; =>?=A! =>1DF! =>1AA! =>B@@ =>EB1 =>D1E =>FBF
473u0
".%<6%# 4-.,#&’
=>=BB C =>=!D C =>!DD C =>1ED! C =>B!A C =>FAB C =>!?1 =>BAB
":; =>DAF =>!BF C =>!@F =>=A! =>!@B C =>=E= =>BDD =>E=A =>DD@
5;7 =>B!1 =>BBF =>!@A C =>FEE =>=E! C =>!FF =>BA! C =>DDB =>B1? =>==1
!!!!G =>=!@!!G =>=E H
v6OX#)9l7q$89@3<*!I)9cO#
33%z_^r:uRS*!3 !I#33%z_
^r:u&eo/jt%)mvu_‘#Boof
%VW8ucYv")WO6I*J*0*)0& !" #$ H*B==!@
:0’&)0& !" #$ H*B==!Xo5{_‘KeIJIuju*
Y)Tis_*ILTi}NvlDe[pK*++
v~2qY)pq5%|pK3o!%/)9#3
3nt5%vB3Gnt2O[Kyz*@^pp[
zs/0)[9mnmX5#LLUxKo
s t , -
gBi*rPl*D<)*F H B==? H H*2qY)HvBqVW H E
’zg()*BF6?XtE@A C 1=! H
;]:*9C.*Du#*F H !@@1 H _‘Y)$j(Hv<#gD H
_‘()))*!16R?Xt!BE C !F!H
S&{*cQ.*S$F H !@@? H ,Yvu0#G5/ZN*JK H
u0))*F@61XtEBB C EB1 H
=EW H !@@1 H A1F C B_‘Y)HvmM2]N H _‘()))*!1
6R?Xt?? C ABH
G a H B==1 H 2q>Pb0!vs2Ae33#VW H ?@At)
>&’F)@>)=7F H
i.5*iS7*%6}*F H B==B H B8C D?S,0_‘Y)#3
3)9VW H 2q>)U>)zg*!E6BXt!B? C !F!H
mnF H B==? H yz) H >?t()9^_ H
mow*3d&*9TS H B==1 H CHhXb":;x K7Iw3#VW H :YE’([*61Xt!1 C !?H
hy/ H !@A! H u0HzgVW#TLmM H u0/6)Y)*Ft
! C ?H
FE5*DWA*iI-*F H B==1 H FcL9$$_Y)\YEG3
3DE2},zl H BEE’))*!@6FXtF?F C F??H
r~G*o=. H !@@A H _‘2nb,-vswo23w3NY/#
VW H FG))*BE6BXt!1E C !1@H
nQH*- s H B=== H u0,[9$< H >?t()9^_ H
7/*J. L*7)*$* LH !@@D H "#4*-*,# *))*M) )4#N&+&N +.- *)N.-6*,# 4#-.2&$*)#
*’$ (<*&*N.% 4#-.2&$*)# *’$ +.- N0%.-.4%*),&N *’$ NM,.).%&N &).#’OM/#) .+
*)N.-6*,# 4#-.2&$*)#) &’ 4%*’,)H 8%*’, K#%% 80M)&.%*FE6FXtD@? C E=DH
K*J/*J P*5*-)N0’#- 3H !@@B H 5*(’#)&#’0*’N# *N,&Q&,&#) .+ )<4#-.2&$# $&)/<,*)# *)N.-6*,# 4#-.2&$*)#**’$
(%<,*,0&.’# -#$!BBB C !BB?H
R&*’’.4.%&,&) K S*T&#) " LH !@?? H "<4#-.2&$# $&)/<,*)#)U.NN<--#’N# &’
0&(0#- 4%*’,)H 8%*’, 80M)&.%*E@6BXtF=@ C F!DH
:0’&)0& I*I*J*0*)0& 7*:0’&)0& L*!" #$ H B==! H 7%JM%*,&’( *(#’,65SVXU
&’$E1F C E1AH
I*J*0*)0& 7*:0’&)0& L*I*J*0*)0& "*!" #$ H B==! H I0# #++#N,) .+
/&N-.(-*Q&,M .’ &’$’!)!/*&*#! H 7$Q "4*N# T#)*BA6DXtEEE C E1! H
!"#$% &’()
@D!! ?# REEFtF_%2qY)uc8#{N*g/*%zUK