免费文献传递   相关文献

Effects of Seawater Stress on Photosynthetic and Antioxidant Characteristics in Portulaca oleracea

海水胁迫下马齿苋光合及抗氧化生理特性研究



全 文 :!!"
"
!
!
#
#
!
$%&!"
!
(%#
!
!
"
!
#
!
$
)*+) ),-./+0) /0(0*)
!!!
123
$
!
4
%
!
56&
!!
!123
!"#
!
21224""
"
7
899:2114;1#"3!1231#1"!
v7¯°~wxyV¤ŠÖHÃhÓR‚WX
ZHº
#
¦
!
I
#
$
!
B
#
[0J
$
^:®÷)*<ö())*
#
Ç© ^:
!22242
&
CD
!
fÖÒHIúG‡0klH&þ`„2Xmn
#
tºÂO¨5Žp2q/Ïûü
$
-"$.*,(1(",%$(1%(
&
ÉÊ
#
ÓS¶z°¹–Ž

j,IJ„½ãCþ°¾äam/2òó
(
Š ¡¢
!
è¬O¨5ŽéêÏûüB-
5"-}§/ùC„äamE
#
.´­:Ié꜇/›}JÈ
(
EFG
!
Ïûü
%
j,IJþ°
%
äamE
%
5Žéê
%
½þ°
HI2JK
!
Z=#34<
!!!!
LMNOP
!
)
!!!!!
LQRK
!
2114;1#"3
$
!123
&
1#;1<4<;13
O))&,0?")3&4=40&1301&??"(6%"0"?
8
(0%&0#,4(!
:(0#"Z#!4(07%414,0&1#?0#,?#(?*+15)090*)4+0940
?R)(,B8E:;
7
8E:
V
#
+)(,(8:
V
#
RT)*G6:
#
*R.(Z6E:;FGE:
$
/IG%%&%Lc%%O/I8M:IM
#
(E:
7
8:
V
@8E%FG6E:
V
*%&M
V
M
#
(E:
7
8:
V
#
58E:
V
96YK%[8:IM!22242
#
*G8:E
&
:>?014,0
!
+GMMLLMIJ9%LO8LLMKM:J9MEXEJMKI%:IM:JKEJ8%:9%:
V
E9M]IGE:
V
ME:OL&6%KM9IM:J
N
EKEDMJMK9E:O
E:J8%]8OE:JM:F
H
DM9XMKM9J6O8MO698:
V
JGMX8&O-"$.*,(1(",%$(1%(E9DEJMK8E&J%
N
K%[8OMJGMJMIG:8IE&KML;
MKM:IM9E:OJGM%KMJ8IE&\E989L%KJGM\KMMO8:
V
%LMI%:%D8I9E&J;J%&MKE:J
N
&E:J9E:O\M\M:ML8I8E&L%KJGM8D;
N
K%[MDM:JE:O6J8&8FEJ8%:%L9E&8:ME&UE&89%8&+GMKM96&J99G%XMOJGEJ-D",%$(1%(M]G8\8JMOG8
V
G
N
G%J%;
9
H
:JGMJ8IMLL8I8M:I
H
E:OE:J8%]8OE:J
N
K%
N
MKJ8M9EJ&%X9MEXEJMKI%:IM:JKEJ8%:JKMEJDM:J
#
XG8IGKMO6IMJGM
OEDE
V
M%LJGM9MEXEJMK9JKM99
@&
8
="1!?
!
-"$.*,(1(",%$(1%(
%
,E9M]IGE:
V
M
N
EKEDMJMK9
%
):J8%]8OE:J
N
K%
N
MKJ8M9
%
/MEXEJMK9JKM99
%
c&6%;
KM9IM:J
N
EKEDMJMK9
678"
!
12#;1>;"1
%
9:8"
!
12#;2!;12
;<=>
!
Ç©B/n)»!OP³F)(@A
%
^:¿÷v()£9IOP³F)(VY
?@AB
!
½wÇ
$
2=4=;
&#
@
#
Ç©q5a
#
,c
#
-Ñ/
#
;.f‡0/212
#
.;DE8&
!
FGE:
V
7
8E:
V
2=4=
!
E&8
H
6:I%D
!!
’“Iòm̟—‡0/›TP
#
œ‡0ù
/oÈ
#
îÖïEcL
(
nZLw]
)
2
*
Ч
#
ù
Ièi½¾‹
">b21
4
GD
!
#
J-‘$y©}
la
(
/0Ew12‡0}HIE
#
œÿž3I
é꜇0A3}Ÿ—

k-HIúG‡0/ù
C(ñ¾õ<=
(
5ŽéêÂIé껑
#
G
/JÛîéŽ
(E
h
„
*&
a
p+}ŸJéê„t]
éê
)
!
*
#
Ì­ž‡0}/›TP
#
«¬‡0ùŽ
(
Ïûü
$
-"$.*,(1(",%$(1%(
&
f»$/hՇ0
#
–ôõ}
5
;"
°½Ý]+•
#
îO;}>t‡
0
(
Ïûüœjk

’“]÷v‰Š\|E"
d
#
(ñ»S}HIE
#
Ïûü›

z&
g
h
–Ž
ÞßIO¨}k-‘’­®
#
ÏûüñBCèI
éê}b5/›
(
‡0èIéê}–Ø

§

/›]HI52
R11212
#
I•éꟗ‡0§qt}
;56z°,}C•
#
!å¹


ûy¾ø°
]
)
";#
*
#
«¬‡0}§qt
#
Q´§‡0}/ù
C«
(
YEK[E:MG
]
)
3
*
12¡¢è¬O¨
(E*&
é
ê
#
Ïûü}•ü݄áᖎ¥|y©
#
ñMÿ
¶Q²§qt
(
¿è5Žéê
#
Ïûü}§

½£äamE74Ч
(
ÕJ12ºÂO¨
5Žéê
#
Ïûüj,IJ£½ãCþ°¾¶
äam/2E
#
ijÏûü\|IÈ}-{„
äamE
#
fQ»˜½/Ièi
#
ÃT‡0äI
ÚV:klH&þ`
(
A
!
STrUV
ABA
!
STÿÓ
l>ÏûüJfÇ©Ißq/ö
(
Jt
12Q
}
R
V
*&
!
áo
21D8:
%™•Ÿ}
#
=‚t“
!#
# ½wÇ]
!
5ŽéêÏûü§¾äam/2E12
ŠŽí:
!#G
‚vÿûš&
#
t
2
"
!R%E
V
&E:O
m”ûh…
#
¬ÏûüÉʛÐ
>
œzÁ³UÂ
#
ì°
$
2>b2"b2!ID
&

3

#
ÒÛa»¼}É
ÊÃ[5Žéêp2
#
p2—FÑ
>
5ŽI¨n
¨
!
³5Ž
$
211Q
&#
31Q
#
"1Q
#
!1Q
#
21Q
#
3Q
5
Ž
#
2
"
!R%E
V
&E:O
f1yœ¤
#
p2ûst
2
"
!R%;
E
V
&E:O
m”û#6
#
ì–p2
"
°
#
m”ûì
!O
J
2
Í
#
“T}Žoé÷O©“ŠŽQ²~™
(
ABC
!
¹¿Î…z3lm
Ãzò÷O‚t
=>Q
ë*Ñÿd9÷v&Q
²z°¹ík
#
]zò³4ly‚
#
gt•¨W
è
>>3
„
>#=:D
p$Ã
SA
ð
#
Wñz°¹
E
#
#
Eh–Ž„
E
"
¦ð
(
ABD
!
V¤P¦çz€3lm
Ò"X¢ž)}Ýj
#
t
*0-)/;!
ÉRŝ
§j,•ÙøùÓSzò}띧±í
$
:MJ
N
G%J%;
9
H
:JGMJ8IKEJM
#
Y
:
&
jüG¨
$
9J%DEJE&I%:O6IJ;
E:IM
#
,
9
&
“«±í
$
JKE:9
N
8KEJ8%:KEJM
#
+
K
&
„ÙÈ
*S
!

$
8:JMKIM&6&EK*S
!
I%:IM:JKEJ8%:
#
*
8
&
]j
,IJþ°
(
*S
!
O¨f
"#1
&
D%&
.
D%&
a2
#
S
!
f
!2Q
#
df
2111
&
D%&
.
D
a!
.
9
a2
(
st²#µ
]
)
>
*
½Qd~ÓS
#
Ý\|}ÏûüyÚzò
#
Ë
±BwzSë
#
]§ð•Ôð9%´S—îïŠ
 
#
=‚Ã(
ABE
!
ÖHÃèN3lm
ËÃ
13
V
zò©®G
Y$Y
5ÿ1Ô
#
kÞ
Ÿûf
31DD%&
.
d
a2
N
R4<
²ÝžŸû
#
ÿ
#^

21111K
.
D8:
a2
ŸL
23D8:
#
ÌØûQ²òó
}ÓS
(
)am0-m$
96
N
MK%]8OMO89D6JE9M
#
/SA
&
éam0$
N
MK%]8OE9M
#
YSA
&
âE

éam
‹$
IEJE&E9M
#
*)+
&
„ZÀ
$
DE&%:
H
&O8E&OMG
H
OM
#
eA)
&
–ŽÛmn½wÇ]
)
4
*
!tJ£Ç¥}d~
ÓS
(
ABF
!
ÿV€

RE:9EJMIG
"|/ù}
ce/;!
|ŸÀ
6Žz
#
ÒXÝÓSÏûücBzòè9\|
!1D8:
‚}Ó[½
$
@
"
&
„ð9½
$
@
+
&#
qt
df
2!11
&
D%&
.
D
a!
.
9
a2
#
S„|Ÿ
#111
&
D%&
.
D
a!
.
9
a2
#
ý
1=
&
D%&
.
D
a2
.
9
a2
#
Èy
"19
(
½ãC)þ°ÌÅWñ
!
ð9m)Ž
Jù}
$
DE]8DE&
N
G%J%IGMD8IE&MLL8I8M:I
H
%LY/
0
#
@
<
"
@
+
&
@
<
"
@
+
i
$
@
+
a@
"
&"
@
+
%
m)8A
$
N
G%;
J%IGMD8IE&
P
6M:IG8:
V
#
R
-
&
R
-i
$
@
+
a`@
&
&"$
@
+
a`
@`
&%
Nm)8A
$
:%:;
N
G%J%IGMD8IE&
P
6M:IG8:
V
#
E-Z
&
E-Z i2a
$
@
+
a`@
"
`
&"$
@
+
a@
"
&%
Qu
m){í
$
EIJ6E&
N
G%J%IGMD8IE&MLL8I8M:I
H
#
.
Y/
0
&
.
Y/
0
i
$
@
+
\a@
&
&"
@
+
%
¡7§;J„h±í
$
M;
&MIJK%:JKE:9
N
%KJKEJM
#
F6H
&
F6H i
+
@
"
@
+
\b
-@Gb13b1<#
(
ABG
!
€ý2s
>R°nt
.]IM&!122
„
/Y//2=1
Š•
Ù
#
t
A6:IE:
2
9
³Èõû~Q²O¦"
(
C
!
pqr2s
CBA
!
v7¯°%wxy¹¿Î…z345
þ¼
2

#
Þß5ŽO¨}y©
#
Ïûüzò
}z°¹–Ž«¬
#
¿Ïûüz°¹
E
"
Ü-
#

5Žéꉊ
#
œz°¹
E
§+}Ÿ—¦z°
¹
®
(
I
A
!
v7¯°%wxy¹¿Î…z345
c8
V
2
!
.LLMIJ9%LO8LLMKM:JI%:IM:JKEJ8%:9%L9MEXEJMK
9JKM99M9%:JGMIG&%K%
N
G
H
&I%:JM:J9%L-D",%$(1%(
b
!
°n˜ÚÛð
j
óÈû¡š
#

>
Í
%
ÂC&–ºÂþÿ¡šûüDE
$
-
#
113
&(
Â
(%JM
!
AEJEEKMDME:9j/A%L>KM
N
&8IEJM9
#
O8LLMKM:J
&MJJMK98:JGM9EDM
V
K%6
N
9G%X98
V
:8L8IE:JO8LLMKM:IM
EJJGM113&M[M&+GM9EDME9\M&%X
CBC
!
v7¯°%wxyV¤R‚345
Þß5ŽO¨}Ü-
#
Ïûü}
-
/
$
¼
!;)
&
6
$
$
¼
!;*
&
‘’«¬
#
 O¨•Ô
211Q
—
#
Ïûü
}띧±íf¹ð
#
=¢5ŽœÿÏûü}§
qt„“«qtñ¢D}‹6qt
#
%J‹6qt
ÞßO¨}Ü-´y©
(
=

$
¼
!;A
&
-A<£ë
§±í¥H}lmla
(
Þß5ŽO¨}y©
#
S
&
=4<
!!
"
!
#
!
$
!
!"
"
$
¼
!;B
&
„
6
$
«¬
#
-
/
«
#
Ïûü­®œ
*S
!
}Mt
#

=

Ü-
(
CBD
!
v7¯°%wxyÿVhÓR‚345
þ¼
"

#
è¬O¨5Ž
#
£œ¤¥¦
#
Ç
I
C
!
v7¯°%wxyV¤hÓR‚345
c8
V
!
!
.LLMIJ9%LO8LLMKM:JI%:IM:JKEJ8%:9%L9MEXEJMK9JKM99M9%:JGM
N
G%J%9
H
:JGMJ8I
N
G
H
98%&%
V
8IE&IGEKEIJMK89J8I9%L-D",%$(1%(
b
!
)
!
띧±í
$
-
/
&#
B
!
jüG¨
$
S
&
&#
*
!
“«±í
$
6
$
&#
A
!
ÙÈ
*S
!
$
=

&
(%JM
!
)
!
(MJ
N
G%J%9
H
:JGMJ8IKEJM
$
-
/
&#
B
!
/J%DEJE&I%:O6IJE:IM
$
S
&
&#
*
!
+KE:9
N
8KEJ8%:KEJM
$
6
$
&#
A
!
0:JMKIM&6&EK*S
!
I%:IM:JKEJ8%:
$
=

&
I
D
!
v7¯°%wxyÿV€345
c8
V
"
!
.LLMIJ9%LO8LLMKM:JI%:IM:JKEJ8%:9%L9MEXEJMK9JKM99M9%:JGML&6%KM9IM:J
N
EKEDMJMK9%L-D",%$(1%(
b
!
)
!
ð9m)ŽJù}
$
@
<
"
@
+
&#
B
!
m)8A
$
R
-
&#
*
!
Nm)8A
$
E-Z
&#
A
!
Qum){í
$
.
Y/
0
&#
.
!
¡7§;J„h±í
$
F6H
&
(%JM
!
)
!
eE]8DE&
N
G%J%IGMD8IE&MLL8I8M:I
H
%LY/
0
$
@
<
"
@
+
&#
B
!
YG%J%IGMD8IE&
P
6M:IG8:
V
$
R
-
&#
*
!
(%:;
N
G%J%IGMD8IE&
P
6M:IG8:
V
$
E-Z
&#
A
!
)IJ6E&
N
G%J%IGMD8IE&MLL8I8M:I
H
$
.
Y/
0
&#
.
!
.&MIJK%:JKE:9
N
%KJKEJM
$
F6H
&
1<<
!#
# ½wÇ]
!
5ŽéêÏûü§¾äam/2E12
½þ°
@
<
"
@
+
#
R
-
#
E-Z
#
.
Y/
0
„
F6H
ûüº
DE
%
è
"1Q
5Žéê
#
@
<
"
@
+
#
R
-
#
E-Z
#
.
Y/
0
„
F6H
•¯fœ¤}
=21#Q
#
#
23<2#
>2!1Q
„
341#
£ œ ¤ ¥ ¦ û ü õ D E
$
-
#
112
&(
L4
#
Ïûüè¬O¨5ŽéêB
‹-5"-}BAJ{턡7§;J„h±
í
#
-.§qtøùãˆ"9ÒJ
#
.´»5"
-}§±í
(
CBE
!
v7¯°%wxyÖHņ‚345

#

#
5Žp2‚Ïûü
/SA
£
*)+
âElmla»¼
#
Û-AFÜ-‚«¬}la
#
´
YSA
âE„
eA)
–ŽUÞß5ŽO¨}Ü-´

(
=¢¬O¨}5Ž9G-.\}âEk-
#
˜ØãÏûüèÕvÑˆJÈ´¾š}âEa„
ø°Úéamù0
eA)
(
Þß5ŽO¨}Ü-
#
†
Ùø°éamI¨y-
#
†ÙøˆÔ}JÈ9€O
(
I
E
!
v7¯°%wxyÖHņ‚345
c8
V
#
!
.LLMIJ9%LO8LLMKM:JI%:IM:JKEJ8%:9%L9MEXEJMK9JKM99M9%:JGME:J8%]8OE:JEIJ8[8J8M9%L-D",%$(1%(
b
!
)
!
)am0-m$
/SA
&#
B
!
éam0$
YSA
&#
*
!

$
eA)
&#
A
!
éam‹$
*)+
&
(%JM
!
)
!
/6
N
MK%]8OMO89D6JE9M
$
/SA
&#
B
!
YMK%]8OE9M
$
YSA
&#
*
!
eE&%:
H
&O8E&OMG
H
OM
$
eA)
&#
A
!
*EJE&E9M
$
*)+
&
D
!
ˆ‰
Ïûü}
*
#
‡0
#
(ñ»Â¾þ†Ù
„e÷Š‹
(
Iéê]Õv
#
*
#
RS"ö
*
"
R
S(ñ99}fa
#
*
#
‡0}§dÅL˜Alf
:Ýݖ Ø R S
$
IKE996&EIME:EI8O DMJE\%&89D
N
EJGXE
H
#
*)e
&#
˜\|÷vlm
#
Ïûü}Ä&
^(ñT•}ojCW
)
<
*
#
-VI/Ïûü³ä
–Ø}ø;
(
Õ12Š Dšè¬O¨5Žéê
#
Ïûü
B-5"-}z°¹–Ž

-
/
„m){í
#

"-Þ]ê56
#
˜­:Iéꜝøùù/}J
È
(
—Ïûü,ë}-.\á
/SA
#
*)+
„
YSA
|OE}y©
#
˜ØãÕvù/}ÒJ
(
L4
5ŽéêÏûüY(ñ"d}䝋6„am
BC
#
-5‡"-}§±í
#
.´­:Iéê
œ‡/›}JÈ
#
&£§5£]
)
=
*
Ч}I•é
ê
#
ÏûüÉÊ(ñ"-}BAJ{í12¥
»¼
(
a95Žéê
#
Lñ{iwxÕ5I’
#

5õq0/›kl2Xmn
)
21
*
#
Ïûüîð‘ÃT
}>#
(ñ»S}H5Žh…}E
#
L
˜ÃTfI’õ}Fº‡0
(
Þßù‰5Ëÿ
½¾}ºOé9
#
‡Ïûü
#
TUI’õ
#
œQ
»˜©@ùõúGL5TU(ñO;}CD
<=
(
IéêÏûü
*
#
-{³ÂmRS¾¶Q
A-{}•J52Dñ¬Q»˜12
#
fÖÒH
2<<
!!
"
!
#
!
$
!
!"
"
IúG‡0klH&þ`„2Xmn
(
E
!

¬O¨5Ž
$
1
"
!1Q
&
éê
#
ÏûüB-5"
-}z°¹–Ž
#
-
/
„
.
Y/
0
#
Þß5ŽO¨}y
©
#
Ïûü
S
&
„
6
$
«¬
#
-
/
«
#
­®œ
*S
!
}Mt
%
@
<
"
@
+
#
R
-
#
.
Y/
0
„
F6H
-…«la
#
E-Z
‚…ŒÜ
%
Ïûüäamøù-./SA
#
YSA
„
*)+
âEyd
#
ñMÿØãø°éamù
0
eA)
#
eA)
Ü-ž€
(
è¬O¨5ŽéêÏ
ûüB-5"-}§/ùC„äamE
#

­:Ié꜇/›}JÈ
(
ŽLM
)
2
*
!
ZLw
#
85

I/‡0¾¶œIò/v}\|/2
)
e
*

,
:
!
()<ڕ
#
!113
!
<
)
!
*
!
Œ%º
#
²4õ
#
­˜M
#
]

5Žéꜧ—uÉÊ/›¾/
2E}Ÿ—
)
5
*

‡0)Ð
#
!121
#
3
$
2
&!
4";4<
)
"
*
!
¶O)
#
¶‘´

IéêœSÕÕÜz°¹§+¾ë§±í
}Ÿ—
)
5
*

õ9I)Ð
#
!12!
#
!<
$
21
&!
232;23<
)
#
*
!
y®¢
#
¶¢í
#
fàµ
#
]
<
–q/lmQJdT#HI
E12
)
5
*

hi)Ð
#
!12"
#
!2
$
"
&!
3#>;333
)
3
*
!
YEK[E:MG-
#
/GEGK%UG+
#
/M
H
MOe R/J6O
H
8:
V
%L9E&8:8J
H
9JKM99MLLMIJ%:
V
MKD8:EJ8%:
#
N
K%&8:M
#
96
V
EK
#
N
K%JM8:
#
&8
N
8OE:O
IG&%K%
N
G
H
&I%:JM:J8:Y6K9&E:M
$
-"$.*,(1(",%$(1%(d
&
&ME[M9
)
5
*
5%6K:E&%L/JKM99YG
H
98%&%
VH
fB8%IGMD89JK
H
#
!12!
#
<
$
2
&!
2)
>
*
!
²#µ
#
ü!!

º/‚‡04ôÞT²O§±í}ÓS
)
5
*

‡0/2)Ð
#
!122
#
4
$
#
&!
#1>;#1<
)
4
*
!
½wÇ
#
Ö6
#
ä³C
#
]

g/nÏûü}§/2E
12
)
5
*

,dæÔ
#
!12"
$
!2
&!
<=;=2
)
<
*
!
¶åã
#
äóÁ
#
[€·

I/Ïûü3ó)12
)
5
*

‡01
2
#
!113
#
!3
$
#
&!
#1>;#1=
)
=
*
!
§5£
#
¶ÔJ
#
V®­
#
]
(E*&
éêœÏûüŸJæç¾
½E}Ÿ—
)
5
*

±,õ)Ð
#
!12!
#
!4
$
/
&!
2";!24
)
21
*
­!!
#
Ü×&
#
G"W
#
]

a95ŽéꍮzÎ#„Ê#
}HIEÌSd~
)
5
*

‡0/2)Ð
#
!12#
#
31
$
!
&!
2#;!!!
$
%&(
!
)
!
*
&
!<<