全 文 ：广 西 植 物 Guihaia 21(2)：103— 105 2001 年 5月
A moss famil y Rhachitheciaceae new
to Guizhou，China
XI()NG Yuan—xin
(DP Pin t protection，GⅢ ^ Uni~ersity，Guiyang 550025t China)
Abstract{The Rhachitheciaceae is a smal[amily with one widly distrihuted and 14 narrowly distributed species be—
longing to seven genera．It was established by Robinson in 1964．Previously，no any species of this fami~was re—
ported from Guizhou province，China．The present paper reports Rhachithecium perpusillum for the first time for
Guizhou province．The description and ilustration of R．perpusillum based on the specimen from Pndin county are
presented ．
Key words：Family Rhachitheciaceae；iOtachidtecium perpusillum；New record；Guizhou
中围分娄号：Q949．709 文献标识码 A 文章编号 ：1000—3142(2001)02—0103—03
The smal mo∞ family Rhachitheciac~ae which ln_
eludes two genera Rhmhithecium Broyh．ex Le．Je1．
and Hylmodontopsis 1wats．＆ Nog．was established by
Robinson in 1964 to include two genera from Or—
thotrichaeeae：The genus Jonesiobryum Allen＆ Pursell
was transferred from Funariaceae to Rhach theciaceae
by Alien and PursclI(1991)and Zandcr(1993) in—
eluded the genus Tis 口m H4 P．de la Varde．from
the Pottiaccae to this family． Goffinet(1977)added
Uleostrum Buch．In this family too．Up to date seven
genus and 15 species have been accepted in the famly
Rhachitheciaceae(Goffinet 1 997)．
The Rhachitheciaceae may be recognized by the
Orthotrichaceae appearance with distinctive sheathing
perichaetial leaves，single peristome consisting of 1 6
teeth fused into 8 pairs and 8 ribbed capsules．(Zande
1993；0’Shea 1997；Goffinet 1997)．
Al the species of this family except Rhachithecium
purpusdlum (Thwait．＆ Mitt．)Broth．are narrowly
distributed in tropical and subtropical regions of both
hemispheres Rhach~hecium purpusillum is widely
spreading in Africa，America and Southeast Asia(Fig．
1)． In China，Rhachitheclum purpusilum has been
recorded from Yunnan province：Yunnanfu；Bei Hsin-
lung 25。24 (507)；Sanyingpan 26。(676)；Zwischen
Hsiangschuiho Sunggwe 26。17 Heding(6546)；and
Sichuan province：Helugo Yalyng Yenyuen(2480)．
(Broth．1929)．
In the htest checklist of m0sses in Guizhou
province(Xiong yuanxin 1997)Rhachitheciaeee was not
includcd．Therefore，the family Rhachitheciaceae in-
duding the genus Rhachithecium and R．purpusillum
are newly reported to Guizhou based on the material
colected from Pudin county of Guizhou．
Rhaehithec-曲ceae RobiB$OB
The family is characterhcd by ha~ng spathulate
or lingulate stem leaves；basal cells hyalme；rectangu—
lar；costa single，extending to the upper portion of the
leaf；perichaetia terminal on the stem~and peristome
singlet 16 teeth in 8 pairs(Zandcr 1993)．
Rhachlthecium Broth．ex Le Jolis．，Mem．Soc．
Sc．Nat．Math．Cherbourg，29：308．1895．
收稿 日期 2000—09—06
作者简介：熊源新(1954一)，男，救授 ，硕士生导师．国际苔藓植物学会 (IAB)~N，从事植物学研究。
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104 广 西 植 物 2l卷
Four species were described in this genus： R．
welwitschf(Dudy．)Zander．is endemic to Angola；R．
papillosum(Wiliams)Wijk．＆ Marg．is endemic to
Philippines and India；And R． ponicum (Toyama)
Wijke＆ Marg．is endemic in Japan．The following is
revised key to species of Rhachithecium：
1．Peristome lacking ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ⋯ ^．nipponicum
1．Peristome present⋯ ⋯ ·⋯ ⋯ ·-⋯ ⋯ ·⋯ ··⋯ ⋯ ⋯ 2．
2．Leaf ceils smooth ··········-·-·-·-·-··-·-·-·-· 3
2．Leaf ce Lb papilose -··-·-·-·-·-···R．papillosum
3．Annulus unistratose ·-·-‘··--R．perpusdtum
3．Annulus bistratose ·-·---·-·-··R．we[zoitschi
Rhachitheciura purpusilhm ‘Thwait．＆ Mitt．)
Broth．，Nat．Pf1．1(3)：11 99．1 909．fPlate I)．
Plant minute， simple，densely matted on tree
bark．Stems very short or a[inosl stemless，radiculose
at base，0．5～ 1．5 mrl high．Including lcaves，yellow．
ish·green or brownish．Lower leaves small，0．4～ 0．7
mm long，0．1～ 0．2 rlm wide：uppcr leaves long，
obovate—spathu[ate，broadly acute at tips，about 0．8～
1．5 mm long，0．4 ～ 0．55 mm wide：upper leaf cels
roundish—hexagonal，10～ 20,um in diameter，smooth
or mammittose；basal Leaf cells rectangular，40~ 90 p-m
long，l5～ 30 Fni wide，lighte or hyaline，thin—walled，
larger，smooth；marginal cel narrow．Costa slender，
ending much below the leaf tip．Propagula abundant on
the upper surface of leaves or the base of costa，light
green．80～ 1 40 pm long，45～ 65 p-m wide，composed
of 2～ 6 cels．Periehaetial leaves 2～ 3，conspicuous，
erec!and convolute—sheathing，oblong一~nceo[ate，1——
2．4 tP_m long．acute at apex；costa ending below apex
of tlie leaf．Seta short，erect or bent at top，smooth．
about 1．8～ 2．8 mrl long，usually twisted when dry．
CapsuIc erect or inclined，ovate—cylindrica1．symlnct
ric，about 0．54 mm high， 0．39 mm in diameter，
strong 8-ribbed and contracted below the mouth when
dry，elliptic and scarcely ribbed when moist．Annula
composed of one row of hyaline cells． OpercuIum
short conica1．Peristome consisted of 1 6 teeth fused at
basc in 8 pairs．smooth．reflexed when dry and in
curved when moist，lanceoIate，about 1 40,um high，
with reansverse striation．Calyptra cucullate，scabrous
by projection of upper cells．Spores rough，spherical，
2l～ 28,Rm In diameter．
Ftg·1 I he dlsrribution patteHts of Rhachlthecium species in the world
● R zw,1wig (hi(Dudy)zand ；▲ pdlo~ {Wili 】I )Wijk＆Ma g；●R up加 ileum(To⋯ a)
Wijk s_M g 1● R．／,e,／，a．~dh cm：Thwait＆Miu )B。rtf1
Specimen cxamined：China，Guizhou province． Yuanxin 98971
．
Pudin cotmty，Xinzhal，ah． 1 1 60 m．2．6。30。N．1 05。 The specimen 0f s c￡es 1hat was c0Ilected fr0m
75 Eon trunk of Pr“ “ ：1 5 Decembcr 1998，Xiong Guizhou is different ff0m that。f thc nTher rc ons w1th
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2期 熊源新等：刺藓科在贵州的发现 105
cell of leaves，propagulas and spores． Rhachithecium
purpusflum is characterized by having smooth or some
what mammilose leaf ceUs；large spores(21~28 gm)
and gemmae on upper sur[aee of leaves．h is different
from that of尺．papillosum and R．nipponicum．
Acknowledgements
1 would like to thank professor Cao Tong for his
comments Oil the manuscript．and professor Lin Pang-
jHail for her helps and literature， and Dr． Gou
Guangqian and Lin Yaoguang for their collaboration
dudng the field survey．This work was supported by
Natural Science and Technology Foundation of
Guizhou province．
References i
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刺藓科在贵州的发现
熊源新
(贵州大学农学院 ，贵州贵阳 650025)
摘 要：刺藓科(RNachkhedaeeae)为 Robinson 1964年从木灵藓科(Orthotrichaeeae)中分出来的 1个小科。现知全世
界有 7属和 15种，除刺藓(Rhachithecium purpusilum)~b．均为热带．亚热带分布狭窄的属种，刺藓科在中国仅剌藓属
1种 ，分布于云南和四川。描述了首次采 自贵州刺藓科的种类刺藓(尺^ ^“m purpusglum)以及刺藓科的形态特
征 。
关键词 ：刺藓科；刺藓；新记录；贵州
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符国瑷：海南锥栗属新植物 图版
．
警 3，』m
：
辫．，
因 A l叶柱A lea~es b~aneh；2雄花亭Inflorescen=e 0．0雄花 Flower ；4雌花序 Infl。⋯ ce阜：
5坚果 Nuc~la；B带坚粜壳斗 Cupula wilh nucula
凹 B 1叶枝Aleaves branch：2雄花Flowel 0；3+雄花序Infl。 ⋯ e 0；4
． 雌花序 lnfl。㈣ ce早；
5带坚果弃 斗Cupula with_1l】⋯I；6 坚果 N⋯
凹 C 1-雄托FIowe：0；2叶桂Aleave brandl； 雌花序Infloresce一 旱；4雄花序 Infl0螂 cencet；
5带坚果壳 斗Cupola with nucula；6坚果 Nucul~．
因 【J 1·带坚果壳斗 Cupula with nucula；2叶杖 Aleaves branch；3
． 雄花序 Inflo⋯ ence0，4雌花序
lafIoresce~ce旱；5坚果N⋯ 1a，6雄花 Flower 0 (王其 绘)
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