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On Paphiopedilum malipoense sp. nov.—An Intermediate Form Between Paphiopedilum and Cypripedium, with a Discussion on the Origin of the genus


Paphiopedilum malipoense S. C. Chen et Tsi is a very interesting new species with its flower similar to that of Cypripedium, especially section Cypripedium.  It belongs
to subgenus Brachypetalum, the most primitive group of Paphiopedilum, but differs from
its allied species in hgniva elliptic-lanceolate sepal with cuspidately acuminate apex, rather
narrow petals and horizontal lip, which are of common occurrence in many cypripediums, but
very rare in paphiopedilums.  Apparently, this is an intermediate form, or a link, between
Paphiopedilum and Cypripedium, but it does not seem to arise from hybridization between
them, because no Cypripedium has been found wherever Paphiopedilum occurs.
      The new species is distributed in southeastern Yunnan of China. In this area, as well
as in river valleys of western Yunnan or the Hengduan Mountains, there have been four
species of the same genus reported before. As we know, the Hengduan Mountains and their
adjacent areas are rich in Cypripedium.  The differentiation of the genus there is remarka-
ble.  All five sections it contains occur there and three of them are quite distinctive.  For
example, the general appearance of the section Bracleosa is dissimilar to that of any other
cypripediums, but closely resembles that of Listera.  It appears that the difference between
sect.  Bracteosa of Cypripedium and sect.  Brachypetalum of Paphiopedilum is not neces-
sarily wider than that between sect. Bracteosa and sect. Cypripedium of the same genus.
Apparently, it is reasonable to consider Paphiopedilum to be an evolutional branch of Cy-
pripedium extending into tropical area, with its primitive group (subgenus Brachypetalum)
still remaining in its northern fringe area.  This primitive subgenus has eight species, dis-
tributed from western Yunnan to the Malay Peninsula.  Five of them, including the inter-
mediate and primitive form published here, are found in the hilly land of southeastern Yun-
nan and the river valleys of western Yunnan.  All these facts suggest its area of origin: the
river valleys of the Hengduan Mountains and the lower hilly land contiguous to the south
of them.


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