免费文献传递   相关文献

THE TEMPORAL AND SPATIAL EXPRESSION PATTERN OF ANTHOCYANIN RELATED GENES IN RICE (Oryza Sativa L.)

水稻花青素合成相关基因的时空表达研究



全 文 :!! " # $
!"#$%!"&"$# $###+ ##$*
!"#$%&"()#*+&$,-$.*#/#$&0*.+%*+1
!"#$
!"#$")#")"$!
%&#$
!"#$")#-)#&
()*
!
./02347C,-
""#$#F%"#("# !
"6•
-
23456
"
"6
#
)78,
""#$$#%##&#
+,-.
!
u¾¤
"$+(=)# !
;
!
./¿U@
!
ÀB7CV
!
vw‘ÁJKãä%&Gü`7C
% 1)2340$[3\367oQ93<:$"=;8<2
/0+,
!
Ô¨
"$+&$)# !
S
!
./>Ã@
!
ÄÅ
!
ÀW
!
vw‘ÁJKÆ^LMkãä%&7C
% 1)2340$@QZ3<:?@X;/WX;86
1234
!$###)(--$""#$%##$)###+)#=
JK|"c؎îL*e`¸Ûç07C
/01
!
233
!
456
!
789
"
./É#R‘!"6)#7Co
!
"6•
)
./0!"#擥ÇD¢È
!
./ TU
!%$##"+#
5
!
6
!
‘a³´µ&M¶·O¸¹º†µ&»¼
!
½¾K&»¼†µ¿GP2†lÀÁ2†&I‘
è©
!
ÄōµÆÇ&vw˜
"
KÈÉÊˇp
IF,
ÌÍÎÏ4ÐGP2†lÀÑ2†&ÒÓIÂ
vw˜
"
ÔÕ
K1D,@! K15?0
l
K15?4
Ö³´µ&2KIÂ2†
!
!×F
#
Ø
#
³Ù•³Ú
$ "
Û&Ü
“+g>IÂ
" K1A%?! K1A%%?
l
K1FAG
¨!Ø+IÂ
!
!×F
#
³Ùl³Ú
$ "
Û
!
Ý
!³Ú
$ "
Û<
!
ÞIÂpTÒsOßW
" K1,)0
!àbláבa&×Fl³+>âpIÂ
!

Ú
$ "
Û<
!
ã!àb+IÂ
!
ÝIÂpTÒsOßW
" K1N$
l
K1N"
äw˜å!áבa&×F+

!K15$
æäwå!³ÙÒIÂ
"
çJ‘a¨©3ˆ&³´µÆǸÁ訩&2†
!
é>4‡&
ÒÓIÂäw˜
"
789
!
³´µ
$
M¶·O
$,.)IF,$
ÒÓIÂ
$
‘a
%"&%&./)$#1#*,+/#%#1&G/$&++)*/#%%&$*)0
#*%")!H#**$&1#%&,-&*&+*$!&"*#+,% -%.(/% 1I#
AGSfJ3)\367!ULC/4)\/4!.SHTCX)\X!BSfN46)Q<67
"O+9@&L"$&/"$9"(<8+=.&%- D$"M.%*+&%6 58.%+1+P.%.1/$9"(,-$.*#/#$+("$)#*+&$H,-$.*#/#$&0*.+%*+1!
4%1/./#/+"()#*+&$,-$.*#/#$&0*.+%*+1! <8+=.&%- :%.M+$1./9! ?&%-;8"#! <8+=.&%-!%$##"+#
#6789:;8$ D36[\38M5V/QQ4<6 0/]/0Q<\QMVX8MXV3036W
V/7X03M5V/QQ4<6Q<36M9<8[3646 3QQ<843M/W 7/6/QY/V/QMXW4/W Y4M9 Q/24)V/30)M42/IF,",.)IF,# M/8964_X/;OMQ9K15?0 36W K15?47/6/QY/V//>5V/QQ/W 46 M9/0/3]/Q! QM/2Q! \0Y/V/M9/Z3Q487/6/Q46]<0]/W 46 36M9<8[3646 Z45V/QQ/W 46 M9/
0/3]/Q! \05V/QQ46 M9/QM/2Q;O6 M9/Q//WQ<\$)"
Y//^Q3\M/V\05V/QQ4<6 0/]/0QY/V/468V/3Q/W Y4M9 Q//W W/]/0<52/6M;K1,)0 Y3Q/>5V/QQ/W Q0479M0[46 M9/
0/3]/Q<\Z038^)7V346 V48/36W 5XV50/)0/3\V48/! ZXM4MQ/>5V/QQ4<6 468V/3Q/W 46 M9/Q//WQ<\$)" Y//^Q3\M/V\05V/QQ/W 46 M9/0/3]/Q<\5XV50/)0/3\V48/! Y9/V/3QK15$ Y3QQ5/8430[
/>5V/QQ/W 46 M9/536480/3M\08<6MV<0/W Z[236[7/6/Q! Y9489 93W M/255V/QQ4<6 Q5/84\484M4/Q;
<=> ?@9A7$36M9<8[3646& Z45V/QQ4<6& V48/
!!
•„@ÙVöÉÊ`)ïŠVöóª`)É
!
@ ÙÓËÌVöãä`-‡
!
dÂÍÎÏý`Ðyq@
+
!!
"
!
#
!
$
"&
¡
ÙoLÍ`æÂzæ
%
JKiIuvw`xyijz
{
!
KòãäÑ¥æÒ
%
Kò`ÓԎvwÔÕ®
ûä
!
Ö*Ä
!

!
šVc
!
Øjä
!
ÙÍ
!
!
)
®šc
µ
!
dÂÙÍÔÕÙÄ
)
Í»Ñ
)
ÚØJy
)
ÛÜcŠ
30^[0V/Qµ
$(
%
»Ñ¤ØjgÝÞ½ÏÒOJŽŠßàÂ
!

ŠÔù
F=)F%)F=
Š
%

F%
•Žn
)
ᤊ±
”Gâ`à!
!
I»ÑÍ
)
»ÑÎÍ
)
ãäÑ
)
å
Ñ
)
›»Ñ
)
|"ccÒÍ`´æçVj
%
½èÀKò
Â
!
vwù–`|"c–éêëÀcŠìíÀc
"!%(
!
dÂéêëÀc`ù†‡ÏìíÀc`ù†
* =(
%
1
Ï|"cÂÞ½ùá~V‘
!
eØdå–ÁI`‡á
¤4
!
‰Á¯îOa¯q=4îO–{f`%
ƒ7C$”
!
èò-±jw‡`kï
&(
!
pˆðq
((
!
pˆðr
+(
!
áÿ_ýG`1*
$#(
!
šñð,
ž¿À
$$(
!
lò5óOŠ5ð,îO
$"!$%(
!
õ
$*(
!
‡ö
$-(
!
‡·÷
$=(
!
‡øù
$&(
!
¾wúEû
ü
$((
%

!
7CJK»Ñc|"c`؎–E϶
·NO`G†
!
qNý ù»Ñ`Kò
%

!
Ã`–ÀKò–Bò
!
èò
!

!
»ò
µ
!
Òý–èòßgÝ7C
!
a¯qèò-±jaþ6
wGü
& $((
%
½
-
–À
.
Kò`LM#Gü
!
à!7C
ü‰°±OžÿÀ`*e+-cy+–à!z“
%
:
¬ªµ
$+(
éIBÀOž4¼¥
$
‰#4*e°±
!
T
ÚdfgϾ
&
"¿À_`
,D(##=
Š
,D"$$(=
z
Ü
%
9!;
"#(
Šÿ֞
"$(
éIèò`ÿÀ¥
$
‰#
4*e°±
!
TÞ½)4
)
#4Štgy+
%
"¸
!
»#$éIJKOžèÀcýG¥
"
‰åg*e°
±
""(
!
:%&Š«C
"%(
ÚdÂ`
DL
*efgÏ
"-^Z
()
!
Š
G&
I!{­*e
%
P*gµ
"*(
éI
OžèÀcù†`LMç®I¯*e°±`+†4¼
LM
%

!
‰|"cVj؎îL*e`7C#Ý
8Iæw
%
½JKÂ
!
|"c`Vj؎+¼vwq,ÐÖ
Ä+¼
%
dVj؎`îL*evw–Š*eŠÝ
É*e
"
O
%
dŠ*eþ§î!-|"cVjØ
ŽîLf`*e
!
vw–,ÐÖÄ؎f*e
"K1D,@! 28+%9&&%.%+&>>"%.& 9&1+#!
ãäÑ؎f
*e
"K15?0! *8&*"%+19%/8&1+#!
ãäћŠf*e
"K15?4! *8&*"%+.1">+$&1+#!
Ȅ
)%p)
æ¤f*e
"K1A%Q?! (&M"%".6H%QH896$"B9&1+#!
Ȅ
)%)
æ¤f*
e
"K1A%?! (&M"%".6H%H896$"B9&1+#!
´æ»Ñξ
Rf*e
"K1FAG! 6.896$"(&M"%"*H$+6#*/&1+#!
|"
c¾Rf*e
"K1,)G! &%/8"*9&%.6.% $+6#*/&1+#!
R
|" c ¾ R f * e
" K1@,G! @+#*"&%/8"*9&%.6.%
$+6#*/&1+#!
|"c؎f*e
"K1,)0! &%/8"*9&%.6.%
19%/8&1+#&
ÝÉ*eþ(Im.±|"cVjç0`
*e
!
vw–
GRN
Š
5$RD
*e&/
%
fQISK
gϾ{"¿À_t
!
q,ÐÖÄ؎+
¼Â`¾{­f
%
½JK`Ò­•{–ç0
!

½9ŠÂ`ç0†uÉ
"-(
%
ØÊ
!k42
µ¾7CŸ
K15?0! K1A%?! K1FAG! K1,)0
µ*e½JK`9
!

!
Jc¥|/
")&)$%W
`ç0†$›
!
€®
K15?0!
K1A%?! K1FAG! K1,)0
½à!•g{–à!Ü7`
ç0
!
£dç0†•¥|/¸Ü`(Bê()
"-(
%
A949
µÚJK`
K15?0! K15?4! K1A%?! K1A% / ?!
K1FAG
Š
K1,)0
*eW¬î+`ìE0]^_Â
!
d
‘J`ÿÀ1¶`»ÀW^IwÀ
"=(
!
ºRÔÕ*
ekÿÀýG–L
% K15$
kJK—1`ÿÀ–
L
"& ! "((
!K1N$
Š
K1N"
kJKJßÿÀ–L
"+(
!

Š|"c؎îL*e{*¬!ÝÉ|"c½JKÂ
`ýG
!
‘êåJK`’Õ•g#¿ýwÀ
%
Y;7
CŸ  Š * e
" K1D,@! K15?0! K15?4! K1A%?!
K1A%/?! K1FAG! K1,)0#
ŠÝÉ*e
"K15$! K1N$!
K1N"#
½ûò
)

)
èòŠwJJKà!•g
!
ºc
¥|/à!¸?O‘Â`ç0$›4
!
I34JK|
"c`Vj؎5f*,
%
$!
„…kG†
BCBD
vw<=
*
Oà!`JK„…
$+%)$$"
ûò
#)AB&"

#)
6Eè™
"
èò
#
Š
O,$--""
wJ
#
Ï
"#$$
ž
=
>7
OÏ./É#ª&~8‹"£
%
H9
)
J
)
|c¥|
/à!¸?`O‘µ„…{¹Ï½LÂ
!
Ñn
%
BC5D$%J/!$
JK„…´
,HS
`-±þ
,HS4Q¡Öº
R
".3^3V3!
ÔY
#
¾5
% 8gHS
`؎þ
IV<2/73
2
:
"
}%
#
`aW;¡Ö<¾5
%
؎`
8gHS
n
H36}%
#
°dÌ7
!
T
={Ýɗ
$##67,
"
K
$`
%
½>f†
IF,

!
n
S8M46
*eÙľ5Ì7ÝÉ
!
§—
S8M46
`êtj„
?7Ø@I@
%
AB/`
8gHS
šÞÏ
*c
CR
!
D
n
%
>f†
IF,
on`djcßæÉQÃç
$%
on
`
S8M46
dj©.`Daµ
%#(
&fQISK! fQFGA
`i
(dj©.
k42
µ
"-(
&K1A%?! K1A% /?! K1FAG!
K1,)0! K15$! K1N$! K1N"
`i(dj©.
A949
#$
!$
? JK|"c؎îL*e`¸Ûç07C
µ
"=(
&fQFGO

KfFofQ#%7=#-#+

!
n
IV42/V
-E#
‰kÖ;dj
%
on
IF,
Ü}I
$ +*c
l^4
-246& +*c
^4
-#Q& =#c"
m
--c#
EF
-#Q& &"c
o
Ì
-#Q!%-
­mn
& &"c
ÙoÌ
&246&
u/šÞÏ
*c%
IF,
›/
!
…jn
"E#b
¶·`pqrsëêt4
°
%
L
BD
xHn
IF,
yz{w|}~€‚ƒ„
.3Z0/$!IV42/VQ36W M9/\V372/6MQ4?/Q<\Q/24),,.)IF,
dj*è
5V42/V632/
~Hdj
\aHdj
V/]/VQ/5V42/V
ᑐQ
\V372/6MQ4?/" Z5#
S8M46 .FSTFSSF.TTTS.TS.S.TTST TFFT..T.TT.TSS.TST.SSF %(-
fQISK TT.FT..FFFTF.F.SFF TTSSFSFF..T..TFSF.FF $#"
fQFGA TTTS.F.FTTSF.TTSSF.F FF.FS.FF.F.FF..T.FFS $#-
fQFGO .FFS.FF.F..FSFFFSF.F TSS.FFSTFSFFTFF.F $%$
fQC%G TSTFSS.TTTSTT..FSSTS F..FTS....FTSFTTSSTS *=+
fQC%qG FFTF.SFST.SFFSTFF..F .TFFSFFS...F.STST..FF *&+
fQgC, FTTT..FSTT..FSTT.SFS .TSSSFFTTSTTTST.SSF %(%
fQSHA TSSTSTTTST.TTTSTTSF. FSTSSTSFTSFFFSTTSTST -"(
fQF$ FTTT..F..F..FFSFTSF FFFTFSSF.TFSF..SSSS. %="
fQB$ TTS.TT.F.FF..TTSF.TS TTT.TTFSTS..FS.FSF.. %="
fQB" T.TTFSS.SSFTSFTSF.FF FT.SFTT.T..TSFTSTT.S %%*
ˆ
$!
|"cîL*e`ÛÜç0$›4
C47;$!.9/Q53M430/>5V/QQ4<6 <\36M9<8[3646 3QQ<843M/W 7/6/Q
n$ +%)$$"
ûò
# &,$ AB&"

# &B$
6Eè™
"
èò
# &I0$ O,,O-""
wJ
#
n$Y94M/)7V346 V48/& ,$ V/W)7V346 V48/& B$ Z038^)7V346 V48/& I0$ 5XV50/)0/3\V48/
"!
ŽkI
5CBD
…†‡ˆ7G>‰ŠL‹ŒJK
|"cîL*e`ÛÜç0$›4qþîL`
Š*eŠÝÉ*e½JKà!•go…V`$›4ç
0
%

$
ÂÃ
!K1D,@! K15?0! K15?4
½ûò
)
B
ò
)
èòŠwJJK`
!
JŠ|”–oç0
%
d
Â
!ISK
½Â`ç0†u‡
!
Ôk
k42
µ
"-(
`$W

&5?0
Š
5?4
½
*
Oà!JK`
)
J
)
|Â`ç
0†$›àR#
% K1A%?! K1A%p?! K1FAG
½
*

!JK`Â{àç0
!
Ò½Jߊ|Â{ç0
!
£½
|Â`ç0†‡ÏJßÂ`ç0†
!
8d½èòJK
`|Â
% K1,)0
½Âàç0
!
À½èòŠwJJK
`Jߊ|–¡†ç0
!
£ç0†œR#$›
%
K1N$
Š
K1N"
À½wJJK`JßÂɆç0
!

K1N"
½èòJK`JßÂÁ–¡†ç0
% K15$
À½
|Âç0
!
ཐŠJßÂç0
%
5C5D
…†‡ˆ7G>ŽŠL‹ŒBJK
|"cîL*e`¸Üç0$›4qþîL`
$$
!!
"
!
#
!
$
"&
¡
ˆ
"!
|"cîL*e`¸Üç0$›4
C47;"!.9/M/255V/QQ4<6 <\36M9<8[3646 3QQ<843M/W 7/6/Q
n$ +%)$$"
ûò
# &,$ AB&"

# &B$
6Eè™
"
èò
# &I0$ O,,O-""
wJ
#
n$Y94M/)7V346 V48/& ,$ V/W)7V346 V48/& B$ Z038^)7V346 V48/& I0$ 5XV50/)0/3\V48/
Š*eŠÝÉ*e½O‘`à!€NGæ`ç0$›
4
%

"
Âê
!K1D,@! K15?0
Š
K15?4
½
*
Oà!
JK`|Âc|/
$ "
9`O‘Â{–Ɇç0
!
ç
0†œR#$›
% K1A%?! K1A%p?
Š
K1FAG
½
*
O
JK`|Â{–ç0
!
d½èò`|Âç0†ª‡
&
|/
$
9
!
BòŠèò`O‘–Ɇç0
&
|/
"
9
!
ûòŠwJJKÂ`ç0†H()
%
´z
!
K1A%?! K1A%p?
Š
K1FAG
`ç0†ŠO‘€N`¸
܎~D
%
|/
$ "
9`O‘
!K1,)0
a›4ƽè
òO‘Âç0
!
£ç0†•€N¸Ü`()ê()
%
%!
ׁ
|"cVj؎½Á¯ijÂ{–7C
!
²I
ò
%$(
!
JK
"=(
!
QJ
%"(
Š‡K
%%(
µ
%
dVj؎L
cÁ¯*e
!
ÔÕ*eÔÕ|"c؎îL`f`
Š*eŠÝÉ|"cVj؎`ÝÉ*e
%

%
`|"cVj؎+¼ŠîLf`*eç
0ÊËÂê
!
,ÐÖÄ؎f
"ISK#!
ãäÑ؎f
"FGA#!
ãäћŠf
"FGO# %
­fqo–»ÑÍj
ä؎`*,
%
ý–2Ù`Þ½Žwå/–`f–M
j¾5{M¥`Vj؎
%

!
Ô
%
­*e½JK
ØÙo–•g{–ç0
!
£ç0†î$à¯
!
eêÂÒ
IBŽ©ç0
% K1A%?! K1A%p?
Š
K1FAG
½BòŠ
èò`O‘Âç0†ª¯
!
W’Á¯Ù͊»Ñͽ
O‘Â`ýG
!
¾êåJKŽž^®ýà!ÿÀ
%
K1,)0
q؎|"c`uæw*e
!
eI2`Ɇç
0
!
å
%)
á|"NŠ
%)
áìíNɆýG
!
¾êåŽ
ž^®wèÀ
%
1Ïûò
!

!
èòŠwJJKzÜ`ÿÀ€
à!
!
Ò*e`ç0Þ½{f`a›4
% K1D,@!
K15?0
Š
K15?4
½
!
J
!
|?c|/
$ "
9O‘Â
{–ç0
!k42
µ
"-(
Á€®
K1D,@
Š
K15?0
½9
)

)
JŠO‘Â{ç0
!
q|"cVj؎`*Yç0
*e
% K1A%?! K1A%p?! K1FAG
Š
K1,)0
½Âàç
0
!
ê½Jß
!
|?Š|/
$ "
9ç0
!
ºRÔÕ
Š*e`ç0å–B™‹Œa›4
%
½|/`à!
¸?Â
!K1A%?! K1A%p?
Š
K1FAG
`ç0†•O‘€
N¸Ü`(Bê()
!
ºRÔÕ*e`ç0kKòÿ
À–L
!
£•„Kò÷ø`ýGê()
% K1,)0

4ƽèò`|/
$ "
9ç0
!
£ç0†ÝÞ()
!
ºR†*ekKòèÀ4¼–L
%
eI
K1,)0
!-
`|"cfwÚR|"cW^Ž|"c
!
e؆*e
`Ɇç0å|"c½èòÂɆýG
% A949
µÚ
K15?0! K15?4! K1A%?! K1A%p?! K1FAG
Š
K1,)0
W
¬î+`ìE0]^_Â
!
d‘JÿÀÁí1RÓ`
»À^Ià!Ü7`wÀ
!
ºRÔÕ*ewùå|"
cýG
"=(
%
K1N$! K1N"
q
D
`µg*e
!

>9*)
©W;

!
‘wJJKJ߀®TÖ×
!
)ü`‹¯½Ï
K1N"
`
-p
PÞ½Ï
K1N$
çQ`è
$#^Z!

K1N"
(¡šj`
F
NP
%*(
%

!
À½wJJKJßÂÉ
†ç0
!
½èòJKJß¡†ç0
!
ê½ûòŠBò
JKJßÂàç0
%
½ˆ
%
Â
!
U–ÿýÔ
"
­W;
e‘`ç0
!
eI-2ÙÝ°R­*e¾àá
â
%
½èòŠwJJK`JßÂ
!K1,)0
`ç0†à
"$
!$
? JK|"c؎îL*e`¸Ûç07C
ˆ
%!
|"cîL*e½d؎+¼Â`ç0ÊË
C47;%!1>5V/QQ4<6 <\M9/7/6/Q46 M9/
36M9<8[3646 Z4n$ +%)$$"
ûò
# &,$ AB& "

# &B$
6Eè™
"
èò
# &I0$ O,,O-"
"
wJ
#
n$Y94M/)7V346 V48/& ,$ V/W)7V346 V48/& B$ Z038^)7V346 V48/& I0$ 5XV50/)0/3\V48/
ç0
&
$àç0
& A$

& K$
J
& C$
|
& A$$
|/
$
9`O‘
& A"$
|/
"
9` O ‘
% FÛ Ü S T f
S& F[364W46$
| " c
&
g49[WV<^3/25\/V<0$
´ æ U V Ù
& g49[WV<_X/V8/M46$
´ æ W ž c
&
1V4XYÙ
& K/X8<8[364W46$
œÀ|"c
& D30<6[0FдST
f
S& H3V467/646$
Z ž c
& H3V467/646 89308<6/$
Z ã ä Ñ
&
I9/6[05V<536<4W 53M9Y3[$
,ÐÖÄ+¼
% SHA$
|"c؎f
& FGO$
ã
äћŠf
& FGA$
ãäÑ؎f
& gC,$
´æ»ÑξRf
& C%G$
Í»
Ñ
)%)
[*WXf
& C%pG$
Í»Ñ
)%p)
[*WXf
& ISK$
,ÐÖĘif
%
/>5V/QQ/W&
$
X6/>5V/QQ/W& A$ QM/2& K$ 0/3\& C$ \036M9<8[364W46 Q[6M93Q/$ FGO$ 89308<6/4Q<2/V3Q/& FGA$89308<6/Q[6M93Q/&
gC,$ W49[WV<\03]<6<0*)V/WX8M3Q/& C%G$ \03]<6<4W)%)9[WV<>[03Q/& C%pG$
\03]<6<4W)%p)9[WV<>[03Q/& ISK$ 59/6[0303646/322<6430[3Q/
¯
!
Òq1Ï
K1N$! K1N"
`ç0
!
åwJJKÂ`|
"cɆýG
!
€°
K1N$! K1N"
Š
K1,)0
Âw{*
¬!ÝÉJKJßÂ|"c`ýG
% G4Vµ¶·W
*e`G†ÁÝý![`
%-(
% K15$
k—1`ÿ
À–L
%=(
!
À½|?¸ç0
!
½d!‹ŒŠ¸?{à
ç0
%
ºR†*eÀ½|Âç0
!
ÁºRJK—1`
ÿÀ½|?¸\ºߐ]
% A34M<9
µ€®™KŠZ
K`
K15$
¾
%
­Ê#‘Þ½
"
Oà!Í©`(^
!
W
’¥Ç_`a`à!
!
¾êW’ç©`¯[4
%&(
%
e
ؽûò
)

)
èòŠwJJKÂ{w4°1†*e
`ç0
%
´z
!
|"cVj؎`îL*eå–{f`¸
Ûç0a›4
!
–ÕkKòÿÀ–L
!
–ÕkJßÿÀ
–L
!
–ÕkJK—1ÿÀ–L
%
‚/+¾{Ñ7C
|"cVj؎îL`W;e‘`Ý°Þ±
)
Ö×è
ò`*ecd$›ç0ހ
!
I(ÝèÀ¬­Kò-
}€‚ƒ
%
st1u
!
$ (!
‹
!
b
!
?
!
c
!
Ô¨
;
JKÓԎLMNO7C¾Ñ
N(;
!"#$
! "##(! """*# $ *-- *`=#
" (!R9367Dn! R9367,C! R9367CU! K4X ,G;I9/6<0485V<\40/Q
36W 36M4<>4W36M38M4]4M[<\Z038^ V48/ZV36 <\W4\/V/6M8<22/V8430[
3]3403Z0/]3V4/M4/QN(;NS7V48C< % (!R9X C! F34JR! B3)4VV3W43M4<6 <6
59/6<0488<25&`&
* (!K//NG;OW/6M4\483M4<6 36W _X36M4\483M4<6 <\36M9<8[3646Q\V<2M9/
7V346Q<\Z038^ V48/"K$9;& 1&/.M& K;# ]3V4/M4/QN(;C<B4 - (!A<25<67,! A4/Z/6936W0)196 A! K46QZ/V7/V)D3VM46 T! B/V79<\/V
1;I9[Q48<89/2483036W 36M4<>4W3M4]/5V<5/VM4/Q<\V/W 36W Z038^
V48/]3V4/M4/Q\V<2.934036W! F946336W AV4K36^3N(;C<"#$$! $"*$ $%" $`*#
= (!.9X66<5 K! F93V0/QCA! A3^W3N! e3663.;S6M4<>4W36MQ36W
36M4<>4W36M38M4]4M[<\Q/]/V305472/6M/W V48/ZV36QN(;.9/NS7V48
C< & (!D46 A n! ,[X A H! k42gG;S6M4)46r3223MV48/! 8[364W46 `% `f`
$
`g)70[88[364W46 36W 5V"#$#! $#$ +-+ +`==
( (!J367J! S6WV/YQDF! GX J! n367gK! P46 J! R9X J! H4G!
K467n G; S6M9<8[3646 />MV38M\V<2 Z038^ V48/Q4764\4836M0[
32/04W[Q0454W/248V3MQ46WX8/W Z[9479 \3MW4/MQN(;NS7V48C<"#$$! -+$ =&-+ =`&=*
+ (!A<6 DN! ,48\/VX048384W <6 M9/70X8N(;NC<$#(!.363^3N! H3^364Q94.! f73Y3k! .QXVX23k! A9423?3Y3D!
%$
! " # $
!"#$%!"&"$# $###+ ##$*
!"#$%&"()#*+&$,-$.*#/#$&0*.+%*+1
A942MV38M36W 36M9<8[364W46Q<\M9/
8<6QM4MX/6MQ5V.% M.M"N(;NS7V48C<$$(!
P*g
!
Pde
!
³f/
!
~=´
!
ghi
!
DTj
;
èò|À
N‰ð,ž¿À·á¤kl`šñin
N(;
ÓÔ#$
!
"##=! "( "%# $ "$= "`"#
$"(!kV4Q` 1M9/VM<6 ID! G/8^/Vk g! B<636<2/S! F<]30A D!
B46^8<25W4Q/3Q/36W 8368/VN(;S2ND/W! "#$$! $$%$ &$ (`(
$%(!n367P! G36 I! R9367D! U43D! R9X G! D3N! GR! K467n;AX550/2/6M3M4<6 <\Z038^ V48/5472/6M\V38M4<6 425V<]/Q
36M4<>4W36M36W 36M4)46\03223M9/3VMW4Q/3Q/N(;SQ43I38NF046 HXMV! "##&! $=$ "+- %`#$
$*(!
bm
!
³BS
!
:C
!
¢n
!
.o
!
p;ª
;
èò|ÀN-±j‰‡
qqyÆWÉrôõHŽ`‚ƒ
N(;
yã)#
!"##(!"+ ""# $
%&= %`&+;
$-(!F9367G! JX B! JX UI! J4K! F9/6 FJ! D4D.! K467n G;
S6M48368/V38M4]4M/Q<\36 36M9<8[3646)V489 />MV38M\V<2 Z038^ V48/
37346QMZV/3QM8368/V8/0Q.% M./$"36W .% M.M"N(;HXMVF<68/V!
"#$#! =""(# $$$"( $`$%=
$=(!
X&M
!
oÞD
!
sHb
!
‹ÞD
;
èò|ÀN-±j‰Ér‡
·÷in
N(;
Â%2¬tV
!"#$$! "&"-# $ =$% =`$-
$&(!
u;S
!
X»V
;
èòÀc‡øù`V¤Þ€c7CÑv
N(;
yãs6)1
!"#$#! %$"=# $ %&" %`&*
$((!GMV38M
\V<2Z038^ V48/"K$9;& 1&/.M& K;!&2"%.*&# <6 89V<64830[308<9<0)
O6WX8/W 04]/VW3237/46 V3MQN(;NS7V48C<%$+$ %`$+=
$+(!
:»
!
Xwx
!
JQ^
!
:6;
!
yz[
!
4=á
;
BòÀc
„Vg¹cd*efg
N(;
7C${
! "##+! "+""# $$& "`$
"#(!
9!;
;
K®èòOž`LMa47C
N(;
|}"6É##
$
! $++-! $("*# $*+( -`#-
"$(!
ÿ֞
;
JKòwÀŠ4LM7C
N(;
LM
! $++(! "#$$#%
$`#*
""(!
"¸
!
»#$
;
JKãO95OžÀc`LMI¹$
N(;
g>"6)#
! $+((! %$- =`
"%(!
:%&
!
«C
;
JKwÀOž*e
IZ
`õ¿fgk,u*e
I
N(;
)#¶$
! "##&! -"""$# $"-$& "`-"%
"*(!
P*g
!
~R
!
Dú€
;
èòKOžÀcù†`LMy+
I
N(;
Â%JK)#
! $++-! +"%# $$*+ $`--
"-(!k42BT! k42NG! D46 A J! A946 kG! k42NG! k42GJ! ,[X
A H! S96 NG;S6M9<8[3646 8<6M/6M46 V48/4QV/03M/W M5V/QQ4<6
0/]/0Q<\36M9<8[3646 Z4""# $ $-= $`=#
"=(!A949 FG! F9X G! .367Kk! A3^32k;! n367DC! K7/6/Q46 V48/\03]<6<4W Z4$`#-*
"&(!.3^393Q94D 1;S630[Q4Q<6 3548X0XQ8<036M9<8[3646 8<0-#$"== %`="
"((!.3^393Q94D1;T/6/3630[Q4Q36W 4MQV/03M/W 5VS7VG<^^34W"+(!A3^32D! HS
30/0/93Q38<250/>Z3Q489/04>)0<<5)9/04>5VN(;I036M8/0594Q4<0! "##$! *""+# $ +(" +`+$
%#(!
`Da
;
JKˆ¤Ä]^*e`fk7C
g(;
TU
$
./É
#
! "##(
%$(!A93V23D! FF9<5V3 A; OW/6M4\483M4<6 <\M9/ D$$ 7/6/ 5V36M9<8[3646 Z4$(("$# $ =+ $`$#
%"(!S92/W H! D3/^3Y3D! H/>5V/QQ4<6 53M/V6 <\36M9<8[3646 Z4Y9/3M8<0/<5M40/QN(;B4<0<743I036M3VX2! "##+! -%""# $""% "`"(
%%(!A949 FG! F9X Ok! J45 n k! K5V/QQ4<6 <\MY<
\03]<6<4W %p)9[WV<>[03Q/ 8gHS 46]<0]/W 46 Z4[36M9<8[3646 59[M<30/>46Q46
Q$*$+
%*(!n3M3VX A! .3^X f! k/4QX^/k! F94^3V3D! S^ 4V3A! D46k3?X94^D
S
30/0/93Q38<250/>/68)0<<5)9/04>5VZ4%-(!G4V36M9<8[3646 388X2X03M4<6 46 MV36Q7/648V48/Z[/>5V/QQ4<6 <\V48/
K1N" X6W/VM9/8<6MV<0<\V48/5JD&","$ 5V<2C<%=(!,/WW[eA! A89/\0/VB1! n4/636W L! n/QQ0/VA ,! ,/WW[S,;
F0<646736W 893V38M/V4?3M4<6 <\M9/V48/9<2<0<7X/<\M9/234?/F$
36M9<8[3646 V/7X03M*+(
%&(!A34M<9 k! f64Q94k! D4^324O! .94W3Vk! A36W4]/VQ4\483M4<6 3MM9/F"fQF$ # 0<8XQ<\Y40W 36W 8X0M4]3M/W V48/$
6X80/$=($ ++& $`##&
*$