免费文献传递   相关文献

Physiological Adaptation of Sorbus pohuashanensis Seedlings to Heat Stress

高温胁迫下花楸树幼苗的生理响应



全 文 :!"#$%&
!"#$$!"%"&#$,#" ,#*
!"#$%&$%$(#)*
!!
!"#$
!$##$)$%*""#$$##&)#,#")#6
Ò¥DEŸFGz»‹+ sœH
f
!
†
$!"
!
ˆ‰Š
"
!
€
!
‹
$
!
FŒD
"
!
!
Ž
(
!
!
!

(
!
‘’“
%
"$a
×Qiª$+M#$$)
!
Vª ×Qi
!*("###% "a
(!"#$%&)!"%&2
!
(3!"4!fÙ{56789
!
:;
!$###$%
(a
:;X…!¸“4uT! 
!
:;
!$###(% %a
O!‚«ä!"4Lʪ`
!
O! «ä
!$(("###
<=>?
$ "#$#)#,)""
@ABC
$
M8„Nv#%)2@w#%"BC¿BABC
"

$"##6#6#
e(3#EÖyÊѬ„
""##,P.4$(P#6#
MNOP
$
’
!
U
"$*(&#!
S
!
5´Y
!
R]%&+
+
!klMN
$
²ÄT
!
%%&ž
!
R]+`…
+2)789:$ X?8>K;?=;8<+8;+;>
67
!
Z
%# À
$
í+›é¨ÓԉŠ
#"%,*$# ?
¢ôõö
!
’Ötõö,L„¸Ð±v

¬ `
3

­‹®
" 4^.#
±v

Úé“j¯“õ
"Dg4#
rv

7Wãåóé“jõ
".3d#
rv

„¸Ð°\½¾´w„¸
™Ú¾”§|+}+“eÝ,4$
(
”•Ÿ 
$
¢ôõöÀ„¸Ð±vÀÄ
! 4^.
±vpú
!
¬ `3pª
!
Ú
é“j¯“õ
"Dg4#
7Wãåóé“jõ
".3d#
±vypú7ÄÕ
%
6iõö,LJD
!
¡ª\vi§v
"/LY
/7#
7—\“$=3
"
*
3D
!
#
\“$±²x¾
"U3#
ßgÀÄd[
!
[\“$±²x¾
"E3b#
gµhd[
(
¢ôõ
ö
# , ?
„¸™Ú¾”§@wjËz+š“
!
å]¢ôõö
* ?!
„¸™Ú¾”§z+‹Pªš“
!
qÔ¦4ë5V
W
!
@³ö\š31Øz­¦4VW
!
@qö\Ç^´µ

Âs¶{|^«
(
89:
!
¢ôõö
%
›é¨ÓÔ
%
„¸Ð
%
„¸Ð°%
7é“xy
%
„¸™Ú¾”§
;<,=$
!D6"a"&
!>?@A
!.
1"25/%&%3/)&:,)4.)./%-%012#0&(2.&*.*3%3!G##,&/-35.% N#).G.*#55
:Q67N"-3
$!"
! DEQ672"-38\+
"
! =5=+("
$
! DE567,*/(-8*"-3
"
! P(
! 014("
(
! 612(-8%*$-3
%
"$+-I:BKBI;B! D?9?Bf9_>9LBCA9H@! D?9?Bf9!*("###! d9>Q98>K! -?9>8% "+0BAB8C;? 1>AH9HJHBIBAB.;8MB7@I5B@N8OIC8HIC@IK8>M -J:H9L8H9I>! DH8HB/ICBAHC@.M79>9AHC8H9I>! PB9Q9>K!$###$! -?9>8%
(+d9A?8> /ICBAH/8C7IKN8>MA;8G9>KDBCL9;BA! PB9Q9>K!$###(! -?9>8%
%+R?B4BA9K> RB87IKU9>K/ICBAHC@PJCB8J! W8>KU9>K!$(("##! 9`:9>! -?9>8#
:85.*).$ R?B;?:ICIG?@:<:JICBA;B>;BG8C87BHBCA! H?B;I>HB>HAIM 78:I>M98:MB?@MB" 4^.#!
AJGBCIe9MBM9A7JH8AB"Dg4# 8;H9L9H@! 8A;ICO8HBGBCIe9M8AB".3d# 8;H9L9H@8>M ;?:ICIG:8AHJ:HC8AHCJ;HJCBIB)
@B8C)I:M N"#G/%B"*/(%*(-$-%+%ABBM:9>KAAHCBAABM O@?9K? HB7GBC8HJCB"%# \# 9> KCISH? ;?87OBCSBCBMBHBC79>BM
HIB:J;9M8HB9HAG?@A9I:IK9;8:CBAGI>ABHI?B8HAHCBAA+R?BCBAJ:HA9>M9;8HBM H?8HH?B;I>HB>HAI;CB8ABM! B:B;HCI:@H9;:B8[8KB8>M 4^.9>;CB8ABM!H?B8;H9L9H@IM .3d9>;CB8ABM 8H<9CAH8>M H?B> MB;CB8ABM
S9H? H?B9>;CB8A9>KH97BIHJ7@9B:M"/LY/7#! 8;HJ8:G?IHI;?B79;8:
B<9;9B>;@"
*
3D
!
#! 8>M G?IHI;?B79;8:UJB>;?9>K;IB<9;9B>H" U3# MB;CB8ABM Be;BGH 9>;CB8ABI<>I>)G?IHI)
;?B79;8:UJB>;?9>K;IB<9;9B>H"E3b#+-?:ICIG:8AHJ:HC8)AHCJ;HJCBS8AABC9IJAM878KBM O@* ? ?B8HAHCBAA! M9AG:8)
@9>K;I:8GABM ;?:ICIG:8AHB>LB:IGBA!O:JCBM KC8>8:87B:8B8>M M9AICMBCBM AHCI78:87B:8C+1H9A;I>;:JMBM H?8HH?B
ABBM:9>KAIA9H9LBHI?9K? HB7GBC8HJCB+1H9AGCIGIABM H?8HH?BABBM:9>KAA?IJ:M OB;J:)
H9L8HBM J>MBCG8CH98:A?8M9>K8H:ISBC8:H9HJMBA+
;#2 <%*,5$ ?9K? HB7GBC8HJCBAHCBAA%N"#G/%B"*/(%*(-$-%+%"T8>;B# TBM:%;?:ICIG?@:;I>HB>H%;?:ICIG?@:<:JICBA)
;B>;B% 8>H9Ie9M8H9I> A@AHB7%;?:ICIG:8AHJ:HC8AHCJ;HJCB
"&
? ’
!
U|
$
¢ôõöÀ›é¨ÓԐ+}”m
›é¨
"N"#G/%B"*/(%*(-$-%+%"T8>;B# TBM:#!
Þ8…››é

ɩݎ
!
ÄÅ#
"0IA8;B8B#
Ÿ•·
#
" 8^:I9MB8B#
›éû
"N"#G/%#
l„+nf
!
^_h
i¥(@?

O!

·Á

óEF

©¸¯

¹u


Tu:¯

Q:

TUu¯w:ºU¯
!
+¥€¶
## " &## 7
áÈRTÝ!
*$+
(
ݎ
¨]:;TM¥»ðג¢™¥¨c
!
C–(ó
j˝Ɠc4IÈMmn¥4X¸“
(
›é¨
Z:;W¨ÈM€¶
$ "## " &## 7
TÈËÍÎ
hi
!
£:;¼ƒÜM…›T

½¾5¿Àܼ
!
Q
:ò|T
%
Ê:;

LÁ|̾©:ÈM4X€¶

$## 7!
/›é¨TM+.€¶Qɪ¥
&## 7!
N`͛騐î“jÙBìPª
(
쥀
¶QÉ
!
-.¤i£>Š

ôì

Åì

¢ÏÌz+
š“
(
Çc+,¤irsÈMn¥!f
!
R1ê¤
iZ¨f+Dz{1ê¬÷«^`Mn’i“c
h
(
(ó›é¨“cjÙ%&&5P
•
(
ZX|
*"+
ú›é¨`
$ *## 7
“4
,## 7
T
È
!
{Ôj٘3îïÆ
(
“c4Հ¶›é
¨ÓÔZ¼·Œ§+#„



Õ|^«
*(+
(
€Ü
c^«^_¤
!
C–%&#j˔~
(
w%
&›é¨êí+/jԉŠ¢ôõö
!
%&‹›
é¨ÓÔZ¢ô„.Àg»Ôxy

„¸™Ú¾”
§

01õxy´w\sxy+}”m/òeš
“¬~
!
š›é¨“cjÙ

Vc¹{eæÞì
s&#$}~á³
(
$!
./012
=+=>
:;IJ
"##*
í
(
Ç
"#
>
!
ú-u¥TUWT›é¨
$
í+Ԙ¥(&š
(# ;7
›/
!
´
6&F
Ùš
s
"&F
Ãך@q
(
Z(!"#$%&)ô9
ó‰ŠÙÚ
(
ô9f¯ËLü
!
,6,®)
!
0ñ]
¡Äk
%
ô9ìÅÆò
!
Ç\sq
%,&
Ç
*$##
,IÌ¡ôš
", \!
I̬Åìš
6&F%$($##
IÌ¡ôš
(( \!
I̬Åìš
&&F(
®ÓÔÆׄy7
!"##*
í
,
Ç
$"
>,¹D[
ê͐
&
S/jÓÔm¸Åì
6#F
\Šœìš
$ $##
#
7I:/7
V"
/A
V$
!
¡ôš
%# YØ¡¸s
320-1n.N"2)(,T#
hT‰Š
"%,*$# ?
¢ôõ
ö
(
`VS²@ãµ
$Y(
p’Æׄà
!
‰Š+}+
“

°\½¾|4$’
!
ÇB4$’Ì5Å
&
Ä
(
¢ôõö–ÓԚŠ
"-5#(
=+@>
pq
$+"+$!
UVWXY@Z[3\]^MR(:
!
g
»qÔ¬Çv

­‹®
" 4^.#
±v’½Š
KÈ|
*%+
}~
(
$+"+"!
_`[(:
!
Úé“j¯“õ
"Dg4#
rv
’½ŠKÈ|
*%+
}~
!

&
Ä
(
7Wã
åóé“jõ
".3d#
rv’½ŠQs+|
*&+

}~
(
´
$ 79>
ó
g4
"#
š“
#a#$
’Zš
$
üõr
Óá{
(
ó8”•µXà
"
#
/K
V$
#
Ÿ—
(
$+"+(!
abMR(:
!
„¸Ð±v’½ŠÊ

*,+
}~
(
$+"+%!
cdbe>89(:
!
n
3.^ )"$##
„¸
а\$’Ötõö,L7—\“$=3
"
*
3D
!
#
\“$±²x¾
" U3#
¡ª\vi§v
"/LY/7#
[\“$±²x¾
"E3b#
|„¸Ð°½¾
!
ǽ¾¾×ÌZ¹’1mÀxyÍ2Ê.+

!
’–„öÔum
(# 79>(
$+"+&!
cd5fghij(
!
’Ötõö,L
y„ö
!
n=~<>ô
$a# 77j$a# 77
+
R¾©R
!
m¸VXò
$F
Ê‹®ÇÈá
’
!
MËåá’
!
µG*
!
MÇ8œ½`
!
Î7n-
騝
23I>)*$"
ÿÌ
!N5P
ýÚqôö
!
Íå˜é
ÎeÏÐå˜5Т
!
Z
2`^ )$%##
ý ÑÀ¥¦„
¸™Ú¾”§
*6+
(
$+"+,!
9klE
!
ùn
D3DD
9H‰Š}Qhqe
ND4
Ì5±P
(
"!
34567
@?=>
Ò¥DE•FGz»‹&7K˕LS
$
$A:
ä©+›œ
›é¨ÓԄqÔ¢ôõö±PQÏ
!
ìˆ
$

$
6õö,LJD
!
qÔÇv>?pª
(
õ
ö
#% ?
7qԐI̬ÇvhTš
$%+*F
$6F%
õö
%*$# ?
hT±õö
# ?
pú‹
(&+$F
&#F,F(
ˆ
$!
¢ôõö›é¨ÓԄÔÇv“”
(#,
!!
"
!
#
!
$
!
%
!
& "
"%
#
ìˆ
"

$
6õö,LJD
!
›é¨ÓԄ
­‹®
" 4^.#
±v>?pú
!
õö
#% ?

4^.
I
̱vhTš
%a(*a&
#
7I:/K
V$
%
õö
%*$# ?
± õ ö
# ?
h T p ú ‹
6"a%F $##a*F
$,&a6F( !
ˆ
"!
¢ôõö›é¨ÓÔ­‹®
" 4^.#
±v“”
@+@>
Ò¥DE•FGz»‹MÉS+›œ
Z¢ôõöÀ
!
›é¨ÓÔ
Dg4
rvŸ^šy
µh7ÀĐd[
"
ˆ
(#(
Z¢ôõö–
" ?!Dg4
rvgµhd[
!
¢ôõö
" ?!Dg4
rvF4¡¢
×
(6%a,( _/K
V$
!
]õö
# ?

$a",
N
( Dg4
r
vpœË=Èä΋˜j™órvé
!
Ӑ‹
Íì@˜j/™†Ø
%
å]6õö,LJD
!
Dg4
rv>?ÄÕ
(
õö
% ?!Dg4
rvÄÕ4
",(
_/K
V$
!
š
# ?
õö
**+&F(
õö
, $# ?!Dg4
rvžŸZ
"%# _/K
V$
Ò±
(
NØ 
Dg4
Mšr
véäÎxy7é“õ
!
Z›é¨ÓÔ¢ôõ
öî?zӋ꒐7é“Mn
(
ˆ
(!
¢ôõö›é¨ÓÔ
Dg4
±v“”
.3d
šäÎ
T
"
g
"
xy
!
7Wãå
V
ÝÔ©Õ
".A.)ZDT#
›-®>õ
**+
(
ìˆ
%

$
õö
# ?!.3d
rvš
,+6& _/K
V$
%
¢ôõö
, ?!.3d
r
vÇ^¡ª×
!
š
$"+"& _/K
V$
!
]õö
# ?

$a*$
N
%, $# ?
Ÿ!ÀÄ
%
õö
$# ? .3d
rvš
(+*(
_/K
V$
!
]õö
# ?

&6F(
6,LJD
.3d
r
v>?ÄÕ
!
OûÕ¥
# ?
õö
(
@+E>
Ò¥DE•FGz»‹&N8ä©+›œ
}Qhq”•
"
Ÿ
$#
Ÿ 
$
„¸Ð
8
„¸Ð
O
„¸Ð
8mO
e„¸Ð
8YO
Ì6õö,LJDÊ
ˆ
%!
¢ôõö›é¨ÓÔ
.3d
±v“”
ÀÄ
%
΋„¸Ð
O!
Ž‰½¾ZÖtõö,LQ
É̖j
(
Ÿ
"
Ÿ 
$
Z¢ôõöÀ›é¨ÓÔÆ×
„„¸Ð±vÀÄ
!
õö,LjD
!
„¸Ð±vjÕ
(
¢ôõö
*$# ?
„¸Ð
8mO
±v–j
":k#+#&#
±õö
# ?
ÄÕ
(
/
# ?
õö¬±
!, ?
õö7„¸
Ð
8
Ç^–jÀÄ
":k#+#&#!
Àċ
"&F(
Z¢ô
õöÀ
!
„¸Ð
O
„gÀÄd[
!
å]QÉz֖j
":l#+#&#(
/„¸Ð
O
±
!
¢ôõöÀ›é¨„
¸Ð
8
ÀÄPš –
!
2´`̈́¸Ð
8YO
×ÄÕ
(
ìÜ,•
!
¢ôõö›é¨„¸Ð
8
“”Pª
(
w
=>
Ò¥DEŸFGz»‹&N8ä©+p<,5
4$ QÉu ÛQI}e Íìì Ì}
!
×
œL
#+#"& & #+##&
„¸Ð
O
ϗ
#+#&# $" #+##% $+$*6
ËÊ
#+#6& $6
œL
#+,$$ & #+$&
„¸Ð
8
ϗ
#+&,( $" #+#%"
(+$%"
!
ËÊ
$+$6% $6
œL
#+& & #+$"
„¸Ð
8mO
ϗ
#+%,( $" #+#(
(+$#,
!
ËÊ
$+#,$ $6
œL
#+%% & #+#
„¸Ð
8YO
ϗ
#+(%" $" #+#"
(+$&"
!
ËÊ
#+6$ $6
!!
ô
$!
#+#&
"&!$6# o"+*$
w
@>
Ò¥DE•FGz»‹O&&N8ä©+›œ
õö,L
Y?
Ī¸Ð±v
Y"7K/K
V$
#
„¸Ð
8
„¸Ð
O
„¸Ð
8mO
„¸Ð
8YO
#"

# $+(%8"$##+## #+%*8"$##+## $+*"8"$##+## "+68
" $+((8"+"# #+%68"6+# $+*$8"+&# "+*(8
% $+$$8O"*"+*# #+%%8"$+6# $+&(8O"*%+$# "+&";
, $+#$O;"6&+%# #+%%8"$+6# $+&$8O"*(+## "+(#B
* #+*;"6(+$# #+(*8"6+"# $+(6O"6&+(# "+&*O
$# #+$;",6+# #+(*8"6+"# $+(,O"6%+6# "+(M
!!
ô
$
ŸtsÖtöVLŸ—Qɖj
":k#+#&#!
!äó¾ö
š/Š…h±
(
%#,
"&
? ’
!
U|
$
¢ôõöÀ›é¨ÓԐ+}”m
@+H>
Ò¥DE•FGz»‹&N8PVk+
›œ
[õöGHÀ¡ª\vi§v
"!LS!?#
š“
g+
!
Ö¦jce+DGH“”
!
õöGHÀ«½
¾ –ÀÄ
*+
(
£ˆ
&

$ !LS!?
Zõö
% ?
7
&Ë2ÀÄ
!
õö
# ? !LS!?
š
#a*$$!
õö
% ?
š
#a66$!
±–NÄՋ
%+F(
Ø ¢ôõö
% ?
7›é¨ÓÔ\xy!
"3D
!
#
&Ç^Öt0ì
Øˆ
(
ˆ
&!
¢ôõö›é¨\xy!¡ª\vi§v
"!LS!?#
“”
7—\“$=3
"
*
3D
!
#
š“¬~£ˆ
,

$
Z¢ôõöGHÀ*
3D
!
KóÄÕ
!
Ötõö,L/
ÖõöQɖj
":k#+#&#!
hT]õö
# ?

6&F6%F ,"F %,F%(F(
Ø 7—n¥\“
$™m\ ivZ=•
!
`ʓ”˜j\s
Mn
(
ˆ
,!
¢ôõö›é¨\xy!\“$=3
"
*
3D
!
#
“”
„¸Ð°\\“$±²x¾
"\:#
]Ÿ^
3D
!
™mv&m0ì5_×
*$#+
(
£ˆ
6

$

ôÀ
!\:
ÒgÀÄd[
!
Ø \™mv&m0ì
ÄÕ
!
å]Ötõö,L/õö
# ?
QÉ֖j
":l#a#&#(
ˆ
6!
¢ôõö›é¨\xy!\“$±²x¾
"\:#
“”
„¸Ð°\[\“$±²x¾
"6:M#!
™t
3D
!™mv[Ö××vçN×Ӑª+
*$$+
(
£
ˆ
*

!
Z¢ôõöGHÀ
!
6iõö,LJ
D
!6:M
>?h¢
(
ÇÖtõö,L/õö
# ?

Qɖj
!
Ê­Ötõö,LQɖj
":k
#a#&#"
Ÿ
(#(
Ø Z¢ôõöÀ›é¨
3D
!úª
‹yçNÝm
(
ˆ
*!
¢ôõö›é¨\xy![\“$±²x¾
"E3b#
“”
w
E>
Ò¥DEŸFGz»‹!
1G
"
$456$75
+p<,5
4$ QÉu ÛQI}e Íìì Ì}
!
×
œL
#+&, & #+$$
*
3D
!
ϗ
#+#"% "% #+##$
$$6+$"
!!
ËÊ
#+,"# "
œL
"+&66 & #+&$&
6:M
ϗ
#+#(" $ #+##"
(#&+&,#
!!
ËÊ
"+,# "%
œL
"+*%# & #+6#
\:
ϗ
#+#* "6 #+##(
"$&+6
!!
ËÊ
"+(# ($
!!
ô
$
!!Z
#+#$
`IµQɖj
(
@+I>
&NCQR÷S
w–hTZ
$
#
7
e
"## >7
cìÀ
!
n Ñ¥
¦
%# ¢ôõöÖt,L›é¨ÓԄ¸™Ú
¾”§
(
ˆ


$
¢ôõö
#, ?
„¸™Ú¾
”§@wjËz+š“
!
ŽŸ^š
$
ßؙ”§þ
ÿ
!
ö\”§ä£

ÂsOÿ
(
¢ôõö
* ?!
„¸™
Ú¾”§z+‹Pªš“
!
qÔ¦4ë5VW
!

ö\š31Øz­¦4VW
!
@qö\Ç^´µ

Â
s¶{|^«
(
(!
38598
¢ô-.kGŒ`͘j„¢zÕ

š™Ùû
ÚØ
!
VWqÔ

„¸™

¢J@™

;³™|óÔx
y
!
Ÿ^š }qEÛ

ÔÇvpœ
*$" V$(+
!
`Ê`Í
˜j™óÌ<ªvrvé
!
Ýg»óÇ^é“õ
ö
!
‰Ê“«+jªhi†Ø
*$%+
(
&#,
!!
"
!
#
!
$
!
%
!
& "
"%
#
.P-
hTš
%# õö
#,* ?!$
#
7
cìÀ›é¨„¸™Ú¾”§
% 42/
hTš
%# õö
#,* ?!
"## >7
cìÀ›é¨„¸™Ú¾”§
% ZN
š@³ö% 3
šg»Ô
% DN
š@qö(
ˆ
!
›é¨ÓԄ¸™Ú¾”§
g»Ôxy]y†ØQϯ{
!
g»Ôy
ø’v™t‹˜jây×Ó
(
¢ôõö×`Íg
»ÔæOvÜP
!
`Ê`Íg»ÛÇvpúe 
}qÝÞ
*$&+
(
w%&z^
!
6¢ôõö,LJ
D
!
›é¨ÓԄ¬ `3–jh¢
(
MšÔ
ó铧jxꐭ‹®
!
Ž5쟗ԝóé“
œìeÔxy¦Øˆ0ì
*,+
!
w%&z^­‹®
±v6¢ôõö,LJDʖjh¢
!
NŸ Ž
g»Ô¦4؈
(
”sÇנѥ¦z^
!
¢ôõ
ö
* ?
›é¨„Ôxyþ7¦4gªVW
( (
c
4$ßØ ¢ôõö×VW›é¨ÓÔg»Ôx
y
!
N / ª Ì ¾ = M j  y õ ö % & ” •
êÍ
*$, V$6+
(
Dg4
Z˜j™órvéäδw/™0
1|})«i5_Mn
(
w%&z^
$Dg4
rv
Z
%# õö
" ?
F4¡ª×
!
x7>?ÀÄ
!
å]
õö
% ?
4
$# ?!Dg4
rvš“zÖª
( .3d
Z
# , ?
rvpª
!
ä΋g»ó̉
T
"
g
"
´w
rvé
!
’qÈ«4‹õUßxyMn
*$*+
(
N/
Z¢ôõöÀ
!
è

àeáâè´wŽéÖt˜je
Ötõö”•¬ß°
*$ V"#+
(
ÒË%&ÓÔ
!

ôõöÀ
Dg4 .3d
”mš(°ÀĐ^
«
*"$ V""+
(
N,×^_]¤šó8㎥¦õö
¢ô0ìeõö,LÖt´w˜jwnõöä
â×ÓÖtÊÉ
(
˜j„¸Ð±v]™t˜j\s×Ӑêü
5_4$
*"(+
!
Z¢ôõöÀ
!
˜S™ó„¸ÐÄ
}õrvpœ
!
v݄¸ÐÄ}
*"%+
!
`Ê=•‹
\vi°<¾v
!
´Éʙóôìó¢R§+
̉Íì@˜jØˆ
*"&+
(
„΢ôv݄
¸ÐÄ}
!
åê˘j„ö„¸Ð±vªªÚÇ
‹\s>3_¶
(
w%&z^
!
›é¨ÓÔZ
%# õöGHÀ
!
6õö,LJD„¸Ð˱v
>?ÄÕ
(
×3®ô]
!
„¸Ð˱vÀĐt
,
!
„¸Ð
8YO
×ÒgÀÄd[
!
Ø ¢ô›é¨
ÓԄ¸Ð
8
“”±„¸Ð
O
“”ª
!

ðU|
*",+
%&¬™
!
/šåõ
*"6+
%&êÍ
!
,
×]ìÖtõöGH`͐
(
ZªÌ¾¬~À
!
¢ôõö^_“”
3D
!Æ×
zÓ
*"*+
(
„¸Ð°\2Ó$wŽ½¾×=>

Q
æ

º†ØȬ4„.\sMne+}‘,
“”
!
Ò]…ð˜j\sVW](¦4†Øsq
4$
*"+
(
yõöxyVW\xyԙmç“õ
!
ŽÄ³VW
3D
!
*(#+
!
¤Ü›é¨ÓԐ
!LS!?
Z
¢ôõö
" ?
–zjËÇ^ –ÀĐ^«
(

Ñ,´H4
!
6iõö,LJD
!
ßؙ¦VW
0ìú5
!
/Üt,
!
VW‹{¥ßؙԵ
3D
!
!
Ý3 *
3D
!
\:
ÀÄ
!3D
!™mv&m0ì
,#,
"&
? ’
!
U|
$
¢ôõöÀ›é¨ÓԐ+}”m
ÄÕ
!

b.
#é“ i=•
(
²³
DHC8A)
ABC
*($+
×vW21ý
!
˜j„öË×v
".PD#!
ê¯h´°\Ýmˆm
!
ª¯hå™mvR˜
(
å™mvR˜×vËê¯hkó
b.
#
é“7^ iÞT
!
Dê¯h´yçNÝmˆm
(
¢ôõöÀ‡\„¸ÐÅsHI
"NT-
!
#
ªv‡
Ü
!
êÕÐ
"n#
±vpú
!
„ÕЛ-Æ×úœ
!
`
Í
6:M
h¢
*("+
úª‹yçNÝm
!
Ë=È01
‹\sxyæOv
(
w%&z^
!
›é¨ÓÔ
6:M
Ç^–jh¢
!
/¢ôõöÀÕæÓÔ\sÈ
v%&”•êÍ
*((+
(
¢ôõöGHÀ
!
›é¨ÓÔ
^_´yçNÝm-ÉÊ\sxy¥¦¢ô
VW
(
êËÊ.
!
“cÈM/§È¡¸GH¬±
°“c¼îÆ
*(%+
!
t¤ì¢€¶ÈM/IÈ
MxL¬o“cÖîÆ
*(&+
(
IÈM¢ô
¼î“«˜j‘}
!
G£:;}mî?jcÕA
¨
",(&(CB( %B$)+"%( W8CM+#
ŠÔ¨¥¦ë5‘
}
*(%+
(
LP›
"=(3-"C+( %+$G".+5+5I;?#
c+
uªÌhiZ€¶
$ ### 7
´µ
!
^Z~̘j
…߉Š‹“c
(
ZՀ¶ÈMc˜,
!
£ºèé
j
!
Œ+DÖs
!
åZ¨ãÀc˜
!
ð+Dsq
!
íp
D,F
(# &# ;7
*(,+
(
w%&Ÿ 
!
Z¢ô„.
À
!
›é¨ÓԐg»Ôxy

„¸™Ú¾”§

0
1õxy´w\sxyߦ4Öt0ìVW
!
Ø
 ›é¨ÓÔ¢ô„.]±PQϐ
(
¤Ü
!
Z
Հ¶“c›é¨m«šŽ¹@+ÈÝ
!
Kê¹@
ãëpRª¨ì}wã
!
Éʼ·¢ô„Øˆ
!
Êݎ<·g^£™„¢
(
›é¨ÓԓcûÕ
€¶Ç^ÖsŸ^]ì̤Ð-.¤i"t€

!
Ž^_“”¤Ðz¹#Ë®¥‰êï
%&
(
VW!>
!
*$+
íð(
!
îä
!
ïòR
!
|
+
(¢|˜j
* +^+

$
fÑÇ

! "##($ &(6 V&%*
*"+
Z
!
X
!
Q±º
+
›é{Ôw“có8îÓ
* +`+
Q:!"#
E
! $* "%#$ $ V%
*(+
Z
!
X
!
ñð 
+
›é“có8¬÷v˔
* +^+
Q:!"
#E
! "##$ ",#$ $6 V$
*%+
KÈ
!
ú4õ
+
˜j+}$784`
* +`+

$
©}ت
$ÇB†
!"##"
*&+
Qs+
!
ñ
!
š
!
DÅ­
+
˜j+}+“78}eEz
* +^+
:;
$
¢|A{ÇB†
! "###
*,+
Ê3É
+
+,$Gn78%&}~/Ez
* +^+
:;
$
“$Ø
"ÇB†
! "##6$ $ V6*
*6+
²¯5
!
3Jñ
!
òó>
!
|
+
>\ô9\ô¤iÕ愸™
Ú¾”§wŽÆא“”
* +`+
mn+,$Ó
! "##(! $%"*#$
$"*6 V$"#
**+ .A8M85+.A;ICO8HBGBCIe9M8AB)8?@MCIKB> GBCIe9MBA;8LB>K9>KB>)
f@7B9> G:8>HA* +`+3?@A9I:3:8>H! $"! *&$ "(& V"%$
*+
~ªç
!
²\-
!
²]ô
+
˜j\sMn\cò
* +`+
˜j+
}$kl
! $"! "*"%#$ "(6 V"%(
*$#+ 58HB^ ! Z9:BAE +`-?:ICIG?@:<:JICBA;B>;B)8GC8;H9;8:KJ9MB* +`+
I`JC>8:I<2eGBC97B>H8:PIH8>@! "###! &$"(%&#$ ,& V,,*
*$$+ 0JO8> .n! TICHI> 3+0BKJ:8H9I> I<>I>)G?IHI;?B79;8:UJB>;?9>K
I<;?:ICIG?@:<:JICBA;B>;B9> G:8>HA* +`+.JAHC8:98> I`JC>8:IH
3?@A9I:IK@! $&! ""$ ""$ V"(#
*$"+ -987GICIL8^ ! 9^AHC9[ 1+R?BJ:HC8AHCJ;HJC8:CBAGI>ABIHIAHCBAAM9H9I>A* +`+2>L9CI>7B>H8:8>M 2eGBC97B>H8:PIH8)
>@! $(! (($ $$ V",
*$(+ -I:9>AZZ! E9BdN! D8?LB9H^ 2+TB8HA?I;[ GCIHB9>A8>M ;?9:)
9>KAB>A9H9L9H@I<7J>KOB8> ?@GI;IH@:A* +`+`IJC>8:I<2eGBC97B>H8:
PIH8>@! $&! %,$ 6& V*#"
*$%+ -?98ND! ;^08B4Z! R?I7GAI> 2`+N9K?H)MBGB>MB>;BIC8UJ8H)9>9H98HBM 7B7OC8>BMBHBC9IC8H9I> 9> OB8> G:8>HA+2L9MB>;B
LI:LB7B>HIH! $*"! &,$
%" V%
*$&+ N9> -]! -?B> ] !^ 5B@` N+DI:JHB:B8[8KB9> AI@OB8> ABBM:9>KA
J>MBCL8C9IJA?B8HA?I;[ CBK97BA* +`+3:8>H-B:3?@A9I:IK@! $*&!
","*#$ $%( V$%*
*$,+
Ô
!
õ
!
ù¼ï
!
úK«
+
©dyõö„ög»Ú¾”§%
&
* +`+
˜j$Ó
! $%! (,"#$ 6(# V6("
*$6+
ö5K
!
D´²
!
ú–œ
+
Dþâyv‘’}~%&
* +`+

="$Ó
! "##$! $%""#$ ," V,&
*$*+ .>> -! 8`;In! TBC78> -+R?BCBMIeAH8HJAIH;B:A".A.
8>M ZDT# 9AAB>A9H9LBHIf9>;97GIABM Ie9M8H9LBAHCBAA9> CIIHA8>M
GC978C@:B8LBAI<:*(%$"C/%L/C3(#+%* +`+3:8>H3?@A9I:IK@8>M P9I)
;?B79AHC@! "##$! ($ ,&6 V,,%
*$+
÷);
!
Õ9Ž
!
úà³
!
|
+
¢ôõöáâèÓԄö1
O+}=m“”
* +`+
u:=!#Eª$$Ó
$
ÍÎ#$B
!
"##%! (""$"#$ (( V(6
*"#+
«øs
!
)
!
Q
!
•´©
!
|
+
¢ôõöy›ùà„ö7
é“xye„¸ÐwŽ°\½¾“”
* +`+
(="#$
!
"##%! (6"*#$ $"%& V$"&#
*"$+
Ž>9
!
²ªú
!
ûT+
¢ôõöüÔ1O+}4$“
”w
.P.
e
, VP.
Žú
* +`+
?ý="$Ó
! "##&! "$
"(#$ $%& V$%
*""+
aÅÁ
!
úvT
!
²
!
Q
!
|
+
¢ôõöþs-²xrÓe
„ö+“4$“”
* +`+
©:=$Ó
! "##"! $6 "$#$ $"6
V$($
*"(+ W9:B[B>AT! -87G Wn! I^>H8KJ ^n!$&(C+DJ:M
J:HC8L9I:BH)P?8LBA979:8CB 70E.8;;J7J:8H9I> I<8>H9Ie9)
M8>HKB>BA9> 6+)"&+(-( BC/?G(3+-+A"C+( N* +`+3:8>H3?@A9I:IK@!
$%! $#,$ $##6 V$#$%
*"%+
ÿ™>
!
ÕËö
!
Õ7—
!
|
+
Ötâ¹v˜jZ¹õöÀ
+D/+}Èv±P%&
* +`+
>?ª$$Ó
$
="/+M#
6#,
!!
"
!
#
!
$
!
%
!
& "
"%
#
$B
! "##,! (""%#$ %"# V%"6
*"&+ 9^;?B:T! /:ICB>;BR+NIAAI<;?:ICIG?@:S9H? :979HBM CBMJ;H9I> I<
G?IHIA@>H?BA9A8A8> 8M8GH9LBCBAGI>ABI O8C:B@:8>MC8;BAHI
?9K?):9K?H8>M ?B8HAHCBAA* +`+.JAHC8:98> I`JC>8:I<3:8>H3?@A9I:I)
K@! $! ",$ &, V&6*
*",+
ðU
!
Ÿ!š
+
¢ôõöèeàg»ÔÇve\s¢Ð
“”
* +`+
¥"˜j$%&
! $6! $&"(#$ "(( V"(6
*"6+
šåõ
!
Õ
!
#
!
Ž\¥
+
$%õöÀ&¥„ö+}™m
ú
* +`+
!"#$
! "##6! %("$"#$ ( V%%
*"*+ D8>H8C9JA5.+D9HBAIA9H9L9H@9> ;?:ICIG:8AHA8>M M9)
H98:9>8;H9L8H9I> I<;@;:9;8>M >I>;@;:9;G?IHIG?IAG?IC@:8H9I> O@
?B8H9>K* +`+` IJC>8:I*"+ 5I;?BL85! N87OCBLG! ZBICK9BLZ!$&(C@2L8:J8H9I> I<;?:ICI)
G?@:<:JICBA;B>;B8>M 7B7OC8>B9>QJC@9> H?B:B8LBAIH9L8CAJ>MBCIA7IH9;AHCBAA* +`+P9IB:B;HCI;?B79AHC@! "##%! ,($
$"$ V$"%
*(#+
ú
!
1
!
¢-ñ
!
Õ:
!
|
+
þi\xy!yõöÈv
* +`+
mn+,$Ó
! "##6! $*"$#$ ,( V,*
*($+ DHC8AABC0 !` RA979:)^9;?8B:^ ! DC9L8AH8L8.+R?B<:JICBA;B>;B
HC8>A9B>H8A8HII:HI;?8C8;HBC9AB8>M A;CBB> G?IHIA@>H?BH9;A87G:BA
* +^YY+]J>JA^ !38H?CB_8>M I^?8>H@3+3CIO9>K3?IHIA@>H?B)
A9A$ B^;?8>9A7! 0BKJ:8H9I> 8>M .M8GH8H9I>+NI>MI>$ R8@:IC8>M
/C8>;9A! "###
*("+ _>:89R>K! d98IK8>KWB>! b9>KH8INJ!$&(C+TB8HDHCBAA1>MJ;BA
8> .KKCBK8H9I> IK-I7G:BeI<3?IHIA@AHB7
!
9> DG9>8;? 3:8>HA* +`+3:8>H3?@A9I:IK@! "##6! $%($ ," V,(*
*((+
QKK
!
G(Ó
!
()©
+
¢ôõöÀÕæÓԐ1O\sÈ
ve
3D
!\“$rvš“
* +`+
˜j+}$kl
! "##6! %(
",#$ $#*& V$#**
*(%+

+
(E¨c“c9~
* +^+
:;
$
#$ÇB†
! $*(
*(&+
ú *
+
!fæÞ{c
* +^+
:;
$
(!"ÇB†
! "##$
*(,+
„Jª
+
(fÚ#¨c
* +^
:;
$
(="ÇB†
! $,$
6& V6*
*#,