免费文献传递   相关文献

Mathematic Classification of 46 Species in Rhododendron with the Morphologic Characters

46种杜鹃花属植物表型性状的数量分类


选择44个在杜鹃分类中的重要性状用SPSS 10.0 进行分析,其中二元性状18个、多元性状13个、数量性状13个。Q聚类采用平均欧氏距离平方系数,R聚类采用相关系数距离,聚类方法采用类间平均链锁法。Q聚类结果将46种杜鹃花属植物分为特征明显的2大类群,即有鳞类杜鹃和无鳞的常绿类杜鹃,在2大类群下的聚类结果与传统的形态分类基本吻合,但美容杜鹃和汶川星毛杜鹃的亚组归属值得商榷。R聚类结果表明性状被分为明显不同的组,鳞片出现部位、花器构造、外部器官尺寸等性状群内有较强的相关关系。主成分分析中,前3个主成分累积贡献率达60.39%,表明杜鹃分类中存在重要性状,主要是有无鳞片、有无腺体和毛被以及叶、花、果的大小等; 对分类贡献不大的性状主要有雄蕊数、萼片是否具毛、萼裂片长短、花冠颜色、花丝基部毛、花冠毛等。主成分分析结果与Q聚类结果基本一致,揭示在杜鹃分类时应选择重要性状,避免过多次要性状的干扰。

The mathematic classification of 46 species in Rhododendron was studied with 44 morphologic characters, including 18 dualistic characters, 13 multi-characters and 13 quantitative characters, by using SPSS( 10.0. The squared Euclidean distance coefficient was used in case clustering and the Pearson correlation was used in variable clustering by within-groups linkage. The 46 species of Rhododendron were divided into two caboodles by case clustering: one was lepidote and the other was evergreen. The different groups were formed under two caboodles and the groups were similar to the traditional classification except R.calophytum and R.asterochnoum. The results of variable clustering showed that the various characters were decided in different groups. Some characters had strong correlativity within a group, such as the position of squama, flower formation, the size of external organs. In principal component analyses (PCA), the accumulative contribution of the first three principal components was up to 60.9%, which showed there were some representative characters which could be used in classification of Rhododendron, such as squama, gland, hair and the size of leaves, flowers and fruits. However, the number of stamen, hair on sepal, size of sepal, color of crown, hair on lower silk and hair on crown etc were not so reliable for the classification. The results of PCA were consistent with that in case clustering, which suggested that we should pay more attention to choose characters in classifying.


全 文 :! !"" ! ##
$ % % &$ # %
& ’ ( )
’()*+,)- ’)./-* ’)+)(-*
/012!"*+02#
-345*
!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!
$ % % &
!6mU’pV_‘#s<{1/K?r"
HCy7 auD7 1 >7 EF)$
A72 TSZ’T)&)F TU 6$"%7!S$2 TSEI~G&’> I~ 67"7%%Y
! "# gh !!AZU’?rB1Lo<{x ’8’’ 7%2% |}?@*^ BrA<{ 7#A5hA<{ 79A5/K<
{ 79A~:/rwx<.*1DE!"r !6mU’pV_‘?u$%$H1 $Tr_*8)"ik91z7?r+|)i*Aa:U’z*S.LU’1)&gV_XP+~;/r!"#$<{g?u$
Hno1&*<]]W@G5p;’~5n@;?#,VL(d 6%29&<*#$U’?rBNZLo<{*2o()&<]5)&-BzLge~k5p5"1T-
=S{?rVLnT1<{2o)7U/5.](<(L5.w]E$5pd$T5pO+@L5pdL=~2*?
?@!"i :/r!"+|au*-LZU’?rk=ghLo<{*TU6h[o<{1R6~
$%&% U’pVS/K?rS2*??@S/r?@
’()*+#’=7#2!& -./01#- -23+#7%%7 > =!##4$%%&5%# > %%6= > %&
vwx#%$%%# > %& > $$~
"QS~u%&e’~
!"#$%&"#’( )*"++’,’("#’-. -, /0 12%(’%+ ’. !"#$#$%&$’#& 3’#$ #$% !-42$-*-5’( )$"4"(#%4+
?@03 .ABCDB47 EAB4 F0B4GDB47 ?@AB4 .D7 H3 ?IJDB4$
A7 5 !"##$%$ "& ’"($)*(+*,-./012 3%(-.0#*0(1# 42-5$()-*+ 61G12 6$"%7!S$5 70-#- !"02*+ ’"($)*(+ 80($10*,-./012 9("5-2.$ 70-#- 67"7%%Y
67+#4"(#% ,@I JAK@IJAKDL L1AMMDNDLAKD0B 0N !6 MOILDIM DB :/";";$2;("2 PAM MK3QDIQ PDK@ !! J0RO@0104DL L@ARALKIRM*DBL13QDB4
7# Q3A1DMKDL L@ARALKIRM*79 J31KDSL@ARALKIRM ABQ 79 G3ABKDKAKDTI L@ARALKIRM*UC 3MDB4 ’8’’A 7%2% 5 ,@I MG3ARIQ *3L1DQIAB
QDMKABLI L0INNDLDIBK PAM 3MIQ DB LAMI L13MKIRDB4 ABQ K@I 8IARM0B L0RRI1AKD0B PAM 3MIQ DB TARDAU1I L13MKIRDB4 UC PDK@DBS4R03OM
1DBVA4I 5 ,@I !6 MOILDIM 0N :/";";$2;("2 PIRI QDTDQIQ DBK0 KP0 LAU00Q1IM UC LAMI L13MKIRDB4%0BI PAM 1IODQ0KI ABQ K@I 0K@IR PAM
ITIR4RIIB5 ,@I QDNNIRIBK 4R03OM PIRI N0RJIQ 3BQIR KP0 LAU00Q1IM ABQ K@I 4R03OM PIRI MDJD1AR K0 K@I KRAQDKD0BA1 L1AMMDNDLAKD0B
IWLIOK : 5 .1#"QDNNIRIBK 4R03OM5 ’0JI L@ARALKIRM @AQ MKR0B4 L0RRI1AKDTDKC PDK@DB A 4R03O*M3L@ AM K@I O0MDKD0B 0N MG3AJA*N10PIR N0RJAKD0B*K@I
MDXI 0N IWKIRBA1 0R4ABM5 )B ORDBLDOA1 L0JO0BIBK ABA1CMIMA8(-Y*K@I ALL3J31AKDTI L0BKRDU3KD0B 0N K@I NDRMK K@RII ORDBLDOA1
L0JO0BIBKM PAM 3O K0 6%2&<*P@DL@ M@0PIQ K@IRI PIRI M0JI RIORIMIBKAKDTI L@ARALKIRM P@DL@ L031Q UI 3MIQ DB L1AMMDNDLAKD0B 0N
:/";";$2;("2*M3L@ AM MG3AJA*41ABQ*@ADR ABQ K@I MDXI 0N 1IATIM*N10PIRM ABQ NR3DKM 5 H0PITIR*K@I B3JUIR 0N MKAJIB*@ADR 0B
MIOA1*MDXI 0N MIOA1*L010R 0N LR0PB*@ADR 0B 10PIR MD1V ABQ @ADR 0B LR0PB IKL PIRI B0K M0 RI1DAU1I N0R K@I L1AMMDNDLAKD0B5 ,@I
RIM31KM 0N 8(- PIRI L0BMDMKIBK PDK@ K@AK DB LAMI L13MKIRDB4*P@DL@ M344IMKIQ K@AK PI M@031Q OAC J0RI AKKIBKD0B K0 L@00MI
L@ARALKIRM DB L1AMMDNCDB4 5
8%9 3-4:+% :/";";$2;("2SJAK@IJAKDL L1AMMDNDLAKD0BSORDBLDOA1 L0JO0BIBK ABA1CMIMA8(-YSL13MKIR ABA1CMDM
U’pVA:/";";$2;("2Y_‘h()3j1.
#w_*(7FM0.#p/SJBx+1}<*
?4\Y/055>ABAyC5=*$%%%YSU’u
8:d*f1g0*{;8Qt6789()Loe
x~BC6Y(7FST1U’?1B7zhGB
7*m6U’lmTU*()4j1\:oxw_*
A/,U’pV_‘mrh5?1Y5J:T*?r
w$\?vw~k91z7?r&&|}D<"
#*uYDK?@*8)aD12.T3*Z89t
?zF&mr1gVdE#NZ?1*qrXU’
pV_‘1\:ox~
k91z7DEiUVLqr!i*z*X/
K?r)~/K?r*6‘?r)q‘aJD<"
#sDK15s,1jU<:]~C$s*/K?r
)(g=x,Dm?rAzq=*$%%=Y5(V8m
=?rA 2I:=*$%%$S-3t=*$%%=S^45
=*$%%"Y589)DEA2I:=*$%%!S*Q3ARQ0 $*
1# 5*$%%$Y=~K{Z/K?r+cd12*??
@(am6x2o7819U?@ab*(4eJ
hAn‘B-L6zW!12oa+*W(=x,
67)AH!C*!""#S9bP=*!""#Y5&’A:7
L=*!"""SHX;=*!"""Syg5=*$%%%Sz<
=*!""#Y=rj1DE**uDE6‘)~^7l
)(1Lo;(~
)uU’pV_‘/K?r1DE=AZCM
nv^@JA~|DE{TS6YQn &’ mm6
U’pV_‘1#s<{|}89/r?@z2*
??@*CZ,DLo?r+;*-L2o<{,=
1MZc8~i<{GG)1\-jv*u|ab
DEm=]lu8~|V_‘1\:oxxx()
=0~
$ 4,iab
!"! &’NO
|DE1 &’mU’B ($mJTS6@Qnw
0*TU! "!" ) ( &(* +*N $*mJBC()F56)
jQ_‘^qa>#FU’^w+~C ,-./+.0
A$%&%S$%1"Y?r89*?V (A)V*[@# $~
= ! %=&’=
!
!!!
#$%& ! #’( )$*(+,$- ./ (01(+,)(2*
C2
234
m0
,5.67.8
w0n
936:-7;<
C2
234
m0
,5.67.8
w0n
!! 936:-7;<
T}U’)V ,/=>.?/8 !"#$%&%’($) ypU’& ,/=8.6;73?!! *&#+&%,-&’&
T}U’& ,.6;73? ./%’01,# !* ypU’ 2 @ 1&#+&%,-&’,# nD!! A:?>B7?>
R.U’)& ,/=8.6;73? 3/4’,%$& N"U’)& ,/=8.6;73?!! *&#+"-/1&4+&
$ -"U’ 2 @ 5&60500 ?) C7?>D7?> !’ 9@U’ 2 @!! )/,-0$0
! a:U’ 2 @ 1&-/+("’,# ?) C7?>D7?> U’)V ,/=>.?/8!! 2(/5/5$%54/%
( T9U’ 2 @ 5$1/4,# A9 E7:??7?> U’& ,.6;73?!! 2(/5/5$%54/%
& * VBQU’ 2 @ 9/%99&)(&%$%)$ Dgn E:.0F:?> !# QKU’ 2 @ /4$/’4$+($) #F!! 93?>6G7
’ R.U’ 2 @ 8/4’,%$0 Dgn E:.0F:?> !1 hD7?>
# jkU’ 2 @ /4:01,-&4$ #F 93?>6G7 !% CkU’ 2 @ 5&605)/%0&%,# A9!! E7:??7?>
1 QPU’ 2 @ /4$/5/;& #F 93?>6G7 (" +LU’ 2 @ 40905,# A9!! E7:??7?>
% *S.LU’ 2 @ &)’$4/1(%/,# a4 H:3I7?> ($ UkU’ 2 @ ’&’)0$%$%)$ bD!! JG:3D/.
QkU’)& ,/=8.6;73? <49"4/+("--& (! ELU’ 2 @ ’401(&%’(,# 6E!! K76G:?>
$" QkU’ 2 @ &49"4/+("--,# ?) C7?>D7?> (( RYU’ 2 @ ",%%&%$%)$ 6E!! K76G:?>
$$ FaU’ 2 @ (,%%$=$--0&%,# #F 93?>6G7 (& *TU’ 2 @ 1/%10%%,# I~!! L/7-7
$! HpU’ 2 @ 8-/40:,%5,# #F 93?>6G7 (* LGU’ 2 @ &,9,)’0%00 I~!! L/7-7
T~U’)& ,/=8.6;73? 7&-0$%)0& (’ xpU’ 2 @ &#$)0&$ a4!! H:3I7?>
$( "oU’ 2 @ :,4$&600 I~ L/7-7 (# "kU’ 2 @ )05$4/+("--,# I~!! L/7-7
$& THU’ 2 @ :&-&%9$%)$ #F 93?>6G7 (1 6G7
$* eWU’ 2 @ =0-’/%00 #F 93?>6G7 (% w-U’ 2 @ &#:09,,# #F!! 93?>6G7
$’ PkU’ 2 @ +(&$/1(4"),# #F 93?>6G7 &" 2pU’ 2 @ -,’$)1$%) #F!! 93?>6G7
RLU’)& ,/=8.6;73? 3&-1/%$4& <$# 92kU’ 2 @ 9&-&1’0%,# a4 H:3I7?> &$ oOU’ 2 @ 4,:090%/),# I~!! L/7-7
$1 TQU’ 2 @ 4$; I~ L/7-7 jQU’)& ,/=8.6;73?!! >&++/%01&
‘pU’)& ,/=8.6;73? ?&1,-08$4& &! IUU’ 2 @ 0%’401&’,# bD!! JG:3D/.
$% g:U’ 2 @ )’4090--/),# a4 H:3I7?> &( ^njU’ 2 @ ’("#08/-0,# Dgn!! E:.0F:?>
!" E6G7 && P6G7
!$ JLU’ 2 @ +&1("’401(,# #F 93?>6G7 KkU’)V ,/=>.?/8!! .)$,5/4(/5/4&)’4,#
XLU’)& ,/=8.6;73? A44/4&’& MpU’& ,.6;73?!! 2(/5/:/’4")
!! S6U’ 2 @ )0B&%9$%)$ ?) C7?>D7?> &* MpU’ 2 @ 4&1$#/),# bD!! JG:3D/.
‘oU’)& ,/=8.6;73? C$4008-/4& KkU’& ,.6;73?!! 74&1("4(/50/%
!( hoU’ 2 @ )+$4&:0-$ #F 93?>6G7 &’ :v;]p 2 @ ($#0’401(/’,# 6E!! K76G:?>
2yU’)& ,/=8.6;73? !!>&%&’&
!& "LU’ 2 @ 10410%%&’,# I~ L/7-7
!"3 xy
$4!4$ l"#$%$& rw0M1 &’mU’e
u &’A?rBV=GAMNOY~%6{U’1Nn
9:5;|1.IUK**+)uU’mi<{1k
]#$~)6{k]#$|}?@~~*(g]Z
U’p?rBTx1 &&A#s<{|}CN*^ B
rA<{ $1A*CN "*$ShA<{ $(A*CN "*
$*!OSeB/K<{ $$ A*UD_7U/_8u
CNSf^/K<{ !A*?f(kQt{Ezp
yt{E*^ BkQt{EukQEik]E,j*
1’ & ’ ( ) &*"
pyt{EupyEipdE,j~a[ZO*k
]UE*kQ[UE*pyEzpdE[+|^qt
u*xt{EG4TUUV6FBkEzdE1r
[qr~i<{CN[@# !~
= ! 12=j.831=
!
!!!
"#$% ! "&’ ()*’ )+ ,)-.&)/)01’2
Z2
"#$
<{
%#&’(#)#*+
CN
,#-.
Z2
"#$
<{
%#&’(#)#*+
CN
!! ,#-.
/ fj 0.1*(2 3 / 4 5*/ 6 7 4 /*8 7 4 ! !9 pdw]/
!
!!!
":4;.& #< ’.2=)>
! zfL 0=1& #? 2.?-.& ;&=?@( & "#?. 5*) AB1>2 / !C .]E D.’=) ).?*2(
nH 5*/ 6 9 44 /*8 9
!! 44 !
9 zf- E)=?- #? 2.?-.& ;&=?@( & "#?. 5*) AB1>2 / !7 .L 0=1& #? >.’=) & "#?. 5*)!! AB1>2 /
C zf< DF:=4= #? 2.?-.& ;&=?@( & "#?. 5*) AB1>2 / !G .- E)=?- #? >.’=) & "#?. 5*)
!
!!!
AB1>2 /
7 k;n H.B2:&. #< ).=<
v; I=’.&+ 5*P; J.=2(.&+ /*’
P; H(1@K ).=2(.&+ ! !L
.< DF:=4= #? >.’=) & "#?. 5*)
!!
AB1>2 /
G kE J.=< ).?*2( !M pdE!! ,&#N? ).?*2(
L kZ J.=< N1-2( !O pdL 0=1& #? @&#N? & "#?. 5*)!! AB1>2 /
M kd< P;Q.&>. >F:=4= & "#?. 5*k R )122). /*g S.?>. ! 95 pd- E)=?- #? @&#N? & "#?. 5*)!! AB1>2 /
O kdL P;Q.&>. (=1& & "#?. 5*l R )122). /*g S.?>. ! 9/ pd< DF:=4= #? @&#N? & "#?. 5*)!! AB1>2 /
/5 kv< T?Q.&>. >F:=4= & "#?. 5*k R )122). /*g S.?>. ! 9! 7U/
!
!!!
":4;.& #< >2=4.?
// kvL T?Q.&>. (=1&
& "#?. 5*l R )122). /*/Z / )=+.&
!*!Z ! )=+.& 9 99
7U<.E
!
!!!
D2=4.? ).?*2(
/! kvLs D2+). #< 1?Q.&>. (=1&
& "#?. 5*"L U#)) /*RL ,:’
!*?f V&=?@( 9 9C
pO+L 0=1& ;.)#N >2=4.?
& "#?. 5*k R )122). /*g
!
!!!
S.?>. !
/9
kQt{E J.=<>2=)K ).?*2(W
J.=< ).?*2( 97
2MP/ ":4;.& #< *.&4.?
!
!!!
&##4
/C pZGH I#>121#? #< =?2(#2=B+
D6 R’1@=) 5*DM6 %1B /*M6
RB1))=&+ ! 9G
2ML 0=1& #? *.&4.? & "#?. 5*)
!!
AB1>2 /
/7 -p/ ":4;.& #< <)#N.&).2 9L 2M- E)=?- #? *.&4.? & "#?. 5*)!! AB1>2 /
/G pZrE J.?*2( #< &=@(1> 9M 2M< DF:=4= #? *.&4.? & "#?. 5*)
!
!!!
AB1>2 /
/L
py t { E I.-1@.) ).?*2(W
,&#N? ).?*2( 9O
2ME
!!
E.&4.? ).?*2(
/M pyL 0=1& #? ’.-1@.) & "#?. 5*) AB1>2 / C5 pPL 0=1& #? ’1>21) & "#?. 5*)!! AB1>2 /
/O py- E)=?- #? ’.-1@.) & "#?. 5*) AB1>2 / C/ pP- E)=?- #? ’1>21) & "#?. 5*)
!
!!!
AB1>2 /
!5 py< DF:=4= #? ’.-1@.) & "#?. 5*) AB1>2 / C! pPE I1>21) ).?*2( 3
pd ,&#N? 5* X pd
,&#N? /*8pd
!
!!!
,&#N? !
!/ pdz D2+). #< @&#N? PQz Y:??.) 5*yz V.))Z)1K. / C9 "z D2+). #< @=’>:).
e D2&=1*(2 5*lN D)1*(2
@:&Q1?* /*N
!
!!!
,:&Q1?* !
!! pT ,#)#:& #< @&#N?
R H1?*. 5*o U.- /*x I:&’). !*
2 [.))#N 9 CC
"E ,=’>:). ).?*2(
/ $! $! \6l"mgh kE5Z5kQEJs&
;|dKLR+ 7 6 !5]kUD$UV<._Sp
dEZ5pyE52ME57UEs&;|dKLR
7Q-pUD$UV<._S"Es&;|dKL
R 7A"UD$UV<._*EG=G.u 44S7
U/z2M/uh[U/!"~sm_‘UD;|
/ 9Aed~
/\!\9 \]^_%o’pq /0Z|}BV,
R*e;+n;+BbA.,)*/OOOY{k]/0|
}e~*e;+B/0UVi PH] ,=1DE8
/~
/\!\C ()pq x DIDD /5\5 <#& N1?-#N>wx<
.*1DEs<{thFGSxtoG2wxtu8/e U/
r*?@<{=1tuFG~! m/r.wxr=
<.vOb~u0/i6FB!!_ z!"_ _1S
J*(?f,D ^/rz U/r1!i{~
/\!\7 *+//> UVi<{=tu8/Xt
u8/JKSu0*?x+!",D2*?SeH
)&=A!55LY1abUV$%sKz2*?sKS
e2*? /sKuT:;*2*? !sKu8:;
e4M|~
! !"i?@
!34 3*$%)<
!\/\/ ^() CGmU’C*1-MT|}/i*/i6F~DE@# 9~J# 9
4@*ST1SJ]WZ/i1! C9b*^ [(!
OG! M# X&;=%CGmU’pV_‘#s<{1/K?r
= ! "3*^V|!!# 4=
$%&’ ! $() *)+%,-. /0 " 1-2.+)3,45 %4*!!
!
!!!!
#
/ibi
!"#$
/ir %&’("#)
%&’("#)
*
%&’("#)
+
S-DE
,-.-/0-("1.2#
DEn
!!3
/ibi
!"#$
/ir %&’("#)
%&’("#)
*
%&’("#)
+
S-DE
,-.-/0-("1.2#
DEn
!!
!!
3
!!* 45 44 +6757 +4 +7 58 +96:+* :6:7:
!!+ 59 4: 56*4; :64:9 +< +: +5 5*6:;8 +6:47
!!5 58 59 <6*75 +6:+7 +; ** *< 5*6+7< :6+:7
!!4 54 57 ;649* *65*8 +7 *: ** 556*;7 *689+
!!< +9 5* ;68;7 :657; +8 5+ 4< 5<68*+ +6;4<
!!; 4+ 45 864:+ *6<5< +9 * < 5;675: :69*8
!!7 4< 4; *:6;:4 +6+:+ 5: *9 +: 5;68;+ :6*5+
!!8 +9 55 **689< *6+9* 5* *: ++ 586:84 *6+++
!!9 +7 5: *46+88 +6595 5+ *5 +4 5965*8 *6+54
!!*: +8 5; *<68*7 *6<+9 55 +7 5+ 4:65*< :6997
!!** *: *4 *;64<+ :6;5< 54 * 8 4*68<+ *6<57
!!*+ +9 54 *76*47 :6;9< 5< *: *9 49675; 76884
!!*5 *+ +* *7659< :6+48 5; +7 4+ <:6;78 :694+
!!*4 5+ 5< *86:*7 :6;++ 57 * 5 <5657: +6;9+
!!*< *; +< *967+< *67:8 58 *: *5 <46:+; :6;<;
!!*; +7 +9 +:6795 *6:;8 59 + *7 <7674; 567+:
!!*7 5 4 +*68;: *6:;7 4: + 9 ;*6<;8 568++
!!*8 *: *; ++6<4< :6;8< 4* * 7 ;;68*8 <6+<:
!!*9 < ; +56544 :6799 4+ + 9 7+6548 <6<5:
!!+: 58 4* +56<+9 :6*8< 45 + *: 8+6;45 *:6+9<
!!+* +7 +8 +;6<;* 56:5+ 44 * + 9:65++ 76;79
!!++ ** *+ +;6;4< :6:84 4< * +7 **76:7< +;67<5
+5 *7 *8 +869<* +65:;
5J/r|A| *Yd4@*!i{ =* r 4;mU’
t?u +ATr_*! *r_A>Yy +:m*|}XE
@)+;Am&*~
!i{ =+ r >r_1 +:m)*rS"r_g?u ?*@*%*A 4 @?%?@?(R
.U’)&1 7Am*@@?<) *m9@U’S%
@?|} *4AmSA@?/RLU’)&1 +Am
zR.U’)&B1a:U’z*S.LU’&
**.ucI*k3T*-p/h~
!i{ =5 r)<1 >r_t?u 1*B*2 5A-
&%1-&<)jQU’)&1 5 Am*^ _f5k
]5pd=.%T-SB-&/Amz<kv&L*pd@T*2M@ES2-&1 4mU’
yoM(zf5k8+5p.52M=$H)L~=5
r"?r_|abt?u 1*B*2 5A-&*>?1(
R.U’)&M1?BS"@r_<) *AmS"%
r_g =5 t?u 1*B*2 5A-&*^ B 2-&) 9
Am*?VQkU’)&5T~U’)&5ypU’
)&5XLU’)&S"A r_g?u + A-&~
s-&mrq@o*7$T}U’)VMm=#s
J:eG@T~
%6{*1T}U’)VM +;Am=<.DE 7+65*:*)+:mU’m=<.DE 5<6+;4*#$)#sdJ:t{@-S4;mU’<.*1u 9:6::*STd ++96958*DE8/n:ZT*}U
OU’pVMm=#s<{?B\?On~
+6*6+ C() *Y<{?r i<{Ctu8/
/TM-‘b/i*/i!"@<{/rm8A|
+Y~/| +4@*!i{r 44A?r<{t?u 4
r~?ry *+ A<{*^ B 7 A<{i<])u*
Gi@G)#Wp;$%S@r<)pd)Li< *A<{S%
r) ;A<{*E@iL)uSAr<{Sh*d +<
A*,4bt?u +A-r*! *-r2oi-B)
u*! +-r+|E(/K<{~
+Y<{=1tu< J C/rGX<{tu
8/JK4@*@?<{,=tuFG@j*^ Bp
.52M5zf5py)&<] 4<{=tu8/V
u **7$3&<{tr:6Skv<])&ic 4
<{tu8/u:6955 :*[[Wok]WS<]1
]Wifj5k5p52M5"=1EG^$H^t
:7 & ’ ( ) 4<"
| ! "#mU’/r|
$%&’! ()*+,-&,./ -0 "# !"#$#$%&$’#&
u*]{_.Z123ed*>?X)T}rU’;-Gi;<[$H@-S2MP/5
-p/ipd52M56yE=<{1tu8/Z
1245ed*>?Xp;’~)aDM@c8SOn*
k(>?;124ed*#$i;!4! 3# X&;=%"#mU’pV_‘#s<{1/K?r
| ! U’<{/r|
"#$%! &’()*+$*,- +. -+*/0+1+$2
)&L7<]7-B5pO+@)Li<5pdw]
/5pPt{E5)&eV‘57UE5p.w]E=
<{i^g<{tu8/[WEZ 3 456 e8*7
$3&<{1]W+|S)yk4O*Zmed?
r$A1t?BexnT~
!"! qS))<
7/r!"(#$@?<{=tu8/4j*
NZ\-LTS@?<{1]W+|S)yk4
!8 & ’ ( ) 69"
O*Zmedr_1t?BS)$Hex*_O)n
oe2*??@~%62*??@*4e]kLT
z?r}>nT1<{**Lo<{bH]s~
!"!"# *+/,$ *?x+?@A# $Y({ $$A
?@$%A.U<{Y*iUV$*C^Z?rB1e
xzrfLft]1$%*?i~1$%_zVL
(~J# $ 4@*! # 2*?GGd\-K]d
$%"&&&’*R (A*?G\-6K1 )%"(*(’*Z^o
$#A*?Gxx1\-Ki (*")%+’*7$ZU’?
rDEBNZLo<{*)noe2*??@~
= ! qS)\])<=
"#$% ! &’#()*+* ,- .,/0,’1’2*
2*?]Z
,-./0.123
04514/6/7
F]$%_ 8/.7.23 6.96/:23;6<
$%_
=.96/:23;6<
VL(
>2-.2/06?’
#UVL(
@;5;327.:6?’
# #+"A$$ $%"&&& $%"&&&
! &")*! #!"*(+ &("$*#
( ("%(+ )"*%! )%"(*(
!"!"! l"m*+/-. rF]_2qXJK
z<{Rx8/#*i?s*XMx,U’?r1
$$A$%<{{=12*?qXK56qXKz<
{Lo<]Z*[@# &~
J# &4R*! # 2*?Zzf<5kv<5p
y<5.<52M< &A<{d.^W4j1^sq
XK*]{_.Z %"*ed*Zkd(<)KS>*u B %"A%+~i<])&t>*! # 2*?
ikE5kZ5pdE52ME5"E=<{.#W@
j1qsqXK*E@26 %"+&~3&<{1\-
qX4j*#$! #2*?2o>?_f(e~k5p5"52M=;<1n@s,*[7$3&
<{{U’?r4Lo~qXK1?Bok[$H
>?]! #2*?r $)mU’?u !Tr_%@T
s1&pd5p.5k](<()-B7Lg=<{dqX
K@j*7$3&<{{U’?r@uLo~
= 3 m.8_qS))<’^56H=
"#$% 3 45#6#.216 (,#7+’8* ,’ 251 06+’.+0#( .,/
!
!
!
!
!
!
!
0,’1’2* #’#()*+*
<{
@C2-2076-<
$%qXK
@C2-2076- 342D./9
6qXK
E4723 342D./9
Lo<]Z
F.9/.G.02/06
<{
@C2-2076-<
$%qXK
@C2-2076- 342D./9
6qXK
E4723 342D./9
Lo<]Z
!! F.9/.G.02/06
zf< FH;252 4/ 76/D6- I-2/0C %"*)! # "z!! F7J36 4G 021<;36 %"+#! !(
py< FH;252 4/ 16D.063 %"*)! ! 7U/!! K;5I6- 4G <7256/ %"+%A !$
pdz F7J36 4G 0-4L/ %"*)! ( pyL!! M2.- 4/ 16D.063 %")*) !&
.< FH;252 4/ 16D.063 %"*)! $ pPE!! ,.<7.3 36/97C %")A) !)
2M< FH;252 4/ 96-56/ %"*)! & zf-!! N32/D 4/ 76/D6- I-2/0C %")++ !+
-p/ K;5I6- 4G G34L6-367 %"AAA ) py-!! N32/D 4/ 16D.063 %")+! !A
"E @21<;36 36/97C %"AA( + zfL!! M2.- 4/ 76/D6- I-2/0C %"))+ !*
kE O62G 36/97C %"AA! A fj!! M6.9C7 %")(+ (%
2ME N6-56/ 36/97C %"A+& * pPL!! M2.- 4/ 1.<7.3 %")(# (#
kv< PI:6-<6 pdE @-4L/ 36/97C %"A$& ## pdw]/!! K;5I6- 4G 16723< %")%( ((
2MP/ K;5I6- 4G 96-56/ -445 %"A($ #! kdL!! PI:6-<6 C2.- %"&*A ($
kvLs F7J36 4G ./:6-<6 C2.- %"A(% #( Qkj!! O62G <723Q?O62G 36/97C %"&A# (&
.- N32/D 4/ <6123 %"A%A #$ 7UE!! F7256/ 36/97C %"&)A ()
pP- N32/D 4/ 1.<7.3 %"A%# #& pd2M- N32/D 4/ 96-56/ %"+*% #) ydj!! ,6D.063?@-4L/ 36/97C %"&$* (A
kvL 8/:6-<6 C2.- %"++& #+ pd-!! N32/D 4/ 0-4L/ %"&## (*
pZGH ,4<.7.4/ 4G 2/7C472RJ %"++( #A .L!! M2.- 4/ <6123 %"$)) $%
kZ O62G L.D7C %"+&$ #* .w]!! F6123 %"$&+ $#
kd< PI:6-<6 pZ6rE O6/97C 4G -20C.< %"+!& !# pO+L!! M2.- I634L <7256/ %"(!% $(
k;n E6R7;-6 4G 362G %"+#( !! pdL M2.- 4/ 0-4L/ %"!&* $$
J<{]ZA# &Yd4@*{U’?rSLo
1n#(<(<7-B7Lg*^ [(k5p52M5
"1T-*^ 6qXK.Z %"Aed*LoG)<{1R #&GSLo<]$ #%G1?f(k
Qt{E57UE5pd<5pyt{E5pd-5p
.L5.w]E5pdT5pO+L5pdL*^ B.
](<(L5.w]E$5pd$T5pO+@L5p
dLZR ( A2*?16qXKBLoH*6qXKqZ %"&e8*{?rVLS-~
!"!"( *+//6%D0 e! #*! 2*?sK
(+! A# X&;=%$)mU’pV_‘#s<{1/K?r
| ! 2*??@4M|
"#$%! &’( )*+,,(- .#+$-+/ 01 23
C2i# 4o~!T}U’)V*"U’)V*#KkU’)V~&’( 56/7(-) +-( #.(5,#*+8 9#,’ ,+78( 4 %
!:67$(56) !"#$%&%’($)*":67$(56) *(+,+,$%,-+%*#:67$(56) .)$/,+-(+,+-&)’-/#
?fuT:;z8:;e4M|A| 0Y~J| 0d
4@*12mU’g$H?u 3 ATr*r7E()
U’r_B*Sr71(KkU’)V1 3Am*y
iU’)Vj@BC*AH^iY~ix 3A2*
?]5j@)+ab4}*!"4\~
! d’
!"# 7C*12^UY
:8(6/(-A4;<;YrpZM61)u 4AKkU’)V*Ah/)’2^r^tIU
’)VU’&Be)&e~A3’+/7(-8+#5 $’ &4 %*
4;;=S3688(5*4;>?S2’#8#@)05 $’ &4 %*4;>AY~E#e
s*KkrU’1gVae(Q’12M,a~J
#s<{df*KkU’)ViU’)V,=,&
p)&,=<.DEu <>CB=*8’j@*fn])
V=1n:~Aa&DE!"#$*ZF&a+ A
)V=,NZ$Hn:~jv$=AA???YwxU<
G:#QFG?@*HLKkU’)VM1 !mU’
?kthS4
2ZA4;;=Y%6{KkU’)V1 =Amke!$
%.I*x]c)V_‘()Q;Z:d5nZ\M
-5MZ\MT5k2auX[;p&=$%*n4
:8(6/(-A4;<;Yr^t?u 4A)V1?r89Sp:
F(\"!"[HL A)V)aDn:AX&;=*
A??>Y~_O*KkrU’?rnG1i~,D*v!
o{YhmrzZnorjeY"I1DE~
!"$ 8R|DS$X
|DEB*<’)&1<.DEu A>C;D>*Z=<.DEu A=C=,3 AA)&z7n:|#4-*)&t?F{n
~W*_O4eDEiHu 4A)&~
Zk91z7?rB*a:U’z*S.LU
’gt?ZT}U’)V8R.U’)&B*AZ
|DE E/rB*c)&1N BmU’Z@-DE
M/iZa?*a:U’z*S.LU’#i,t
D@\*SgiRLU’)&1TQU’z92k
U’/iu 4r~C?rN]#*R.U’)&1
2o$%(zf&L*k)-B7JLSk@-*E
nM A? */S2M)-BA4$8=*4;;mU’io&1^g BmU’)$Htf%zf)
LSk<.E26 A! */*T126 !? */S2Mz
pP&-B*Ge(1(<=gIR.U’)&_
XP+~
!"! )*.8^ST
!/rz2*??@!"q-L?r<{H%
UhU(*a@?<{17UE5pd)&eV‘5
pyE5p.L5pd$T5pO+@)Li<5pd
w]/5pPE5)&eV‘=<{ZV8?r=x
dex4-*ZU’?rk=ghLo<{*Yhn
r})&<]5)&Lg~rs5k5p52M5"1
T-=*TU6h[o<{{?r1R6~
J K - .
jv$*^E#*QBr " #$$$ " %mU’pV_‘1?k?BD
E " Y6_‘*#$A&Y%’(( ) ’*#"
4$8*a$^*u?P*= " +%%& " BC_‘"%! ,(" #?3 " Q
R%()]DE*# ) ’"
H)&*‘Ba " #$$( " -.-- 2*??@B1$%sKUVw$ "
9UbK*A’Y%+$ ) ++"
H!C " #$$( " LK<_0t67)ptZ’zZJ)p{(vw "
U‘Z’()*’,A#/Y%*’%$ ) *’%#"
HX;*QrG " #$$$ " a6bVm=wm 0+ i]|p"/K<{
1?kJ:?@ " &’()DE*+’A’Y%#%$ ) #%& "
-3t*o i*HK4*= " #$$( " BCPV_‘1/K?rD
E " V4_‘)DE*#,A#Y%+&% ) +,("
9bP*ct$ " #$$( " d_67891*iv9‘AWsDE " B
C67Z’)J*+,A’Y%+’* ) +&+"
:7L*eC’*^B0*= " #$$$ " Qfk/K<{1?k?@ "
&’()*’/A,Y%&( ) ,’"
42Z " +%%/ " KkU’)Vke!$%i&1t? " RYg"T
))J*+/A&Y%/# ) /’"
.,) " +%%% " 6‘/) " QR%()]DE*,+ ) ($"
2I:*z = " #$$# " BC2‘V_‘/K?r1DE " 6‘/)
)J*+(A’Y%’*$ ) ’*’"
2I:*‘=6*Qg%*= " #$$& " BC@?ntm2_‘t8]
lu81DE " V4_‘)DE*##A’Y%##/ ) #’$"
^45*G-h*.t>*= " #$$, " M2z7;#/r?@DE "
M2DEsB*#&A+Y%#, ) #*"
z q*ifo*^ u/ " #$$( " jsDm1/K?rDE " ^d)
J*’&A&Y%%&( ) %,&"
z <*ko9 " #$$( " DQE&’*i6l5pDE " &’)pw
$*#(A’Y%#(# ) #(,"
yC5*ua; " #$$$ " U’p(_‘}<*?~exLxDE|
] " V4_‘)DE*+*A/Y%,$% ) ,+&"
yg5*4bl " +%%% " 2*?/rZ‘atm2^l-B1Bx "
GK&)F)J*+%A+Y%’’ ) ’/"
X&;*QS~*^ i " #$$* " #/mU’pV_‘p:z7~?
r)DE " &’()*&&A#Y%,, ) /’"
1234567839: ; 0* <=34 !* >7?6:@ A* !" #$ " +%%/ " B26 ?6:CD
%&’(’(!)(*’)* 9@D E83DD9F9E3@9G: 3:H D=:G:=4=" IH9:5C7?2%!G=38
JG@3:9E A37H6: IH9:5C7?2"
1C886: K" +%*$ " > 76L9D9G: GF %&’(’(!)(*’) M" -C5?6:CD %&’(’(!)(*’)
-6E@9G: %&’(’(!)(*’) 3:H +’,’)#)"&-. " NG@6D !G=38 JG@3:9E A37H6:
IH9:5C7?2*’%A+Y%+ ) #$* "
IHC37HG M* JG75 >*A67G7?6 K* !" #$ " #$$# " 08G738 3:H L6?6@3@9L6
4G7O2G46@79ED GF F9L6 +$!-*’"&#$$/0AP7E29H3E636YDO6E96D%1G77683@9G:
Q9@2 @3RG:G4=*O2=8G?6:=*?6:6@9E L37935989@= 3:H OG889:3@9G: D=D@64D"
>::38D GF JG@3:=*%$%#+% ) #’$ "
.2989ODG: S N*.2989ODG: T !" +%*#" > O768949:37= D=:GOD9D GF @26 ?6:CD
%&’(’(!)(*’) " NG@6D !G=38 JG@3:9E A37H6: IH9:5C7?2*&$A+Y%##, )
##("
-86C467 <"+%&% " I9: D=D@64 H67?3@@C:? %&’(’(!)(*’) M" JG@ K3275 -=D@*
(&%,++ ) ,,’"
-86C467 <"+%*$ " .3D@ 3:H O76D6:@ @3RG:G49E D=D@64D GF %&’(’(!)(*’) 53D6H
G: 43E7G4G7O2G8G?9E38 E2373E@67DUUMC@6=: K M*PGJ796: S I"
1G:@795C@9G:D @GQ37H 3 E83DD9F9E3@9G: GF %&’(’(!)(*’) " N6Q VG7W%N6Q
VG7W JG@3:9E38 A37H6:*+% ) #/"
!"#$% QRS)
,(! *# X&;=%&/mU’pV_‘#s<{1/K?r