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Effects of Provenances on Growth Characteristics of Sapwood and Heartwood of Larix gmelinii Trees

兴安落叶松边材心材生长特征的种源效应*


以生长在相同环境条件下的7个代表不同气候地理条件(7个种源: 塔河、满归、根河、新林、三站、乌伊岭、鹤北)的26年生兴安落叶松为对象,比较不同种源树木的边材/心材生长和异速生长关系,探索种源地气候条件对边材/心材生长特征的影响。结果表明: 胸径(DBH)、边材宽度(SW)、边材面积(SA)、心材半径(HR)、平均边材生长速率(MSGR)等生长特征参数的种源间差异显著,其中最南种源地鹤北的生长特征参数的平均值最大,三站的平均值最小。种源也显著地影响SA、心材面积(HA)与MSGR的关系,但对其生长特征参数与DBH的异速生长关系和心边材比率(HSR)没有显著影响。SW,HR,HSR和MSGR分别与DBH呈极显著的线性关系,而SA和HA与DBH则呈极显著的幂函数关系。除了三站种源之外,种源地的年均温和年降水量都显著地影响兴安落叶松的SA和MSGR。兴安落叶松边材/心材生长特征因长期适应不同种源地的温度和降水等环境条件而发生了显著变化,这种变化主要是通过生长速率差异而不是改变其生长格局实现的。

Response and adaptation of tree growth characteristics of sapwood and heartwood to climate changes are essential to exploring mechanisms of tree water relation and carbon metabolism, developing tree growth yield models, effectively using timber, etc. In this study, we compared the growth of sapwood and heartwood and allometric relationships of 26yearold Larix gmelinii trees growing under same environmental conditions but originated from seven provenances (Tahe, Mangui, Genhe, Xinlin, Sanzhan, Wuyiling, Heibei), representing significantly different geographical and climatic conditions, to examine effects of climatic conditions of origins on the growth characteristics of sapwood and heartwood. There were significant differences in such growth parameters as diameter at breast height (DBH), sapwood width (SW), sapwood area (SA), heartwood radius (HR) and mean sapwood growth rate (MSGR) among the provenances. The mean values of the parameters from the most southern provenance: Heibei were the greatest, whereas those from Sanzhan were the least. Provenances also significantly influenced the relationships between SA, heartwood area (HA) and MSGR, but did not affect the allomteric relationships of the parameters against DBH, and the ratio of heartwood area to sapwood area (HSR). The SW, HR, HSR and SGR were significantly correlated to DBH; and SA and HA were related to DBH as a power function. The mean annual air temperature and precipitation of the origins significantly affected the SA and MSGR of the larch trees except for the Sanzhan provenance. We concluded that significant changes in the growth characteristics of sapwood and heartwood of L. gmelinii due to the longterm adaptation to changes in temperature and precipitation of the origin, which were presented in terms of changes in growth rates instead of growth patterns.


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