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Effects of Deferred Grazing on Grassland Community Characteristics and Productivity in Alpine Meadow

春季延迟放牧对高寒草甸草地群落特征及生产力的影响



全 文 :!23" !5#
 Vol.23  No.5
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犱狅犻:10.11733/j.issn.10070435.2015.05.025
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correspondence,Email:hualm@gsau.edu.cn
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:S812    >?@AB:A     >CD=:10070435(2015)05106805
犈犳犳犲犮狋狊狅犳犇犲犳犲狉狉犲犱犌狉犪狕犻狀犵狅狀犌狉犪狊狊犾犪狀犱犆狅犿犿狌狀犻狋狔犆犺犪狉犪犮狋犲狉犻狊狋犻犮狊
犪狀犱犘狉狅犱狌犮狋犻狏犻狋狔犻狀犃犾狆犻狀犲犕犲犪犱狅狑
WANGQiaoling,HUALimin,WANGGuizhen,LIULi
(ColegeofGrasslandScience,GansuAgriculturalUniversityKeyLaboratoryofGrasslandEcosystemoftheMinistryof
EducationCentersforGrazingLandEcosystemSustainability,Lanzhou,GansuProvince730070,China)
犃犫狊狋狉犪犮狋:Winterspringpastureisseverelydegradedrangelandbecauseofgrazingforlongtimeandinthe
stageofreturninggreen.Theexperimentsofdeferredgrazingwith4paddocksandanintervalof10days
betweenpaddockswereconductedinalpinemeadow.Theplantheight,coverage,biomassaswelasthe
numberoffloweringandseedsettingweremeasuredandrecorded.Theresultsshowedthattherewasno
significantdifferenceintheplantheight,coverage,biomassandtheimportantvalueofdominantspeciesa
mongfourpaddocksatthebeginning.However,theplantheights,coverage,biomassofaltestedpad
docksincreasedwiththegrazingtimedeferred.Theplantheight,coverageandbiomassofthemostdef
erredgrazingareawere12.07cm,97.67%,348.98g·m-2,respectively.Thenumberoffloweringinthe
mostdeferredgrazingpaddockwassignificantlylowerthanthatinotherthreepaddocks(犘<0.05),
whereasthetotalnumberofseedsettingwassignificantlyhigherthanthatinothers(犘<0.05).Theim
portantvalueof犘狅犪狊狆.and犈犾狔犿狌狊狀狌狋犪狀狊increased,whereastheimportantvalueof犔犲狔犿狌狊狊犲犮犪犾犻狀狌狊
and犅狉狅犿狌狊犻狀犲狉犿犻狊decreasedinaltestedpaddocks.Theimportantvalueof犎犲狋犲狉狅狆犪狆狆狌狊犪犾狋犪犻犮狌狊and
犕犲犱犻犮犪犵狅狉狌狋犺犲狀犻犮犪increasedin1st,2ndand3rdpaddocks,whereasdecreasedin4thpaddock.Thesere
sultssuggestthatthestrategyofdeferredspringgrazinginwinterspringpastureisagoodmanagement
measureforcontrolingtherangelanddegradationofalpinemeadow.
犓犲狔狑狅狉犱狊:AlpineMeadow;Deferredspringgrazing;Grasslandproductivity;Communitycharacteristics
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Timeofleaving
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Numberoffinesheep
N–/Â
Areaofgrassland/hm2
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Fig.2 Theeffectsofdeferredspringgrazing
onthecoverageofvegetation
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Table2 Theeffectsofdeferredspringgrazingontheimportantvalueofmajorplantspeciesofvegetation
^X
Treatment
CDõImportantvalue
&$É
犘狅犪狊狆.
]^LMN
犈犾狔犿狌狊狀狌狋犪狀狊
/N
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Vc5d
犅狉狅犿狌狊犻狀犲狉犿犻狊
jkGHIo
犎犲狋犲狉狅狆犪狆狆狌狊
犪犾狋犪犻犮狌狊
TUh
犕犲犱犻犮犪犵狅狉狌狋犺犲狀犻犮犪
5% 0.659a 0.757a 0.532a 0.579a 0.439a 0.291a
1Ý 6% 0.646b 0.717a 0.527a 0.517b 0.516a 0.615a
Plotone 7% 0.705b 0.646a 0.324a 0.303b 0.211a 0.400a
8% 0.897a 0.817ab 0.308a 0.367bc 0.588a 0.546a
5% 0.749a 0.670a 0.500a 0.639a 0.371a 0.376a
2Ý 6% 0.655b 0.594b 0.408b 0.691a 0.600a 0.446ab
Plottwo 7% 0.703b 0.637a 0.182b 0.332b 0.253a 0.545a
8% 0.845a 0.835a 0.174b 0.463ab 0.572a 0.480a
5% 0.749a 0.733a 0.452a 0.674a 0.379a 0.263a
3Ý 6% 0.660b 0.575b 0.336b 0.636ab 0.504a 0.498ab
Plotthree 7% 0.728b 0.695a 0.267a 0.485a 0.229a 0.502a
8% 0.910a 0.814ab 0.276a 0.522a 0.512a 0.476a
5% 0.801a 0.604a 0.479a 0.611a 0.359a 0.325a
4Ý 6% 0.896a 0.504b 0.291b 0.169c 0.239b 0.374b
Plotfour 7% 0.911a 0.689a 0.258a 0.180c 0.218a 0.437a
8% 0.969a 0.692b 0.183b 0.275c 0.262b 0.285b
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[1] ;Ɗ,;¼?,û6L,!.J¤ìÞìæÝNñ:FŽA
À,<˜™
[J].Q}@Ë,2005,27(5):633640
[2] ª™ó,1¨,^ ¼Ê,!.J¤ìÞ¬­:|àáû8
ðæ
[J].ñX+w,2012,67(1):312
[3] oRJ,yZz,Yî°,!.J¤ìÞNñ,<ÞEs¥[J].
N)°+
,2007,24(5):2026
[4] ;ðB,HG¾,—.®,!.J¤ìÞìæNß,<©ÀBz
ÝRS
[J].N)+w,2012,22(2):110
[5] Öúú,Ó_½,$:,!.J¤ìÞìæNß,<8».’M
‡/+89f—BSˆížKy
[J].:+w,2013,33
(15):46834691
[6] ’Z|,H2:,uF®.ìæNßO,)|ü8@kXW8

[J].´†°+,2000,22(4):5061
[7] ͍0,uS‚,’Z|,!.J¤ìÞìæNñB,<†Ò’
M
[J].N)8O,,2008,11:0109
[8] ‘›,_ÔÎ.3¤ìÞNñ,<†ÒáEz{[J].+¬KË
ûü°+
,2008,6(1):112116
[9] ButtolphLP,CppockDL.Influenceofdeferredgrazingon
vegetationdynamicsandlivestockproductivityinanAndean
pastoralsystem[J].JournalofAppliedEcology,2004,41(4):
664674
[10]NieZN,ZolingerRP.Impactofdeferredgrazingandfertiliz
eronplantpopulationdensity,groundcoverandsoilmoisture
ofnativepasturesinsteephilcountryofsouthernAustralia
[J].GrassandForageScience,2012,67(2):231242
[11]IsmailSS,LarsOE,OysteinH,犲狋犪犾.Theeffectsofadef
erredgrazingsystemonrangelandvegetationinanorthwest
ern,semiaridregionofTanzania[J].AfricanJournalof
Range&ForageScience,2013,30(3):141148
[12]™u.N)°+˜™NÓ[M].4¥:+¬()¡¢,1998
[13]o„®.NÞ:O,)ìi:ú•–[M].78,78°+
•–¡¢
,2013
[14]$„Æ,h»3,’£L,!.­î»,KNñ8 B67[J].
Nñ+w
,2008,16(3):166286
[15]ÖJ$,’·,1œ,!.­î»,KNÞ=­O:úçB6
7
[J].Nñ+w,2005,13:5366
[16]c›A,Heo,$;,!.îàg»,KWXò,›¨šN
ñ89f—RSB67
[J].Zb()°+,2013,50(6):
11451149
[17]’·,ÖJ$,HÛH,!.­î»,KÞR=Nñ:úgâB
67
[J].÷ã()•–+0+w,2006,19(1):17
[18]Y%S.Þî»,Kc¯N–8 B67[J].+¬Nñ,1999
(6):6869
[19];J<,ɬÊ,’²,!.NßNÞWXò,›¨KË̍
›ÆÌëB67
[J].N)°+,2007,24(10):610
[20]MuhammadSH,AngusM,AmjadAC,犲狋犪犾.Effectofgraz
ingsystemsonrangeconditioninPabbiHilsReserveForest,
Kharian,Punjab,Pakistan[J].ChileanJournalofAgricultur
alResearch,2011,71(4):560565
[21]²J„,}ì£,¬S.ò,›¨KJ¤ìÞ¬¤ìæNß
89f—RSB67
[J].N)+w,2009,18(5):256261
[22];Lø,H_J,¶´´,!.WXò,›¨šìæNß8 R
S=ËÌâzn<˜™
[J].Nñ+w,2010,18(4):511516

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