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Alpine Meadow Succession and Plant Species Diversity Characteristics under Different Grazing Gradients

不同牧压梯度下高寒草甸演替及植物物种多样性分异特征



全 文 :!23" !5#
 Vol.23  No.5
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ACTA AGRESTIA SINICA
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犱狅犻:10.11733/j.issn.10070435.2015.05.006
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犃犾狆犻狀犲犕犲犪犱狅狑犛狌犮犮犲狊狊犻狅狀犪狀犱犘犾犪狀狋犛狆犲犮犻犲狊犇犻狏犲狉狊犻狋狔犆犺犪狉犪犮狋犲狉犻狊狋犻犮狊
狌狀犱犲狉犇犻犳犳犲狉犲狀狋犌狉犪狕犻狀犵犌狉犪犱犻犲狀狋狊
OUYANGJingzheng1,2,LIYikang1,2,LINLi1,DUYangong1,
ZHANGFawei1,GUOXiaowei1,LIQian1,LIUShuli1,2,SUNYanan1,2,
KEXun1,2,CAOGuangmin1
(1.NorthwestInstituteofPlateauBiology,ChineseAcademyofSciences,Xining,QinghaiProvince810008,China;
2.UniversityofChineseAcademyofSciences,Beijing100049,China)
犃犫狊狋狉犪犮狋:Acomparativestudyonfouradjacentplantcommunityspeciescharacteristicswithdifferentgraz
ingintensitieswasconductedinalpinemeadows.Thevariationsofplantspeciesdiversityatdifferentsuc
cessionalstageswerediscussedunderdifferentgrazinggradients,aimingtoprovidethebasisofvegetation
protectioninthisregion.TheresultsshowedthatShannonWienerindexchangedindistinctivelybetween
thestageof犓狅犫狉犲狊犻犪犺狌犿犻犾犻狊communityandmatticepipedoncrackingstageof犓.狆狔犵犿犪犲犪community,
however,itchangedsignificantlyamongtheotherthreestages.Simpsonindexdidnotsignificantlychange
betweenthematticepipedoncrackingstageof犓.狆狔犵犿犪犲犪communitywithGramineaeand犓.犺狌犿犻犾犻狊
communitystage,matticepipedonthickeningstageof犓.狆狔犵犿犪犲犪communityand犓.狆狔犵犿犪犲犪communi
ty,whereassignificantlychangedamongotherstages.TheevennessindicesbasedontheShannon-Wie
nerindexdidnotchangesignificantlybetween犓.狆狔犵犿犪犲犪communityandmatticepipedonthickening
stageof犓.狆狔犵犿犪犲犪community,whereasitchangedsignificantlyamongtheotherthreestages.Grazing
measuresignificantlychangedtheconstructivespeciesanddominantspeciesofmeadow,resultinginthe
changeofspeciesimportantvalue.Withtheincreasesofgrazingpressure,theplantcommunitywassignif
icantlychanged.ThecommunitysuccessionofthealpinemeadowwasfromGramineaeand犓.犺狌犿犻犾犻狊
communityto犓.狆狔犵犿犪犲犪community,matticepipedonthickeningstageof犓.狆狔犵犿犪犲犪communityto
matticepipedoncrackingstageof犓.狆狔犵犿犪犲犪community.TheimportantvalueofCyperaceaespeciesin
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creasedwiththesuccessionsofcommunitiesinthealpinemeadow,whereastheimportantvalueofGra
mineaespeciesdecreased.
犓犲狔狑狅狉犱狊:Grazingpressuregradient;Alpinemeadow;Succession;Speciesdiversity;
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Table1 Variationsofecologicaldominanceinalpinemeadowcommunity
Nñ©<ðà
Successionstage
wØNN…PæUC#
MC
wØNN…Pæ H#
MT
äØN“#
KH
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GK
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Note:significantdifferencebetweenmeanvalues(犘<0.05)
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[1] ;€¬,­5®.8 :+—f—8:AÀ[M].4¥:+
¬FŽ°+•–¡¢
,1997:512
[2] ;§¯,Y°.:9‹&gBXW8=¹[M].4¥:+¬F
Ž°+¡¢
,1994:167
[3] Ñìñ,±­ž,,!.J¤ìÞìæNñ:AÀ£¤ô
õ¥Þ
[J].Üñ+w,2003,21(1):5055
[4] NaeemS,LiS.Biodiversityenhancesecosystemreliability
[J].Nature,1997,390(4):507509
[5] TilmanD,DowningJA.Biodiversityandstabilityingrass
land[J].Nature,1994,367(27):363365
[6] Ö­z.ò,ÜûKìæNß89f—RSB67[J].p2
()°+
,2010,38(5):128132
[7] ²J„,}ì£,¬S.ò,›¨KJ¤ìÞ¬¤ìæNß
89f—RSB67
[J].N)+w,2009,18(5):256261
[8] ˆá‰,$Za,ÖH$,!.ìæäØNNß9Y‹&g8
â‹&gK‹€:úçB67
[J].:+NZ,2009,28(6):
9991005
[9] 1›,Ó|®,Ö_³,!.ò,›¨KFJ3©ìæNÞf—
9Y‹&g=:úçB67
[J].Nñ+w,2012,20(6):
10331038
[10];Lø,H_J,¶´´,!.WXò,›¨šìæNß8 R
S=ËÌâzn<˜™
[J].N)+w,2010,18(4):510516
[11]oÖþ,⵶,$w·,!.WXîàí¨ò,KìæNß
89f—RSB67
[J].3489+w,2014,34(2):349
357
[12];u¸,0gŽ,;‹ ,!.ò,ûQšìæNß89‹&
g

:úçKËÌâz‘n[J].:+w,2008,
28(9):41454151
[13]H›~,Ü>¹Î,ÖЛ,!.º,»,K¤4ìæNß9Y
‹&g=:9ëB67
[J].+¬()°•Nw,2013,15(3):
143149
[14]Íáâ,‹3ãä,åþþ,!.J¤ìÞìÜØNNß89‹
&g=ËÌâzKò,B7ž›L
[J].Nñ+w,2012,20
(6):10261032
[15]Ö}×,£d,Hӛ,!.wØNf—ìæNßñ&g8 
ò,dçšB>”tÃf—
[J].Üñ+w,2010,28(3):257
265
[16]Ö}×,£d,Hӛ,!.ØNNß,<=’M‡/+¾D,
N©À=ñPRSn<
[J].N)+w,2010,19(5):179185
[17]£d,Ö}×,Hӛ,!.ìæäØNf—,<©ÀAÄÇ、
r:<+Áë+RS
[J].:+w,2013,33(17):5245
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[18]—.®,ª«,Ú¬á,!.ìæØNNßB Q8¾Q,<
zoRS†Ò@:›X
[J].Üñ+w,2007,25(6):641648
[19]™u.N)°+˜™NÓ[M].4¥:+¬()¡¢,
1998:129
[20]Ö6›,ÍZÈ.1QNÞݼwñ89f—©À‡/+9Y
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[J].‚œc()*++w,2002,23(4):2631
[21]š£.ÀÁ:+[M].4¥:+¬£)¡¢,2009:6173
[22]¨w©,ª½£,HG¾,!.¬ý7ÜWX,<ðàìæNß
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[23]YŽ,¶‹G,=à%,!.@a¿c¯£f—l%RS˜™
[J].£)°+˜™,2010,23(1):9397
[24]À˜ç,ªŠ,¶w5,!.nËÁ!Ýc¯f—B89àá、
89‹&g†Ò8FŽE7B@A
[J].Nñ+w,2012,20
(5):609615
[25]Í+¨,֘S,ÍÂ,!.º,KãÄ1QNÞ9Y‹&
gB67
[J].:+NZ,2008,27(8):13071312
[26]ÍÂ,Hò,Í+¨,!.Simpson«p=ShannonWiener«p
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[27]=cƒ,Y].+¨ò,ÜûKNñ:9‹&g67BÑ>
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[J].õ()°+,2007,35(21):65676568
[28]ِÆ,Ç;°,;‹‡.XËQ”,m8kâ
[J].N)°+,1999,16(2):59
[29]—.®,0gŽ.ò,ìæØNNßBÇkg†®0Iü›L
[J].+¬()ð_,2009,30(4):553559
[30]£d,—.®,Ö}×,!.»yPQKJ¤ìÞìæäØNN
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[31]uF®,;‹‡,HÈJ,!.WXò,›¨šìæNß8 ©
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[32]ö# ,;‹‡,bñ­,!.n&†Ý».Nñf—RS、‹
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