Cloning and Analysis of BnMPK4, A Novel MAP Kinase Gene Induced by Oligochitosan in Brassica napus Differences in Root Microscopic Structure of Root Mutants lrn1, prl1 and Wild Type in Oilseed Rape (Brassica napus L.) Expression Analysis and eQTL Mapping of BnTT3 Gene in Brassica napus L.