免费文献传递   相关文献

Observations on Megasporogenesis, Microsporogenesis and Development of the Male and Female Gametophytes of Arundinaria simonii f. heterophylla

异叶苦竹大小孢子及雌雄配子体的发育


Embryological characters of Arundinaria simonii f. heterophylla were reported in this article. The anthers were tetrasporangiate in which the cytokinesis of microsporocyte in meiosis was successive type and the produced microspore tetrads were bilaterally symmetrical. The cells of tapetum layer with dense cytoplasm were secretory type, and they began to degenerate during meiosis division. The mature pollen was trinucleate. The single ovule with parietal placenta was anatropous, double integument and tenuinucellatae. The development of the embryo sac was polygonum type. Generally the florets close to the base of spikelets bloomed earlier than others, and development of both stamens and pistils within a spikelet was synchronized.


全 文 :第 !" 卷 第 # 期
$ % & ’ 年 # 月
林 业 科 学
()*+,-*. (*/0.+ (*,*).+
0123!"!,13#
.456! $ % & ’
718"&%6&&9%9:;6&%%&<9!##6$%&’%#$!
CFVF8ZF7 7OEF" $%&$ =&$ =$&# CFZ8GF7 7OEF" $%&’ =%! =$A6
@14L7OE81L UT1;FVE" G4UU1TEF7 YP,OE81LO2,OE4TO2(V8FLVF@14L7OE81L 1S)N8LO%’&%%%$"!&6
!)1TTFGU1L78L5O4EN1T" g8L5R421L56
异叶苦竹大小孢子及雌雄配子体的发育!
林树燕B丁雨龙
%南京林业大学竹类研究所B南京 $&%%’9&
关键词" B小孢子发生# 大孢子发生# 异叶苦竹
中图分类号! (9!9BBB文献标识码! .BBB文章编号! &%%& =9!###$%&’$%# =%&># =%#
X<-(*V"+$1.-1.!(3"-)1*13(.(-$-! !$7*1-)1*13(.(-$-".@R(V(K1)%(.+14+6(
!"K(".@0(%"K(Q"%(+1)6>+(-14B$",2*,)$*) &*51,*43+#(#$16+/0)
/8L (N4POLBg8L5R421L5
%;.,L""0$1$.+39 4(1-*-2-$!:.(<*(% >"+$1-+76(*)$+1*-7B:.(<*(% $&%%’9&
;<-+*"7+" B+WYTP12158VO2VNOTOVEFTG1SN+2(/*(.+*. 1*,"(*S69$-$+"O97#. XFTFTFU1TEF7 8L EN8GOTE8V2F6-NFOLENFTG
XFTFEFETOGU1TOL58OEF8L XN8VN ENFVPE1‘8LFG8G1SW8VT1GU1T1VPEF8L WF81G8GXOGG4VVFGG8ZFEPUFOL7 ENFUT174VF7
W8VT1GU1TFEFETO7GXFTFY82OEFTO2PGPWWFET8VO26-NFVF2G1SEOUFE4W2OPFTX8EN 7FLGFVPE1U2OGWXFTFGFVTFE1TPEPUF! OL7
ENFPYF5OL E17F5FLFTOEF74T8L5WF81G8G78Z8G81L6-NFWOE4TFU12FL XOGET8L4V2FOEF6-NFG8L52F1Z42FX8EN UOT8FEO2
U2OVFLEOXOGOLOET1U14G! 714Y2F8LEF54WFLEOL7 EFL48L4VF2OEOF6-NF7FZF21UWFLE1SENFFWYTP1GOVXOGU12P51L4WEPUF6
fFLFTO2PENFS21TFEGV21GFE1ENFYOGF1SGU8‘F2FEGY211WF7 FOT28FTENOL 1ENFTG! OL7 7FZF21UWFLE1SY1EN GEOWFLGOL7 U8GE82G
X8EN8L OGU8‘F2FEXOGGPLVNT1L8\F76
=(> ?1*@-" BW8VT1GU1T15FLFG8G# WF5OGU1T15FLFG8G# N+2(/*(.+*. 1*,"(*S69$-$+"O97#.
B B -NF28SFN8GE1TP1SWOLP YOWY11GUFV8FG8G
VNOTOVEFT8\F7 YPOL 4L4G4O2S21XFT8L5NOY8E% 8^OL5!
$%%9# R8! &""9# KFL! &""’&6-NFP1SEFL S21XFTOL7
78FG8W42EOLF14G2P1ZFTX87FOTFOGOE21L58LEFTZO2G
% ’ =&$% PFOTG&! OL7 ENFL TF5FLFTOEFST1W GFF7 1T
TN8\1WF% O^L\FL! &"9># /8FGF! &"#A# /8$-.#=! $%%>#
-OP21T$-.#=! &"### &""’# .YF$-.#=! $%%&&6-NFTF
XFTFO2TFO7Pu48EF21EG1SU4Y28VOE81LG7FO28L5X8EN
YOWY11S21XFT8L5VPV2FGOL7 G1WFNPU1ENFGFGXFTF
58ZFL %/O8! $%%&# k8O$-.#=! &""A# g4 $-.#=! $%%%#
]V)24TF! &""’&6D1XFZFT! ENFTFXFTFSFXFTTFU1TEG
OY14EFWYTP1L8V7FZF21UWFLE1SYOWY11%J8O1$-.#=!
&"#!# D4 $-.#=! &""!# KOL5$-.#=! $%%># /8L $-.#=!
$%%"& 74FE1ENF21L5S21XFT8L58LEFTZO2G6
N+2(/*(.+*. 1*,"(*S=9$-$+"O97#. XOG8LET174VF7
ST1W O^UOL 8L &"#! OL7 5T1XL 8L ENFYOWY115OT7FL 1S
,OL;8L5 @1TFGETP iL8ZFTG8EP! )N8LO6 *ENOG YFFL
S21XFT8L5GU1TO78VO2PG8LVF$%%!! Y4ESFXGFF7GVOL
YFVOUE4TF76*L 1T7FTE17FEFTW8LFXNFENFTENF21X
GFF77FZF21UWFLE 1VV4TTF7 8L 78SFTFLE FWYTP1L8V
7FZF21UWFLEUFT817! ENFS21TO2W1TUN1215POL7 ENF
XN12FUT1VFGG1SW8VT1GU1T15FLFG8G! WF5OGU1T15FLFG8G
OL7 5OWFE1UNPEF 7FZF21UWFLE 1S N= 1*,"(* S=
9$-$+"O97#. XFTF7FGVT8YF7 OL7 V1WUOTF76
CD!"+(*$"K-".@%(+61@-
BB@8F27 1YGFTZOE81L OL7 V12FVE81L 1SENFWOEFT8O2
XFTFVOTT8F7 14E74T8L5S14TV1LGFV4E8ZFPFOTG%$%%"’
$%&$& OEENFG8EF1SYOWY115OT7FL 1S,OL;8L5@1TFGETP
iL8ZFTG8EP! 8^OL5G4 IT1Z8LVF! )N8LO6
-NF WOEFT8O2G 1S N+2(/*(.+*. 1*,"(* S6
9$-$+"O97#. ST1WENFP14L58LS21TFGVFLVFGOL7 S21XFTG
8L ZOT814GGEO5FGS1TFWYTP12158VO2GE47PXFTFV12FVEF7
FZFTP7OPYFEXFFL #"%%’&%"%% OW 74T8L5ENFUFT817
]OTVN $L7’.UT82$’T7 ST1W$%%" E1$%&$6@21XFTY47G
OL7 S21XFTGXFTFS8QF7 8L W178S8F7 @.. %S1TWO28Lm
OVFE8VOV87mO2V1N12bAm>m#"&6-NFW1TUN1215P1SENF
!第 " 期 林树燕等! 异叶苦竹大小孢子及雌雄配子体的发育
7-6\+93\’36K3+9S+, R6(5/(4643-],49/(058+7-6\+9/(0
3+’36( 796: A$$A 56A$$DTM8+7/;+, :’5+9/’-\’3
59+’5+, \/58 R6(S+(5/6(’-:+586,367,+8],9’5/6(#
59’(3<’9+(R]# ’(, +:K+,,+, /( <’9’7/(TM/334+3\+9+
3+R5/6(+,# K658 59’(3S+93+-]’(, -6(0/54,/(’-]# ’5" N
?$ ::58/Rh(+33\/58 ’J+/R’265’9]B+R5/6(+943/(0
35’(,’9, 5+R8(/i4+3T B+R5/6(3\+9+35’/(+, \/58 ’
3’79’(/().’35d9++( R6:K/(’5/6(TM8+3+R5/6(3\+9+
6K3+9S+, \/58 ’( j-]:<43PI)A :/R963R6<+ ’(,
<8656:/R9609’<83 \+9+ 5’h+( \/58 <865609’<8/R
7’R/-/5/+3’336R/’5+, \/58 58+:/R963R6<+T
./0T?!.-6\+9/(0R8’9’R5+93678%$*9(*’%(’ :(&;*(7"<#,#%;=<>-’
’TM8+7-6\+9/(0R-4:< 678":(&;*(7"<#,#%;=<>-’’ K# RTM8+-’5+9’-/(7-69+3R+(R+’(, 58+0+(4/(+3-+::’’(, 645)359+5R8/(035’:+(3’ , N8TM8+3594R549+676(+3’(, 6(+0](6+R/4:\/58 3869535]-+K+’9/(05\6K9’(R8+36735/0:’’ /TB++,367,/7+9+(53/g+\+9+5].69+;’:/(/(0 <6-+( 7+95/-/5]# 5\6 ,/7+9+(5
:+586,3\+9+43+,T./935# ’(58+93\+9+R6-+R5+, 796:
3M8+ <6-+( 7+95/-/5]\’3+35/:’5+, K]R64(5/(058+
(4:K+9367(69:’-’(, ’K(69:’-<6-+( 09’/(3/( 58+
7/+-,367:/R963R6<]<+93796:5+( 3’75+97-69+56<+(/(0# =’9(6]*336-45/6( 769AD 8# 59+’5+, \/58 % :6-+JN?
U’jI769?A 8# 58+( 35’/(+, \/58 $1?c ’(/-/(+K-4+T
M8+ :/R963R6<+’(, 58+(4:K+967<6-+( 09’/(3’5’R8+, 56
’(, 0+9:/(’5+, 6( 58+35/0:’\’3R64(5+,TM8+:+’(3
796:5+( 58+<96R+,49+’9+’S’/-’K-+/( I4 %?@@D&T
>"@-56:15)04)0):8535
>A$"%07:/2-5.-0.-B/2C,/:/98!M8+7-6\+9K4,367
8":(&;*(7"<#,#%;=<>-’ 675+( ,/7+9+(5/’5+/( &’9R8
’(, K-66: /( H<9/-# \8/R8 /3 3+:+-’4R5’(5
/(7-69+3R+(R+TB R:-6(0# \8/R8 /(3+95
+’R8 (6,+677-6\+9/(0K9’(R8+3# -/(+’9# 09++(# 7-’5#
0-’K9643’(, <+,/R+-/3A ND R: -6(0# \8/R8 675+(
R6(5’/( % N?$ 7-69+53# \/58 A +:<5]0-4:+3’5/53K’3+#
9’R8/-’ ,/3’95/R4-’5+,# D N " :: -6(0# 7-’5#
<4K+94-63+TJ+::’?D NA$ :: -6(0# <49<-+’(,
09++(TL’-+’?A N?# :: -6(0# \8/5+-6,/R4-+E#
35’:+( E# ]+-6\# > N?$ ::-6(0# 7/-’:+(537/-/769:#
?$ N?% ::-6(0TjS’9]-6(0+-/<5/R# A NE ::-6(0#
@%?
林 业 科 学 !" 卷B
GEP2FG8L52F! & =$ WW 21L5! GE85WOG’! XN8EF!
U24W1GF! ’ =! WW21L56)OTP1UG8G&$3" =$%3’ WW
%@856&O=8&6
ABAD01*%"+$1.14".+6(*?"KB-NFOLENFTG1SN=
1*,"(*S=9$-$+"O97#. OTFEFETOGU1TOL58OEFGET4VE4TF
OL7 XFTFENFEPU8VO21SW1L1V1EP2F71L14GGUFV8FG6.E
ENFFOT2P 7FZF21UWFLEO2GEO5F! O L4WYFTT1XG1S
OTVNFGU1T8O2VF2GYF5OL E178SFTFLE8OEFG4TT14L78L5YP
ENFOLENFTFU87FTW8G%@856$O&6*EXOGL1EF7 ENOE
L1TWO2W8E1G8G1VV4TTF7 8L OTVNFGU1T8O2VF2GE1S1TW
UT8WOTPUOT8FEO2VF2GOL7 UT8WOTPGU1T15FL14GVF2G
%@856$Y&6
-NFOLENFTXO2XOGWO8L2P7FT8ZF7 ST1W UOT8FEO2
VF2GV1LG8GE8L51S%ST1WENF14EFTE1ENF8LLFT&1LFT1X
1SFU87FTW8G! 1LFT1X1SFL71ENFV84W! 1LFT1X 1S
W8772F2OPFT! OL7 1LFT1X1SEOUFE4WENOEXOG8L ENF
S1TW1SGFVTFE1TPEOUFE4W %@856$F&6-NFFU87FTW8G
VOL YFTFEO8LF7 ENT145N14EENF5OWFE15FLFG8GUT1VFGG6
D1XFZFT! ENFW8772F2OPFTVF2G7F5FLFTOEF7 5TO74O2P
74T8L5ENFV14TGF1SENFS1TWOE81L 1SW8VT1GU1TFW1ENFT
VF2G! X8EN 1L2PTF28VGTFWO8L8L5YPENFE8WF1SENF
W8VT1GU1TFEFETO7G7FZF21UF76-NFEOUFE4WVF2GXFTF
VNOTOVEFT8\F7 YPENFUTFGFLVF1S1LF1TEX1L4V2F8OL7
ENF7FLGFVPE1U2OGW6-NFEOUFE4WTFOVNF7 8EGWOQ8WO2
G8\F74T8L5WF81G8G78Z8G81L OL7 ENFL 7F5FLFTOEF7 OL7
FZFL V1WU2FEF2P78G8LEF5TOEF7 4LE82ENFS1TWOE81L 1S
ET8L4V2FOEFU12FL 5TO8LG6KNFL WOE4TFU12FL 5TO8LG
7FZF21UF7! ENFOLENFTXO21L2PV1LG8GEF7 1SFU87FTW8G
OL7 FL71ENFV84W OL7 ENF FL71ENFV84W VF2 XO2
EN8V‘FLF7 TO78O2POL7 YFVOWFS8YT14G6
ABED!$7*1-)1*13(.(-$-".@%"K(3"%(+1)6>+(B*E
XOG1YGFTZF7 ENOE74T8L5S4TENFT7FZF21UWFLE! ENF
UT8WOTPGU1T15FL14GVF2GXFTF518L5 ENT145N ENF
W8E1G8G78Z8G81L E1S1TW GFV1L7OTPGU1T15FL14GVF2G
ST1WXN8VN ENFW8VT1GU1T1VPEFGXFTF7FT8ZF7%@856$V!
7! F&6-NFW8VT1GU1T1VPEFUOGGF7 ENT145N FOVN GEO5F
1S WF81G8G 78Z8G81L E1 S1TW W1GE2P Y82OEFTO2P
GPWWFET8VO2EFETO7G%@856$ S=1& OL7 ENFVPE1‘8LFLG8G
XOGG4VVFGG8ZF6*EXOG87FLE8S8F7 ENOEENFW8VT1GU1TFG
F8ENFT8L LF85NY14T8L5GU1TOL58O1TX8EN8L ENFGOWF
GU1TOL584W XFTF L1E GPLVNT1L8\F7 8L ENF8T
7FZF21UWFLE6 .SEFTENFGFUOTOE81L 1SEFETO7G! ENF
WOE4TFW8VT1GU1TFGXFTFS1TWF7 X8EN 7FLGFVPE1U2OGW
OL7 1LFUT1W8LFLEOL7 VFLETO2PU2OVF7 L4V2F4G%@856$
U! u&6.GENFW8VT1GU1TF7FZF21UF7 S4TENFT! O2OT5F
ZOV412FU42F7 ENFVPE1U2OGWOL7 ENFL4V2F4GE1U12FG
%@853$T! G&6 g4T8L5 ENF G4YGFu4FLEW8VT1GU1TF
78Z8G81L! EX14LFu4O2PG8\F7 L4V2F8XFTFUT174VF76
-NF2OT5FT1LFX1427 YFENFZF5FEOE8ZFL4V2F4GOL7 ENF
GWO2FTX1427 YFENFTFUT174VE8ZF1LF6-NF8WWOE4TF
U12FL 5TO8L XOG4L8L4V2FOEFOL7 8L ENFWOE4TFU12FL
5TO8L EX1 GUFTW L4V2F8 XFTF S1TWF7 ST1W ENF
TFUT174VE8ZF L4V2F4G YP 518L5 ENT145N O W8E1E8V
78Z8G81L %@856$E&6.GOTFG42EENOEENFWOE4TFU12FL
7FZF21UF7 8LE1 O ET8L4V2FOEF 5TO8L X8EN 1L2P 1LF
5FTW8LO2OUFTE4TF%@856$4&6
ABGD!(3"-)1*".3$&% ".@%(3"-)1*13(.(-$-B-NF
OTVNFGU1T8O2VF2 YF21X ENFL4VF24GFU87FTW8GXOG
78Z87F7 8LE1OUOT8FEO2VF2OL7 OGU1T15FL14GVF2YP
ENF UFT8V28LO2 78Z8G81L OL7 ENF GU1T15FL14G VF2
78SFTFLE8OEF7 8LE1OWF5OGU1T1VPEF%@856’O! Y&6-NF
WF5OGU1T1VPEFXOGG4TT14L7F7 YP1L2P1LFT1X 1S
L4VF24G VF2G OL7 ENFTFS1TF ENF 1Z42F XOG
EFL48L4VF2OEOF6-NFWF5OGU1T1VPEFXOGVNOTOVEFT8\F7
YPO2OT5FL4V2F4GOL7 7FLGFVPE1U2OGW6g4T8L5ENF
G4YGFu4FLE7FZF21UWFLEENFWF5OGU1T1VPEFF21L5OEF7
2FL5ENXOPGOL7 FLEFTF7 8LE1WF81G8G78Z8G81L %@853’
V=5& E178TFVE2PS1TW ENF7PO7 %@856’N & OL7 ENF
28LFOTEFETO7 1SWF5OGU1TFG % @856’8&6 [L2PENF
WF5OGU1TFOEENFVNO2O\O2FL7 XOGENFS4LVE81LO2
5OWFEF! XN82F1ENFTG5TO74O2P7F5FLFTOEF7 %@856’;&6
AB8D/%<*>1 -"7".@4(%"K(3"%(+1)6>+(B-NF
S4LVE81LO2WF5OGU1TFX8EN 7FLGF VPE1U2OGW OL7 O
UT1W8LFLEL4V2F4GS1TWF7 O$XN8VN 78Z87F7 TOU872P8L S4TENFT7FZF21UWFLEOL7 1LF
#O!FWYTP1GOVXOG1SENFD"#7%"(2,EPUF! 8L XN8VN ENTFF
L4V2F828F7 OEENFW8VT1UP2OTFL7! EX1LFOTENFVFLEFT!
OL7 ENTFFOEENFVNO2O\O2FL76[LFW8VT1UP2OTL4V2F4G
YFVOWFENFF55OL7 EX1YFVOWFENFGPLFT587G! XN8VN
S1TWF7 ENFF55OUUOTOE4G%@856’U&6-NFEX1VFLETO2
L4V2F8YFVOWFENFU12OTL4V2F8! XN8VN 2OEFTS4GF7
%@856’u&6-NFVNO2O\O2L4V2F8YFVOWFENTFFOLE8U17O2
VF2G%@856’1&! XN8VN UFTG8GEF7 4U E1ENFWOE4TF
FWYTP1GOVGEO5F6.SEFTENF#OLE8U17O2VF2G 78Z87F7 TOU872P E1 7FZF21U W1TF
OLE8U17O2VF2G%@856’T&6
ABND’6(@(V(K1)%(.+"K71**(-)1.@(.7(*(K"+$1.-6$)
<(+?((.-+"%(.-".@)$-+$K-B-NFS21XFT8L5E8WF1S
%9&
!第 " 期 林树燕等! 异叶苦竹大小孢子及雌雄配子体的发育
!!
./0TA!M8+:/R963<6960+(+3/3’(, ,+S+-6<:+(567:’-+0’:+56<8]5+678%$*9(*’%(’ :(&;*(7T<#,#%;=<>-’
’TM8+59’(3S+93+3+R5/6( 67]64(0’(58+9/( \8/R8 ’9R8+3<69/’-R+-3\+9+(65+’3]56K+764(, ’558/335’0+#58+645-’]+9/3+:/563/367’9R8+3<69/’-R+-%’9R& ’ RTL9/:’9]3<6960+(643R+-% <3& ’ ,TB+R6(,’9]3<6960+(643R+-%33& ’ +T&/R963<69+:658+9R+-%:/R& ’(, 7649
-’]+9367’(58+9\’-%+M+-6<8’3+67:+/63/3O%5+-O& ’ -TL9+:+/5/R<8’3+OO% <9+OO& ’ :T&+5’<8’3+67:+/63/3OO%:+5OO& ’ (TH(’<’3+67:+/63/3OO%’(’OO& ’ 6TM+59’,3
%5+5& ’ :/563/367:/R963<69+%:/R& ’ 5Te+0+5’5/S+(4R-+43%SR& ’(, 5\63<+9:(4R-+/%3(& ’ 4T&’549+<6-+(T
?>?
林 业 科 学 D@ 卷!
./0TE!M8+:+0’3<6960+(+3/3’(, ,+S+-6<:+(5677+:’-+0’:+56<8]5+678%$*9(*’%(’ :(&;*(7"<#,#%;=<>-’
’TH9R8+3<69/4:%’9& ’ K# RT&+0’3<69+:658+9R+-%::R& ’ , N0T&+/63/367:+0’3<69+:658+9R+-%::R& ’ 8TV]’,’
/# CTM+59’, 67:+0’3<69+’ hTM\6)(4R-+43+:K9]63’R%5(+3& ’ -TM8+:/563/367+:K9]6(4R-+43%:/5& ’ :T.649)(4R-+43
+:K9]63’R%7(+3& ’ (TF/085)(4R-+43+:K9]63’R%+:K3& ’ 6T&’549++:K9]63’R%:+:K3& ’ M\6<6-’9(4R-+% <(& ’ 9TH(5/<6,’-R+-3%’R&T
A>?
B第 # 期 林树燕等" 异叶苦竹大小孢子及雌雄配子体的发育
78SFTFLES21TFEG8L 1LFGU8‘F2FEXOG8LV1LG8GEFLEOL7
5FLFTO2PENFS21TFEGV21GFE1ENFYOGFY211WF7 FOT28FT
ENOL 1ENFTG6
@T1W -OY6&! 8EVOL YF GFFL ENOEW1GE1S
7FZF21UWFLEO2GEO5FGYFEXFFL GEOWFLGOL7 U8GE82GXFTF
GPLVNT1L8\F7! FQVFUES1TENFGEO5FXNFL ENF2FL5EN 1S
ENFGU8‘F2FEGXFTFOY14E$ =’ VWOL7 ENFOLENFTGXFTF
OY14E% =$ WW6KNFL ENF2FL5EN 1SOLENFT8G! =A
WW! Y1EN ENFU8GE82OL7 ENFGEOWFL XFTF8L WF81G8G
78Z8G81L6KNFL OLENFTWOE4T8L5! ENFFWYTP1GOVO2G1
EFL7F7 E1WOE4TF! XN8VN 8L78VOEF7 ENOEENFGU8‘F2FEG1S
N=1*,"(*S=9$-$+"O97#. YF21L5F7 E1O78VN15OWP6
’"<\CD’6(*(K"+$1.<(+?((.+6(K(.3+614-)$^(K(+".@+6(@(V(K1)%(.+14-+"%(.".@)$-+$K
/FL5EN 1S
GU8‘F2FEG:VW
/FL5EN 1S
OLENFT:WW
(EO5F1S
W8VT1GU1T15FLFG8G
(EO5F1S
WOVT1GU1T15FLFG8G
$ =’ % =$ (U1T15FL14GVF2 .TVNFGU1T8O2VF2
’ =’3A $ =’ ]8VT1GU1T1VPEF ]F5OGU1T1VPEF
’3A =!3A ! INOGFG1SWF81G8G gPO7! EFETO7
! =A A R14L5! STFFW8VT1GU1TF $A => > =9 ]OE4T8L5W8VT1GU1TFG !> =93A # =" ?8L4V2FOEF5TO8L #9 =#3A &% =&& -T8L4V2FOEF5TO8L ]OE4TF7 FWYTP1GOV
BB
AB] D P1K(. 4(*+$K$+> 14B$",2*,)$*) &*51,* 4+#(#$16+/0)B[ZFT9%c 1SENFU12FL 5TO8LG1SN=
1*,"(*S=9$-$+"O97#. XFTFL1TWO2OG8L78VOEF7 YP
GEO8L8L5X8EN e*<*$ OL7 OL828LFY24F6.Y14E’%c GEFT82F
U12FL 5TO8LGXFTFGEO8LF7 285NE2POL7 OYL1TWO2P!
XNFTFOGENFEPU8VO2L1TWO2U12FL 5TO8LGGEO8LF7 7OT‘2P
X8EN e*<*$ GEO8L8L5OL7 XFTFT14L7 8L GNOUF6(EO8L8L5
X8EN OL828LFY24F5OZFV1LG8GEFLETFG42EGOL7 SFTE82F
U12FL 5TO8LGS241TFGVF7 GET1L52P6-N4GW1GEWOE4TF
WO2F5OWFE1UNPEFGXFTF8L78VOEF7 E1YFSFTE82F6
EDR$-7&--$1.
BB -NFOLENFTXO21SN=1*,"(*S6 9$-$+"O97#.
V1LG8GEF7 1SS14T2OPFTG% FU87FTW8G! FL71ENFV84W!
W8772F2OPFTOL7 EOUFE4W&! XN8VN YF21L5F7 E1ENF
W1L1V1EP2F71L14GEPUF6]FOLXN82F! ENFEOUFE4W XOG
1S EPU8VO2 52OL742OT EPUF! ENF VPE1‘8LFG8G 1S
W8VT1GU1T1VPEF8L WF81G8GXOG1SG4VVFGG8ZFEPUF6-N8G
S8L78L58G8L O5TFFWFLEX8EN ENOE1SKOL5$-.#=
% $%%> &6 -NF W8VT1GU1TF EFETO7GXFTF Y82OEFTO2P
GPWWFET8VO2EPUF! XN8VN XOGG85L8S8VOLE2P 78SFTFLE
ST1WENOE1SKOL5$-.#=%$%%> & OL7 D4OL5$-.#6
% &""" &6 -NFP TFU1TEF7 TFGUFVE8ZF2P ENOE ENF
W8VT1GU1TF EFETO7G XFTF EFETONF7TO2EPUF 8L Y1EN
X$(/+"3.#.,211*(*321OL7 D97#"1-.3971O+.$3"@=.E
ENFGOWFE8WF! D4OL5$-.#=%&"""& U18LEF7 14EENOE
ENFTFXFTFEX1T1XG1SW8772F2OPFT8L ENFOLENFTG1S
D9=O+.$3"@OL7 1LF1SENFW8772F2OPFTG7F5FLFTOEF7
OL7 78GOUUFOTF7 8L G4YGFu4FLE7FZF21UWFLE6
g4T8L5U12FL 7FZF21UWFLEEOUFE4W 2OPFTGFTZFG
W1GE2POGOE8GG4FS1TWF81VPEF:GU1TFL4ET8E81L6*L
O778E81L E1EN8GWO8L S4LVE81L! ENFEOUFE4W NOG1ENFT
S4LVE81LG! LOWF2PENFUT174VE81L 1SENF21V42OTS2487!
ENFUT174VE81L OL7 TF2FOGF1SVO2OGF! ENFV1LZFP8L51S
I3.3(6U1G8E8ZFWOEFT8O2E1XOT7GENF21V424G! ENF
S1TWOE81L 1SFQ8LF UTFV4TG1TG! Z8GV8L ENTFO7GOL7
1TY8V42FG % biY8GVN Y178FG&! ENF UT174VE81L 1S
GU1T1UNPE8VUT1EF8LGOL7 FL\PWFG! OL7 1SU12FL‘8E:
ETPUN8LF%IOV8L8$-.#=! &"#A&6*L ENFUTFGFLEGE47P!
ENFEOUFE4WVF2GYF5OL E17F5FLFTOEFOEENFUTFWF81E8V
GEO5FOL7 XFTFOYG1TYF7 V1WU2FEF2P8L ENF7FZF21UWFLE
1SENFW8VT1GU1TFG6(8W82OTETFL7GXFTFTFU1TEF7 YP
D4OL5$-.#6%&"""&6
(8L52F 1Z42F X8EN UOT8FEO2 U2OVFLEO XOG
OLOET1U14G! 714Y2F8LEF54WFLEOL7 EFL48L4VF2OEOF6
-NFWF5OGU1T1VPEF4L7FTXFLEWF81E8V78Z8G81L E1S1TW
S14TWF5OGU1TFGOTTOL5F7 8L O28LF! OL7 ENFW8VT1UP2OT
WF5OGU1TFXOGENFS4LVE81LO21LF6-NFWOE4TFFWYTP1
GOVU1GGFGGF7 1LFVFLETO2VF2! 1LFF55VF2! OL7 EX1
GPLFT587GLFOTENFW8VT1UP2OTOL7 WOLPOLE8U17O2VF2G
XFTF1YGFTZF7 LFOTENFVNO2O\O2FL76-NFTFS1TF! ENF
7FZF21UWFLE1SFWYTP1S121XGENFD"#7%"(2,EPUF! OG
7FGVT8YF7 S1TH+*-*32,.$1-*)2, %D4! &"#A# $%%A&$
.VV1T78L5E1ENF7FZF21UWFLEO2V1TTFGU1L7FLVF
TF2OE81LGN8U YFEXFFL GEOWFLGOL7 U8GE82G! 8EVOL YF
7TOXL ENOE74T8L5ENFFOT2P7FZF21UWFLEGEO5F! ENF
7FZF21UWFLEO2GEO5F1SU8GE82GXOGFOT28FTENOL ENF
OLENFTG! XN8VN 8G G8W82OTX8EN ENF TFU1TEOY14E
X$(/+"3.#.,211*(*321 YP KOL5$-.#= % $%%> &6
D1XFZFT! ENF7FZF21UWFLE1SOLENFTGOL7 U8GE82GYF5OL
’9&
林 业 科 学 !" 卷B
E1YFGPLVNT1L8\F7 XNFL ENF2FL5EN 1SENFGU8‘F2FEGXOG
OY14E’ =’3A VW6*E8L78VOEF7 ENOEENF7FZF21UWFLE
GUFF7 1SENFOLENFTGXOG8WUT1ZF7 8L EN8GUFT8176
KOL5$-.#=% $%%> & V1LG87FTF7 ENOE8EO2XOPGXOG
UT1E15PLP74T8L5ENFXN12F7FZF21UWFLE8L GU8‘F2FEG1S
X=1*(*3214L28‘FN=1*,"(*S=9$-$+"O97#.=
iG4O2PYOWY11GNOZFO21L5S21XFT8L5UFT817!
Y4EENFL4WYFT1SST48EGUTFGFLEOEO58ZFL E8WFU18LE8G
4G4O2PZFTP21XOL7 78SFTGYFEXFFL GUFV8FG6(1WF
TFGFOTVN 7OEO8GOZO82OY2F! S1TFQOWU2F! GFF7 GFE8G
!c S1T;.19.(*. &.+%$1*! A3#c S1TD97#"1-.3971
)*"#.13$(1! OL7 >3&c S1TD97#"1-.3971*+*/$13$(1
%]V)24TF! &""’ &6 @1TG1WF GUFV8FG! G4VN OG
;.,L21. )2#%.+*1! XN8VN 8GX87F2PV42E8ZOEF7 OT14L7
ENFX1T27! OL7 ;.,L21. L.#3". 8L ?OL52O7FGN! GFF7G
NOZFLFZFTYFFL GFFL G1SOT6-NFGFTFG42EGG455FGEENOE
F8ENFTST48EGFE! 1T8EGWOE4TOE81L! 8GTF2OE8ZF2PTOTF8L
Y1EN E8WFOL7 GUOVF6*L W1GEYOWY11GENFS21TFEG
UT174V8L5WOE4TFST48EGOTFW1TF1T2FGGUT1WUE2P
TF2FOGF7 YPOYGV8GG81L ENFTOVN82OOL7 SO2 E1ENF
5T14L76 .L8WO2G! UOTE8V42OT2P VFTEO8L Y8T7G OL7
T17FLEG! NOTZFGEENFW ST1WENFU2OLE1TENF5T14L76
.LP1SENFGFSOVE1TG! F8ENFT1L ENF8T1XL 1T8L
V1WY8LOE81L! W85NE YF TFGU1LG8Y2F S1T ENF 21X
STFu4FLVP1SGFF7 GFEYPYOWY11GUFV8FG6fFLFTO2P!
ENFWO;1TSOVE1TGOTF28‘F2PE1YFV28WOE8VSOVE1TG!
OY1TE81L 1SOU1TE81L 1SENFU12FL 5TO8LG! OL7 ENF
L4WYFT1SS21XFT8L5U2OLEG5T1X8L5OEOG8EF6[4T
UTFZ814GGE47PGN1XF7 ENFGF2S1*,"(*S=9$-$+"O97#. 8G21X% 4LU4Y28GNF7 7OEO&6*L
ENFUTFGFLEGE47P! V1WUOTF7 E11ENFTYOWY11GUFV8FG!
N=1*,"(*S=9$-$+"O97#. U12FL OUUFOTGE1NOZFO
N85NFT2FZF21SZ8OY828EP % OY1ZF9%c&! G18E8G
8WU1GG8Y2FENOEENFU11TGFF7 GFE1SN=1*,"(*S=
9$-$+"O97#. 8GOGG1V8OEF7 X8EN U11TU12FL Z8OY828EP6*L
ENF,OL;8L5@1TFGETPiL8ZFTG8EPYOWY11V12FVE81L!
1L2PGFZFTO2U2OLEGS21XFTF7 8L ENFGEOL7G1SN=1*,"(*
S69$-$+"O97#.6KFS14L7 ENOEENFL4WYFT1SS21TFEG
1UFL8L58L OG8L52F7OP8G28W8EF7 74T8L5ENFS21XFT8L5
UFT8176]1TF1ZFT! S1TOG8L52FS21TFE! ENFUFT817 ST1W
ENF1UFL8L5E1ENFV21G4TF1SENFUO2FOOL7 2FWWO8G
GN1TE%1L2P& =$ N&6[SV14TGF! ENFV28WOEFVOL O2G1
OSFVEGFF7 GFE# S1TFQOWU2F! V1LGEOLE2PTO8LPXFOENFT
74T8L5OLENFG8GX82TF74VFENFSTFu4FLVP1SGFF7 GFE6
.EENFGOWFE8WFENF8LV87FLVF1S7FGET4VE81L 1S
8WWOE4TFST48EGYP8LGFVEG8GZFTPN85N 8L S21XFT8L5
LOE4TO2GEOL7G1SN=1*,"(*S=9$-$+"O97#.=
*L G4WWOTP! XF NOZF 7FGVT8YF7 ENF YOG8V
TFUT174VE8ZF 7FZF21UWFLEO2 VNOTOVEFT8GE8VG 1S
N+2(/*(.+*. 1*,"(*S=9$-$+"O97#. OL7 1YGFTZF7 ENOE
ENFU128LOE81L FSFVE8ZFLFGG8GU11TYFVO4GF1SENF
GWO2 L4WYFT 1S S21XFT8L5 YOWY11 U2OLEG6 KF
GUFV42OEFENOEEN8GW85NEYFOWO;1TVO4GFS1TENF
GUFV8FG5 21XGFF7 GFE6
参 考 文 献
.YF]! ]854VN8D! ,O‘OGN8\4‘O-6$%%&6.L 8LEFTOVE8ZFFSFVE1S
G8W42EOLF14G7FOEN 1S7XOTSYOWY11! VOL1UP5OU! OL7 UTF7OE1TP
T17FLEG1L YFFVN TF5FLFTOE81L6+V1215P! &$9%$& " $#& =$#>6
g4 @OL %杜凡 & ! k4F 8^OT1L5%薛嘉榕 &6$%%%6(E47P1L S21XFT8L5
UNFL1WFL1L OL7 8EGEPUF1SYOWY118L R4LLOL 8L UOGES8SEFFL PFOTG6
(V8(82(8L%林业科学& !’>%>& " A& =>#6
D4 )NFL5N4O%胡成华& !R4 @45FL %喻富根&6&""!6[YGFTZOE81L OL7
GE47P1L FWYTP1215P1S!9*,"("L.,L21. ,.+,"+$.=^?OWY11
CFGFOTVN%竹子研究汇刊& !&’%!& "> =&’6
D4 (N8P8%胡适宜&6&"#A6+WYTP1215P1SENFOL581GUFTWG%被子植物胚
胎学&6?F8;8L5" D85NFT+74VOE81L ITFGG%高等教育出版社& ! A# =
9%6
D4 (N8P8%胡适宜 &6&""!6]FEN17 1SUTFUOTOE81L 1SG287FG4GF7 E1
FQOW8LFENFU12FL 5FTW8LOE81L 1L ENFGE85WOOL7 U12FL E4YF5T1XEN
8L ENF GEP2F6 )N8LFGF?42FE8L 1S?1EOLP% 植 物 学 通 报 & !
&&%$& "A# =>%6
D4 (N8P8%胡适宜&6$%%A6CFUT174VE8ZFY81215P1SOL581GUFTWG%被子植
物生殖生物学&6?F8;8L5" D85NFT+74VOE81L ITFGG%高等教育出版
社& ! &%#6
D4OL5 8^OLu8L%黄坚钦& ! D4OL5D4ON1L5%黄华宏 & ! DF 8^S4 %何福
基& ! $-.#=&"""6-NFS1TWOE81L 1SW8VT1GU1TFOL7 ENF7FZF21UWFLE
1SWO2F5OWFE1UNPEF1SD97#"1-.3971O+.$3"@= 1^4TLO21S?OWY11
CFGFOTVN%竹子研究汇刊& ! &#%’& " AA =A#6
O^L\FL gD6&"9>6KNPYOWY11GXO8EG121L5E1S21XFT3 .LL CFZ+V12
(PGE! 9" ’!9 =’"&6
8^OL5MFN48%江泽慧&6$%%96?OWY11OL7 TOEOL 8L ENFX1T27%世界竹
藤&6?F8;8L5" )N8LO@1TFGETPI4Y28GN8L5D14GF%中国林业出版
社& ! &A =&96
/O8f4OL5N48%赖 广 辉 &6$%%&6(4UU2FWFLEOTP7FGVT8UE81L 1SS21TO2
W1TUN1215P 1S EFL GUFV8FG 8L ENF 5FL4G D97#"1-.3971
%?OWY4G187FOF&6 K^4NOL ?1ECFG%武汉植物学研究& ! &"%&& "
9 =&’6
/8MNO1N4O! gFL8VN ]! ?1TGVN -6$%%>6(8W42EOLF14GS21XFT8L51S
4WYTF2OYOWY11%>.+%$1*. ,2+*$#*.$& OE8EGLOE8ZFN1WF8L )FLETO2
)N8LO6 1^4TLO21S@1TFGETPCFGFOTVN! &9%!& "$"’ =$"96
/8FGFK6&"#A6?OWY11G’?81215P! G82Z8VG! UT1UFTE8FG! 4E828\OE81L6
f-M! +GVNY1TL!&%’ =&$%6
/8L (N4POL%林树燕 & ! DO1 4^OL;4OL %郝娟娟 & ! k8L D4O%辛华 &6
$%%"6 -NF WF5OGU1T15FLFG8G! W8VT1GU1T15FLFG8G OL7 ENF
7FZF21UWFLE 1S ENF8T SFWO2F OL7 WO2F 5OWFE1UNPEF 8L
!9&
B第 # 期 林树燕等" 异叶苦竹大小孢子及雌雄配子体的发育
5$(1-+2"3.#.,211*392.($(1*1=^,OL;8L5@1TFGETPiL8Z%南京林业
大学学报& ! ’’%’& "" =&$6
]V)24TF@.6&""’6-NF?OWY11G6KOGN8L5E1L OL7 /1L71L" (W8ENG1L8OL
*LGE8E4E81L ITFGG! "A ="96
IOV8L8+! @TOLVN8f f! DFGGF]6&"#A6-NFEOUFE4W" *EGS1TW!
S4LVE81L! OL7 U1GG8Y2FUNP215FLP8L ’,L+7"O97-.6I2OLE(PGEFWOE8VG
OL7 +Z124E81L! &!"%’ :!& "&AA =&#A
J8O1(N8P8%乔士义 & ! /8O1f4OL524 %廖光庐 &6&"#!6+WYTP1215P
1YGFTZOE81L 1SD97#"1-.3971O2L$13$(1=?OWY11CFGFOTVN %竹类研
究& ! ’ %&& "&A =$’6
-OP21T.D! J8L M(6&"##6CF5FLFTOE81L ST1WGFF7 1SI*(.+2(/*(.+*.
&.(%*.(.! OYOWY11! 8L ENFK121L5f8OLEIOL7OTFGFTZF! (8VN4OL!
)N8LO6.W ?^1E! 9A" &%>A =&%9’6
-OP21T.D! J8L M(6&""’6?OWY11TF5FLFTOE81L OSEFTS21XFT8L58L ENF
K121L5f8OLEIOL7OTFGFTZF6?812)1LG! >%’& " $’& =$’!6
KOL5(N454OL5%王曙光& ! I4 k8O12OL%普晓兰& ! g8L5R421L5%丁雨
龙&6$%%>6-NFGET4VE4TFG1STFUT174VE8ZF1T5OLGOL7 7FZF21UWFLE1S
ENFSFWO2FOL7 WO2F5OWFE1UNPEF1SX$(/+"3.#.,211*(*321=?42FE8L
1S?1ECFGFOTVN%植物研究& ! $> %’& " $9% =$9!6
KFL -O8N48%温太辉&6&""’6)121TF7 824GETOE81LG1S?OWY4G187FOF8L
)N8LO%中国竹类彩色图鉴 &6-O8UF8" (N4Q8L ITFGG%淑馨出版
社& ! "’ ="A6
k8O,8OLNF%夏念和 & ! 8^O/8OL5\N8%贾良智 &6&""A6(1WFLFXOL7
8WUFTSFVE2P‘L1XL GUFV8FG1SYOWY11GST1W)N8LO6^-T1U (4YET1U
?1E%热带亚热带植物学报& ! !%&& " $’ =’%6
R8-1L5UF8%易同培&6&""96?OWY11GS21TO1S(8VN4OL%四川竹类植物
志&6?F8;8L5" )N8LO@1TFGETPI4Y28GN8L5D14GF%中国林业出版
社& ! $#! =$#>6
!责任编辑B徐B红"
)林产化学与工业*征订启事
)林产化学与工业*%双月刊!&"#& 年创刊&!由中国林业科学研究院林产化学工业研究所(中国林学会
林产化学化工分会共同主办!为全国林产化工行业的学术类期刊$ 报道范围是可再生的木质和非木质生物
质资源的化学加工与利用!包括生物质能源(生物质化学品(生物质新材料(生物质天然活性成分和制浆造纸
等!主要包括松脂化学(生物质能源化学(生物质炭材料(生物基功能高分子材料(胶黏剂化学(森林植物资源
提取物化学利用(环境保护工程(木材制浆造纸为主的林纸一体化和林产化学工程设备研究设计等方面的最
新研究成果$
本刊现被美国1工程索引2%+*&(美国1化学文摘2%).核心&(荷兰1文摘与引文数据库2 % (V1U4G&(美国
.乌利希国际期刊指南/(英国1英联邦农业和生物科学文摘2%).?.YGETOVEG&(英国1林产品文摘2%@I.&(英
国1全球健康2(英国1皇家化学学会系列文摘2%C()&(俄罗斯1文摘杂志2 %I8&等国外数据库收录#被.中
国科学引文数据库%)()g&/(.中文核心期刊/(.中国科技核心期刊/(.C))(+中国核心学术期刊%.&/(
.中国农业核心期刊/(.中国期刊全文数据库/(.中国学术期刊综合评价数据库/(.万方数据’’’数字化期
刊群/(.中文科技期刊数据库/(.中国核心期刊%遴选&数据库/(1中国学术期刊文摘2等国内 &% 多种大型
刊库收录!$%%#($%&& 年连续两届被评为.中国精品科技期刊/$
本刊为双月刊!逢双月月末出版!大 &> 开!定价"国内RcC8B99 元:期%全年 M9B99 元#国外(-C8699 美元:
期%全年(-M9699 美元$
本刊刊号"*((,%$A’ =$!&9!),’$ =&&!":($ 国内外公开发行!国内邮发代号"$# =A"#国外发行代
号"JA"!&$
地B址!$&%%!$ 江苏省南京市锁金五村 &> 号 林化所内BBBBB电B话!%%$A&#A!#$!"’
传B真!%%$A&#A!#$!"’BBBB/T%"$K" V8SUˆZ8U6&>’6V1WBBB6++)"::XXX6V8SU6OV6VL
A9&