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Dynamics of Species Composition and Generation Pattern in the Gaps of Karst Forest in maolan,Guizhou Province

茂兰喀斯特森林的林隙物种组成动态及更新模式*


分析茂兰喀斯特森林林隙物种组成和种间关系随林隙发育的动态变化,并对林隙主要树种的更新模式进行探讨。结果表明:在发育时间为10、20、30和40年的序列上,林隙中乔木层优势度最大的物种分别是小叶柿、黄连木、掌叶木和云贵鹅耳枥;灌木层优势种包括贵州悬竹、革叶铁榄、湖北十大功劳和长梗核果木;草本层优势种包括石韦、瓦韦、翠云草和庐山楼梯草;藤本优势种包括鞘柄菝葜、四川清风藤、藤黄檀和圆锥清风藤。非林隙林分的优势种不明显。在林隙发育过程中乔木层、灌木层、草本层和藤本的平均相似系数分别为0.31、0.42、0.27和0.32,说明在林隙发育过程中,灌木种类具有较高的连续性和稳定性;草本植物则随着林隙环境条件的改变而出现较大的波动性,藤本植物和乔木的稳定性和连续性介于灌木和草本植物之间。依据树木幼苗、幼树和林冠层数量的差异,可将林隙树种的更新模式分为4种类型。

Dynamics of species composition and the correlation coefficients of species in each layer were studied and generation patterns of major tree species were discussed among developmental stages of gaps in Karst forest in Maolan Nature Reserve,Guizhou Province.Along the gap developmental series of 10,20,30 and 40 years,the dominant species in the arbor layer were Diospyros dumetorum,Pistacia chinensis,Handeliodendron bodinieri and Carpinus pubescens.The dominant species in the shrub layer were Ampelocalamus calcareous,Sinosideroxylon wightianum,Mahonia confusa and Drypetes longipes,while the dominant species in the herb layer were Pyrrosia lingua,Lepisorus,,Selaginella uncinata and Elatostema stewardii.As for the liana,Smilax stans,Sabia schumanniana,Dabergia hancei and Sabia paniculata were the common species.In the non-gap stands,there were no dominant species.The similarity coefficients of trees,shrubs,herbaceous species and liana species with different gap stages were 0.31,0.42,0.27 and 0.32,respectively.This indicated that the composition of shrub species remained relatively stable during the past 40 years of gap development,while the composition of herbaceous species fluctuated greatly.Trees replacement patterns could be classified into 4 groups according to quantitative dynamics of seedlings,saplings and trees in different gap stages.


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