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Six new species and two new varieties of Ranunculaceae from southwest China

华西南毛茛科六新种和二新变种



全 文 :书广 西 植 物 Guihaia Sept.2013,33(5):579-587           http://journal.gxzw.gxib.cn
DOI:10.3969/j.issn.1000-3142.2013.05.001
王文采.华西南毛茛科六新种和二新变种[J].广西植物,2013,33(5):579-587
Wang WT.Six new species and two new varieties of Ranunculaceae from southwest China[J].Guihaia,2013,33(5):579-587
华西南毛茛科六新种和二新变种
王文采
(中国科学院植物研究所 系统与进化植物学国家重点实验室,北京100093)
摘 要:该文描述了自华西南东部发现的毛茛科翠雀属2新种和1新变种、唐松草属1新种、银莲花属1新种
和1新变种,以及毛茛属2新种。
关键词:毛茛科;翠雀属;唐松草属;银莲花属;毛茛属;新分类群;华西南
中图分类号:Q944  文献标识码:A  文章编号:1000-3142(2013)05-0579-09
Six new species and two new varieties of
Ranunculaceae from southwest China
WANG Wen-Tsai
(State Key Laboratory of Systematic and Evolutionary Botany,Institute of Botany,
Chinese Academy of Sciences,Beijing 100093,China)
Abstract:Two species and one variety of Delphinium,one species of Thalictrum,one species and one variety of
Anemone,and two species of Ranunculus(Ranunculaceae)are described as new from southwest China.
Key words:Ranunculaceae;Delphinium;Thalictrum;Anemone;Ranunculus;new taxa;southwest China
  今年春季,哈佛大学D.E.Boufford博士寄来
一批近年采自西藏东部和四川西部的毛茛科植物标
本托我鉴定,从中发现翠雀属2新种和1新变种以
及银莲花属草玉梅的1新变种。今年夏季,同事杨
永博士、刘冰博士等到西藏东部进行植物区系考察,
也将采集的毛茛科植物标本托我鉴定,从中发现唐
松草属和银莲花属各1新种,以及毛茛属2新种。
  长苞翠雀花 图1
Delphinium longibracteolatum W.T.Wang,sp.
nov.Fig.1
(sect.Elatopsis Huth)
Species nova haec est afinis D.densifloro Duth-
ie ex Huth,quod foliorum laminis basi profunde corda-
tis,pedicelis supra medium 2-bracteolatis,bracteolis
minoribus linearibus ca.1.5cm longis 1-2mm latis
densiuscule pubescentibus et ciliatis,calcari sepalino
conico sepalis multo breviore,staminodiorum limbis
ventre dense longeque barbatis facile distinguitur.
Perennial herbs.Stems ca.50cm tal,below 6mm
thick,sparsely puberulous,simple,1-leaved.Basal
leaves ca.4,long petiolate;blades papery,subsemi-or-
bicular,(7-10)cm×(12-16.8)cm,3-lobed,apex
rounded-truncate,base subtruncate,central lobe smal,
ovate,3-3.5cm long,with upper margins lobulate or
incised and dentate,lateral lobes larger,obliquely fla-
belate,unequaly 2-3-lobed,with upper margins in-
cised and dentate;surfaces glabrous or abaxialy near
base on nerves with sparse hairs;petioles 8-17cm
long,sparsely spreading-pubescent.Terminal raceme ca.
20cm long,below loosely above densely flowered;bracts
greenish-white,blue-nerved,shortly petiolate,lanceolate,
(2-3)cm×(0.45-0.6)cm,glabrous or sparsely
收稿日期:2012-11-22  修回日期:2013-01-16
作者简介:王文采(1926-),男,山东掖县人,研究员/教授,中国科学院院士,著名植物分类学家,长期从事有花植物分类学研究。
图1 长苞翠雀花 A.开花茎;B.退化雄蕊;C.雄蕊;D.3枚心皮。(根据模式标本,孙英宝绘图)
Fig.1 Delphinium longibracteolatum A.habit;B.staminode;C.stamen;D.three carpels.(from holotypes,Drawn by SUN Ying-Bao)
pubescent;rhachis sparsely spreading-pubescent;pedi-
cels 0.8-2.5cm long,sparsely spreading-pubescent
or subglabrous,near apex 2-bracteolate;bracteoles sim-
ilar to bracts,narrow-lanceolate or linear-lanceolate,
(1.4-2.4)cm×(0.2-0.4)cm,glabrous or with
few hairs.Flower ca.3cm long:Sepals abaxialy
greenish-white,purple-nerved,adaxialy dark purple
(according to the colectors),entirely glabrous,abaxial-
ly sparsely pubescent,upper sepal eliptic-ovate,ca.1.
5cm×0.7cm,spur conical-subulate or subulate,1.3
-1.4cm long,with base 4-4.5mm thick,lateral
sepals ovate-oblong,ca.1.6cm×0.6cm,lower sepals
lanceolate,ca.1.4cm×(0.3-0.5)cm.Petals
black,glabrous,2-lobulate at apex.Staminodes black;
085 广 西 植 物                  33卷
limbs ovate,ca.7mm long,adaxialy and abaxialy
glabrous,unequaly 2-lobulate,near apex sparsely cili-
ate;claws as long as limbs,above base minutely 1-ap-
pendiculate.Stamens ca.1cm long,glabrous;filaments
lanceolate-linear,ca.8 mm long;anthers black,ob-
long,ca.1.2mm long.Carpels 3;ovaries oblong,ca.
5mm long,densely puberulous;steyles 1.5mm long,
glabrous.
Tibet:Jiangda County.Narrow gorge and
rocky slopes ca.6km E in a straight line from the
center of Jiangda City.31°30′19″N,98°17′22″
E;4 400-4 800m.Kobresia meadow giving way
to slopes with dwarf(<5mtal)Potentilla fruti-
cosa and Caragana,then rocky slope and rocky
ridge.On slope at base of shrubs of Caragana.
Tepals outside pale greenish white with purple vena-
tion,inside dark purple.August 4,2009.D.E.Bouf-
ford,B.Bartholomew,D.A.Eaton,X.H.Li,R.H.
Lee,B.Xu,J.P.Yue,J.W.Zhang & X.X.Zhu
No.41846(holotype,PE;isotype,GH,not seen).
本种在亲缘关系上稍接近密花翠雀花D.den-
siflorum Duthie ex Huth,与后者的区别在于本种
的叶片基部近截形,花梗顶端生2小苞片,小苞片较
大,狭披针形,近无毛,萼距近钻形,退化雄蕊腹面无
毛。在密花翠雀花,叶片基部深心行,花梗在中部之
上生2小苞片,小苞片较小,条形,长约1.5cm,
宽1-2mm,稍密被柔毛和缘毛,萼距筒状圆锥形,
基部粗5-6mm,退化雄蕊腹面被长髯毛(Munz,
1967;Wang,1979)。
  左贡翠雀花 图2:A-E
Delphinium zuogongense W.T.Wang,sp.nov.
Fig.2:A-E.
(sect.DelphinastrumDC.)
Species nova haec est afinis D.umbroso Hand.-
Mazz.,quod caulibus retrorse strigilosis,inflorescentis
racemosis,rhachidibus dense albo-strigilosis et patule
luteolo-glanduloso-puberulis,floribus majoribus,sepal-
is 8.5-15mm longis,calcari sepalino 20-40mm
longo,carpelorum ovaris pilosis difert.
Perennial herbs.Stems ca.80cm tal,below ca.
7mm thick,glabrous.Lower cauline leaves long peti-
olate;blades papery,pentagonal,(4-5.8)cm×(8-
9)cm,3-parted nearly to base,base deeply cordate,
central lobe narrow-rhombic,1.8-2cm broad,with
acute apex and lobulate and dentate upper margins,
lateral lobes larger,obliquely flabelate,unequaly 2-
parted;surfaces adaxialy appressed-puberulous,abaxi-
aly on nerves puberulous;petioles 7-12cm long,
sparsely spreading-pubescent or subglabrous.Panicle
terminal,ca.34cm×15cm,abunding in flowers;
bracts shortly petiolate,lower bracts foliaceous,ca.
3cm long,3-parted to 3-sect,upper bracts narrow-
lanceolate,1-1.8cm long;rhachis below glabrous,
near apex densely yelowish-glandular-puberulous;ped-
icels 1-4cm long,near apex 2-bracteolate and densely
yelowish-glandular-puberulous;bracteoles 2-5 mm
remote from flower,linear or narrow-linear,(4-7)cm
×(0.2-0.4)mm,subglabrous.Flower smal,ca.2
cm long.Sepals purple(according to the colectors),
adaxialy glabrous,abaxialy yelowish-glandular-pube-
rulous,upper sepal ovate,7-10mm long,spur subu-
late,1.1-1.3cm long,arcuate-decurved,with base 3
-4mm thick,lateral sepals narrow-obovate-oblong,7
-9mm long,lower sepals as long as lateral sepals,
narrow-obovate or narrow-oblanceolate.Petals gla-
brous,apex entire.Staminodes 2;limbs deep blue,quad-
rate-ovate,ca.4mm long,unequaly 2-lobed,sparsely
ciliolate,adaxialy above base white-barbate;claws as
long as limbs,glabrous,above base minutely 1-appendic-
ulate.Stamens ca.5mm long,glabrous;filaments lan-
ceolate-linear,ca.4mm long;anthers black,oblong,1
mm long.Carpels 3,ca.3mm long,glabrous.
Tibet:Zuogong County.S of Zuogong City along
the Yu Qu(Yu River)Y 563.Along smal tributary,
29°29′40″N,97°56′38″E;4 000-4 100m.Deforested
ravine along smal stream (originaly Picea forest),
with Rhododendron,Ribes,Juniperus.Abies forest
and beside road.Tepals purple.July 18,2009.D.E.
Bouford,B.Bartholomew,D.A.Eaton,X.H.Li,
R.H.Ree,B.Xu,T.P.Yue,J.W.Zhang &X.X.
Zhu No.40721 (holotype,PE;isotype,GH,not
seen).
本种接近阴地翠雀花 D.umbrosum Hand.
-Mazz.,与后者的区别在于本种的茎无毛,花序圆锥
状,花序轴下部无毛,花较小,萼片长7-10mm,萼
距长11-13mm,心皮无毛。在阴地翠雀花,茎被反
曲短糙伏毛,花序总状,花序轴全部密被白色短伏毛
和黄色开展短腺毛,花较大,萼片长8.5-15mm,
1855期            王文采等:华西南毛茛科六新种和二新变种
图2 A-E.左贡翠雀花 A.中部茎生叶;B.顶生圆锥花序;C.退化雄蕊;D.雄蕊;E.3枚心皮。F-G.光果拟
螺距翠雀花 F.中部茎生叶;G.花。(根据模式标本,孙英宝绘图)
Fig.2 A-E.Delphinium zuogongense A.middle cauline leaf;B.terminal panicle;C.staminode;D.stamen;E.three
carpels.F-G.D.buleyanumvar.leiogynum F.middle cauline leaf;G.flower.(from holotypes,Drawn by SUN Ying-
Bao)
萼距长20-24mm,心皮子房被疏柔毛 (Munz,
1967;Wang,1979)。
  光果拟螺距翠雀花 图2:F-G
Delphinium buleyanumDiels var.leiogynum W.
T.Wang,var.nov.Fig.2:F-G.
(sect.DelphinastrumDC.)
A var.bulleyano difert carpelis glabris.
Sichuan:Danba County:SE of Danba City,on
285 广 西 植 物                  33卷
road(highway 303)to Daofu and Kangding at Ding-
guo Shan.30°36′40″N,101°45′22″E;2 800-2 950
m.Remnant Tsugaforest with scattered Picea and
Abies and shrubs and smal trees of Malus,Salixand
Betula and shrubs of Rosa.Open areas and at edge of
forest.Herbs,ca.1mtal,flowers blunish purple.
Scattered throughout.24July,2007.D.E.Bouf-
ford,K.Fujikawa,S.L.Keley,R.H.Ree,B.Xu,
J.W.Zhang,T.C.Zhang & W.D.Zhu No.38028
(holotype,PE;isotype,GH,not seen).
本种与模式变种的区别在于心皮无毛。模式变
种,心皮的子房被柔毛 (Munz,1967;Wang,1979)。
  美花唐松草 图3:A-E
Thalictrum calianthum W.T.Wang,sp.nov.
Fig.3:A-E
Species nova haec est afinis T.delavayi Franch.,
quod foliolis majoribus usque ad 3cm longis 2-2.5cm
latis,sepalis minoribus usque ad 9(-12)mm longis,
antheris oblongis,stylis ovaris multo brevioribus sub
fructu apice haud hamatis,achenis ad suturas dorsales
ventralesque anguste alatis difert.
Perennil herbs.Stems ca.1.5mtal,glabrous,
branched.Middle cauline leaves short petiolate,3-4-
ternately compound,glabrous;blades deltoid or trian-
gular in outline,16-18cm long,22-25cm broad;
leaflets numerous,thin papery,terminal leaflets broad-
obovate,7-10 mm long,5-9 mm broad,3-
lobed,apex long apiculate,lateral leaflets obliquely
ovate,obliquely quadrate or obliquely narrow-obovate,
4-9mm long,3-9mm broad,3-lobulate or undivid-
ed,apex acute;petioles ca.5cm long,base brown-
vaginate.Terminal thyrse ca.30cm long,ca.30-
flowered,glabrous;bracts foliaceous,2.5-7cm long;
pedicels slender,2-4.5cm long.Flower 2-3cm in
diam.,beautiful;sepals 4(5),purple(according to the
colectors),spreading,long eliptic or narrow-ovate,11
-16mm long,5-10mm broad,glabrous,apex mu-
cronate;stamens 22-28,5-8.5mm long,glabrous,
filaments filiform,anthers yelow,narrow.linear,1.5
-2mm long,apex apiculate;carpels 10-15,glabrous,
ovaries green,eliptic,ca.1mm long,styles slender,
dark-blue,ca.1mm long,stigmas yelow,narrow-line-
ar,0.8-1mm long,basal stipes 0.5-0.8mm long.
Achenes flattened,sublunate,ca.5mm long,1.5-2
mm broad,glabrous,on each side prominently and lon-
gitudinaly 1-3-nerved,basal stipes ca.1mm long,
persistent styles 2-2.2mm long,apex hooked.
西藏 (Tibet):米林 (Milin),派区 (Paiqu),格噶
村 (Gega vilage),alt.3 400m,灌丛中 (in shrub-
bery),植物高1.5m (pl.1.5mtal),花紫色 (fls.
purple),2012-08-01,杨永,刘冰,林秦文,赖阳均,
冯涛 (Y.Yang,B.Liou,Q.W.Lin,Y.J.Lai &T.
Feng)471(holotype,PE)。
本种与偏翅唐松草T.delavayi Franch.近缘,
与后者的区别在于本种叶的小叶较小,长4-10mm,
宽3-9mm,萼片较大,长11-16mm,花药狭条形,
花柱细长,与子房等长,果期伸长后顶端呈钩状弯曲,
瘦果无翅。在偏翅唐松草,小叶较大,长达3cm,宽
达2-2.5cm,萼片较小,长达9(-12)mm,花药长
圆形,花柱比子房短数倍,果期顶端不弯曲,瘦果沿
背缝线和腹缝线具狭翅(王文采,1979)。
在1991、2001年我分别到瑞典乌普萨拉和瑞士
日内瓦访问时,曾在两地的植物园中看到栽培有特
产我国云南、四川的偏翅唐松草,本种的花比偏翅唐
松草的花还要大,呈紫色,也更加美丽,值得引种作
为观赏植物。
  米林银莲花 图3:F-K
Anemone milinensis W.T.Wang,sp.nov.Fig.3:F-

Species nova haec est afinis A.rivulari
Buch.-Ham.ex DC.,quae folis utrinque strigosis,flo-
ribus pluribus in cymam compositam bis vel ter ramo-
sam dispositis,bracteis petiolatis,floris sepalis albis
pluribus(6-)7-8(-10)eliptico-obovatis dorso
sparse pubescentibus apice dense puberulis facile re-
cedit.
Perennial herbs.Rhizome unknown.Basal leaves
several,long petiolate,glabrous;blades papery,deltoid
or orbicular-ovate,1.2-1.5cm long,1.7-2.6cm
broad,3-sect,base deep-cordate,central segment
broad-ovate,ca.1.5cm broad,3-parted,partitions
with 2-3ultimate ovate lobules,lateral segments ob-
liquely flabelate,unequaly 3-parted;petioles 9-13.5
cm long.Scape 12-16cm tal,glabrous;involucre
consisting of 3sessile bracts,which are 1.1-1.3cm
long,2-3-fid,with lobes narrow-triangular or sublin-
ear,entire or 3-lobulate.Flower solitary,terminal,or 2
flowers arranged in a terminal simple monochasium;
3855期            王文采等:华西南毛茛科六新种和二新变种
图3 A-E.美花唐松草 A.茎中部叶;B.聚伞圆锥花序;C.雄蕊;D.心皮;E.瘦果。F-K.米林银莲花 F.基生叶;G.花
葶;
H.花;I.雄蕊;J.雌蕊群;K.心皮纵切面。(根据模式标本,孙英宝绘图)
Fig.3 A-E.Thalictrum calianthum A.middle cauline leaf;B.thyrse;C.stamen;D.carpel;E.achene.(from holotype) F-K.A-
nemone milinensis F.basal leaf;G.scape;H.flower;I.stamen;J.gynoecium;K.longitudinal section of carpel.(from holotypes,Drawn by
SUN Ying-Bao)
pedicels 1-12mm long,glabrous;sepals 5,dilutely
blue-purple(according to the colectors),broad-obo-
vate,ca.6mm long,4-4.5mm broad,glabrous,apex
rounded or subtruncate;stamens ca.18,glabrous,fila-
ments white,linear,1-2mm long,anthers dark-blue,
narrow-oblong,1.2-1.5mm long;carpels ca.16,gla-
485 广 西 植 物                  33卷
brous,ovaries green,narrow-oblong,1-2mm long,1-
ovulate,styles 0.5-1mm long,apex hooked.
西藏(Tibet):米林(Milin),派区(Paiqu),多雄
拉山(Mount Duoxiongla),alt.4 200m,高山草地
(alpine meadow),花淡蓝紫色(fls.dilutely blue-
purple),2012-07-30,杨永,刘冰,林秦文,赖阳
均,冯涛(Y.Yang,B.Liou,Q.W.Lin,Y.J.
Lai &T.Feng)319(holotype,PE)。
本种与草玉梅 A.rivularis Buch.-Ham.ex
DC.近缘,与后者的区别在于本种的叶无毛,花单
朵顶生或2朵组成一顶生简单单歧聚伞花序,苞片
无柄,花的萼片淡蓝紫色,5枚,宽倒卵形,无毛。在
草玉梅,叶两面被糙伏毛,花较多,组成2-3回分枝
的复杂聚伞花序,苞片具柄,花的萼片白色,较
多,(6-)7-8(-10)枚,椭圆形状倒卵形,背面疏被
柔毛,顶端密被短柔毛(王文采,1980)。
  疏毛草玉梅  (变种)
Anemone rivularis Buch.-Ham.ex DC.var.pilo-
sisepala W.T.Wang,var.nov.
A var.rivulari difert cyma simplici 3-flora,se-
palis apice sparse pilosis.
Tibet:Mangkang(Markham).Between Mang-
kang and the Jinsha Jiang on highway 318,along Xi
Qiu(Xi River).29°43′78″N,98°46′37″E;3 300m.
Ravine with dwarf Quercus and mixed shrubs on
steep,rocky slopes.Among rocks in thickets and
clearings.Sepals abaxialy purple,adaxialy white.Ju-
ly 16,2009.D.E.Bouford,B.Bartholomew,D.A.
Eaton,X.H.Li,R.H.Ree,B.Xu,J.P.Yue,J.
W.Zhang &X.X.Zhu No.40614(holotype,PE;i-
sotype,GH,not seen).
本变种与模式变种的区别在于本变种的花序为
简单聚伞花序,具3朵花,萼片顶端疏被柔毛。在模
式变种,花序为2-3回分枝的复杂聚伞花序,具较
多花,萼片顶端密被短柔毛(王文采,1980)。
  多雄拉毛茛 图4:A-D
Ranunculus duoxionglashanicus W.T.Wang,sp.
nov.Fig.4:A-D
Species nova haec est afinis R.petrogeito Ulbr.,
qui foliorum basalium segmentis centralibus 3-fidis,
floris staminibus numerosis,antheris luteis difert.
Perennial smal herbs.Stems ca.3fascicled,12-
18 cm tal,appressed-puberulous,simple or 1-
branched,1-2-foliate.Basal leaves several,long petio-
late;blades thin-papery,pentagonal,1-1.2cm long,
ca.1.8cm broad,3-parted or 3-sect,base deep-cor-
date,central segment obovate,with rounded-obtuse
apex,undivided or inconspicuously 3-lobulate,lateral
segments obliquely broad-obovate,unequaly 2-lobu-
late;surfaces adaxialy appressed-puberulous,abaxialy
glabrous;petioles 6.5-9cm long,glabrous.Lower
cauline leaves shortly petiolate,their blades 1.3-1.8
cm broad,3-lobed or 3-parted,with oblong entire
lobes;upper cauline leaves smaler,shortly petiolate or
subsessile,oblong-eliptic,1-1.4cm long,entire.
Flower solitary,terminal,7-8mm in diam.;recepta-
cle glabrous;sepals 5,long eliptic,3-5mm long,1-
2mm broad,abaxialy pubescent,apex slightly acute or
obtuse;petals 5,yelow,broad-eliptic,3-4mm long,2
-2.8 mm broad,glabrous,longitudinaly 3-5-
nerved,apex rounded or obtuse,basal nectary pit mi-
nute;stamens ca.10,whitish,1.8-2mm long,gla-
brous,filaments narrow-linear,anthers oblong or nar-
row-oblong,0.6-0.7mm long;gynoecium broad-o-
void,ca.3.2mm long;carpels ca.50,flattened,gla-
brous,ovaries obliquely broad-ovate,ca.1mm long,
styles 0.6-0.8mm long.
西藏(Tibet):米林(Milin),派区(Paiqu),多雄
拉山(Mount Duoxiongla),山口 (mountain pass),
alt.4 200m,高山草地 (alpine meadow),2012-
07-30,杨永,刘冰,林秦文,赖阳均,冯涛 (Y.
Yang,B.Liou,Q.W.Lin,Y.J.Lai &T.Feng)
281(holotype,PE)。
本种与太白山毛茛R.petrogeiton Ulbr.近
缘,与后者的区别为本种的基生叶中全裂片不分裂
或不明显微3裂,花的雄蕊较少,约10枚,呈白色。
在太白山毛茛,基生叶的中全裂片3裂近中部,花的
雄蕊多数,花药黄色(Wang &Gilbert,2001)。
  十蕊毛茛 图4:E-I
Ranunculus decandrus W.T.Wang,sp.nov.Fig.
4:E-I
O b petala oblonga et carpela glabra species
nova haec est affinis R.micronivali H and.-
M azz.,qui caulibus hum ilim is 0.6-5cm al-
tis saepe efoliatis,folis m inoribus,lam inis 1.8
-5mm longis 3-7mm latis,flore m inore 0.
6-0.8mm in diam etro,stam inibus
5855期            王文采等:华西南毛茛科六新种和二新变种
图4  A-D.多雄拉毛茛 A.植株全形;B.花瓣;C.2雄蕊;D.心皮。E-I.十蕊毛茛 E.植株全形;F.2花瓣;G.2雄蕊;H.
心皮;I.瘦果。(根据模式标本,孙英宝绘图)
Fig.4 A-D.Ranunculus duoxionglashanicus A.habit;B.petal;C.two stamens;D.carpel.(from holotype) E-I.R.decandrus
 E.habit;F.two petals;G.two stamens;H.carpel;I.achene.(from holotypes,Drawn by SUN Ying-Bao)
pauciori-bus 3-7,stylis ovaris leviter longioribus dis-
tinguitur.
Perennial smal herbs.Stems ca.4fascicled,6.5
-18cm tal,puberulous,1-2-branched,occasionaly
simple.Basal leaves several,long petiolate;blades pa-
pery,orbicular-ovate or pentagonal,0.5-1.5cm long,
1.1-2.5cm broad,3-parted or 3-sect,base deep cor-
date,central segment broad-obovate,undivided or 3-
685 广 西 植 物                  33卷
lobulate,lateral segmemts obliquely flabelate,une-
qualy 2-lobed,margin ciliate;surfaces glabrous or ad-
axialy sparsely appressed-pubescent;petioles 3-10cm
long,glabrous.Lower cauline leaves slightly short pet-
iolate,3-parted;upper cauline ones smal,subsessile,3-
lobed.Flower solitary,terminal,7-9mm in diam.;
receptacle glabrous;sepals 5,long eliptic,2-4mm
long,1-2mm broad,abaxialy sparsely pubescent;
petals 5,yelow,oblong or cuneate-oblong,3.2-5.5
mm long,1.4-2mm broad,glabrous,apex rounded-
obtuse or truncate,claws 0.5-0.8mm long,nectary
pit minute;stamens ca.10,whitish,3-3.2mm long,
glabrous,filaments narrow-linear,anthers oblong-elip-
tic,ca.1mm long;gynoecium ovoid,ca.5mm long;
carpels ca.60,glabrous,ovaries flattened,obliquely
broad-ovate,ca.1.2mm long,styles 0.3-0.6mm,
abaxialy stigmatose.Aggregate fruit narrow-ovoid,
ca.8mm long,at base 5mm thick;achenes bilateraly
compressed,obovoid,ca.1.2mm long,l mm broad,
glabrous,with persistent styles ca.0.5mm long.
西藏(Tibet):林芝(Linzhi),色季拉山(Mount
Sejila),alt.4 400m,流石滩(on screes),2012-08-05,
杨永,刘冰,林秦文,赖阳均,冯涛(Y.Yang,B.Liou,
Q.W.Lin,Y.J.Lai & T.Feng)644(holotype,
PE).
本种的花瓣长圆形,心皮无毛,而与窄瓣毛茛
R.micronivalis Hand.-Mazz.相近,与后者的区别在
于本种的茎较高,高6.5-18cm,具2-3叶,基生叶
较大,叶片长5-15mm,宽11-25mm,花较大,直径
7-9mm,雄蕊约10枚,花柱比子房短2-4倍。在
窄瓣毛茛,茎低矮,高0.6-5mm,常无叶,叶较小,叶
片长1.8-5mm,宽3-7mm,花较小,直径0.6-0.
8mm,雄蕊较少,3-7枚,花柱长度稍超过子房
(Wang &Gilbert,2001)。
致谢 蒙D.E.Bouford博士惠赠珍贵标本,杨
永博士、刘冰博士提供标本,孙英宝先生绘图。
参考文献:
王文采.1979.Delphinium.中国植物志[M].北京:科学出版
社,27:326-462
王文采,王蜀秀.1979.Thalictrum.中国植物志[M].北京:科
学出版社,27:52-592
王文采.1980.Anemone.中国植物志[M].北京:科学出版社,
28:1-56
Munz PA.1967-1968.A synopsis of the Asian species of Del-
phinium,sensu stricto[J].J Arnold Arbor,48:249-302,476
-545;49:73-166,233-259
Wang WT,Gilbert MG.2001.Ranunculus[M]//Wu CY,Raven
PH(eds).Flora of China.Beijing:Science Press;St.Louis:
Missouri Botanical Garden Press,6:391-431
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7855期            王文采等:华西南毛茛科六新种和二新变种