Abstract:Sporogenesis and the development of male and female gametophytes inAnoectochilus roxburghii are studied. The mature ovules are anatropous, bitegminous andtenuinucellatae. The development of the female gametophyte is of polygonum type. Matureembryo sac has 7 cells. In the development of mega- and microsporogenesis, calloseappears. The microspore tetrads are tetrahedral isobilateral, decussate, T-shaped and remainin massulae. At shedding stage the pollen grains are 2-celled. The anther wall consists offour-layers with secretory tapetum.