免费文献传递   相关文献

Cytological Advances on Tribe Polygonateae (Asparagaceae)

天门冬科黄精族细胞学研究进展



全 文 :书!"#$%&
,2016,36(4):0834-0845
犃犮狋犪犅狅狋.犅狅狉犲犪犾.犗犮犮犻犱犲狀狋.犛犻狀.
  !"#$:10004025(2016)04083412               犱狅犻:10.7606/j.issn.10004025.2016.04.0834
  %&(:20151205;)*&%+(:20160303
  ,-./:()*+%,-(31270273,31560060)
  0123:.(/(1990-),0,12,3456789:%;<=>?%@A。Email:wjedu@163.com
  BCDE:F G,H2,IJK,12@ALMN,3456789:;<=>?%@A。Email:mengyingnie@163.com
456789:;<=>?@A
!"#

$%&

 (

OPQ% L$RS;TU+%%V#$RSWX;YZ[\]^_
,`
aOP416000)
B C:bcdefgh:ijklmnopq,rs,tuvwxijkyz{|qlmno}g~q€
‚ƒ„…†‡ˆ

‰5lmnqŠ~g‹h~;>?Œx>g9:<Žd‘†ijkuy’q“S
g”x>•–

ijk—˜ij’

™š›’

œij’gžŸ’

 ¡100¢£,¤¥™š›’(x=18)、œ
ij’
(x=16)gžŸ’(x=20)qlmn,o¦§,¨ ij’lmn,o©ª«Q,34¬x=8~16,­wŠ~
®w‹h~¯°

lmnop±²ijk4³’qlmn„´µy¶·¸,¹º†ijkulmn5»,
o¼½,o”q‚ƒ

y’ulmnq”34¾n¯b¿~nÀÁsqÂÀœ

Š~bÄk¥¶Å3M


Çij’uÈw‹ÉÊËq‹h~¯°

78%Ìp;<=>?ŒqˆÍΫÏÐ

¬ijku’|Ñ
ҒÓq>?Œ;„ÔՆ[4qÖ×RØ

DEF

ijk

lmn,o

lmn„

‹h~
GHIJ$
:Q343.2   !KLMN:A
犆狔狋狅犾狅犵犻犮犪犾犃犱狏犪狀犮犲狊狅狀犜狉犻犫犲犘狅犾狔犵狅狀犪狋犲犪犲(犃狊狆犪狉犪犵犪犮犲犪犲)
WANGJiajian,NIEZelong,MENGYing
(KeyLaboratoryofPlantResourcesConservationandUtilization,ColegeofBiologyandEnvironmentalSciences,JishouUniversi
ty,Jishou,Hunan416000,China)
犃犫狊狋狉犪犮狋:BaseonthesurveyofcytologicaldatafromPolygonateae,wereviewedchromosomenumbersand
theirvariationsoftaxafromthetribePolygonateae,includingtheirpolyploidizationandaneuploidizatione
vents,andthecorrelationwithphylogeneticrelationshipsandgeographicdistribution.Thetribeincludes
ca.100speciesfromfourgeneraof犕犪犻犪狀狋犺犲犿狌犿,犇犻狊狆狅狉狅狆狊犻狊,犎犲狋犲狉狅狆狅犾狔犵狅狀犪狋狌犿,and犘狅犾狔犵狅狀犪狋狌犿.
犘狅犾狔犵狅狀犪狋狌犿showsawidevariationofchromosomebasenumbersmainlyfrom8to16 withboth
polyploidizationandaneuploidizationsimultaneously,quitedifferentfromtheotherthreegenerawithstable
basicchromosomalnumbers,i.e.,犕犪犻犪狀狋犺犲犿狌犿 (x=18),犎犲狋犲狉狅狆狅犾狔犵狅狀犪狋狌犿 (x=16),and犇犻狊狆狅狉狅狆狊犻狊
(x=20).Chromosomaldatasupporttherecognitionofthe4generainthetribe,eachshowingadistinct
chromosomalpattern.Wealsofindthatadecreasingtrendofchromosomalbasicnumberoftaxainthe
tribe.Thechromosomalevolutionofthetribeisdominantlyonthediploidylevelwithlesspolyploidization
andtheaneuploidizationeventsareonlyfoundin犘狅犾狔犵狅狀犪狋狌犿.Cytologicaldataarehighlyconsistentwith
thephylogeneticrelationshipsandprovideusefulimplicationsfortheevolutionofthetribe.
犓犲狔狑狅狉犱狊:Polygonateae;chromosomebasenumber;chromosomalevolution;aneuploidization
  ijk(Polygonateae)ْÚÛÜÝ+[13],3
4— ˜ Þ ß ™ š › ’
(犕犪犻犪狀狋犺犲犿狌犿 F.H.
Wigg.)、žŸ’(犇犻狊狆狅狉狅狆狊犻狊Hance)、œij’
(犎犲狋犲狉狅狆狅犾狔犵狅狀犪狋狌犿M.N.Tamura&Ogisu)gi

(犘狅犾狔犵狅狀犪狋狌犿 Mil.), ¡100£。ijkD
¬à³)*qá>z{

®âã†<=>?%Ìp
qäå

æçkq>?Æèébêë

ì?sْÚ
íÐ+
[45],
<=Ìpäåîï5íÐ+<ð

ñò
èÚóô+¥
[67],
õö÷øïùbqóô+úQã
ûüý+
[1,89]
ÑÒ¯bqÛÜÝ+
[23]。
ijkÞ
ŽÚ"þÿ

!"Ãq"#$<Ž

¤¥™š›’g
ij’!¯%&|<Ž´µ

(w)*qL$9
:%@A+,

ijkQŠo£z<ŽÚ-.9/

¥!a9/q01»2g34/¬¤[4q<
Ž¥56à
[1011]。
<=>?Œ@A±²çkB
É<¬¿Q<ä

ÆÚh³ijk,7q™š›’
;—˜ij’

œij’gžŸ’89qàä:
¬;[1216]。
Þßq™š›’—˜ì?q=>’
(犛犿犻犾犪犮犻狀犪
Desf.)g™š›’(犕犪犻犪狀狋犺犲犿狌犿s.str.),¡35
£

-.w20¢£(5?@A、BÄã-CDEF9
/
),
¤¥¡13£f¥<ŽÚ¥!aq34g
CDEF9/

"GHw5£,¤¢£z<ŽÚ¥G
%I$4/
[14,1728]。
ì?=>’g™š›’ÇbJ
,osébKL
[15]。
ì?=>’qJ,o¬M,
o

™š›’qJ,o¬¿,o

™š›’£zb
8Nsw«QKL
[13,2930],
ÑOLqPQJR¬3
4ST

lmno}Š¬2n=36,‰wŠ~qU
V
[31]。
’u£|q/<34WpžXY

JRÒJ
qST

ü8ZST„…/<

æ¾£|x>[w)
Šêë
[1213,2930,3233]。Hara[29]WpJ\]^、_`
aLÆè

Jbcá

Jd8N

îì?=>’
q.%z{<¬4³e。fg[30]WpžXY8N
”qhi

YLüqoj

JRqˆk

J,o
q€œî™š›’<¬2³.’5³e:lXžY


mJe

nJe

oXžYe
)、
7pžY.
’

M,oJeg¿,oJe
)。
ij’Ñ"þÿ#$/q<Ž¬3

¾ijk
ñQq’

 ¡60¢£,34<Žb.%(¥、B
ÄãCDEF9/
),
Ñ¥!a9/g-".¬Š
r<Ž¥5

w6£<Žãs%(2£¬s%S
w
),2£<ŽÚ"G[28,34]。ij’¶Çù—t£z
ñŠ

¤8NsqKLuvwx

SL¾JRqS
T

tÚij’’u£|q[3537]。
BakerWpüRqSTîij’<¬3³e::ü
e
(犃犾狋犲狉狀犻犳狅犾犻犪)、yüe(犞犲狉狋犻犮犻犾犾犪狋犪)gtüe
(犗狆狆狅狊犻狋犻犳狅犾犻犪)[35]。z„WpüR、{^qQ|g
ˆk

JqQ|goj

J}ZSTîij’<¬3
³e 8 ³>::üe (犃犾狋犲犾狅犫犪狋犪、犃犾狋犲狉狀犻犳狅犾犻犪、
犅狉犪犮狋犲犪狋犪、犘狌狀犮狋犪狋犪)、yüe(犎狅狅犽犲狉犻犪狀犪、犓犻狀犵犻
犪狀犪、犞犲狉狋犻犮犻犾犾犪狋犪)、t ü e (犗狆狆狅狊犻狋犻犳狅犾犻犪)[38]。
TamuraŒ¯ij’qlmno}gÂÃ;J~q
ST!€·x

pøij’<9yüe
(Sect.
犞犲狉狋犻犮犻犾犾犪狋犪,x=14、15,J~7)gije(Sect.
犘狅犾狔犵狅狀犪狋狌犿),ijeÓ<93³>:Ser.犘狅犾狔犵狅
狀犪狋狌犿 (x=9,J~s7v‚)、Ser.犅狉犪犮狋犲犪狋犪(x=
10,J~¥7v‚c¶v‚)gSer.犐狀犳犾犪狋犪(x=
11,J~Ó7v‚)[37]。
v

ijk¥[—˜2³£z«ƒq|’:
žŸ’gœij’

žŸ’6£,34<ŽÚ¥
p„ÑaÒ-a.q.I$ÒI$9/
[3839],
¤
<Ž¥5b¥a…†ÑÒp„‡q
[40]。
ž
Ÿ’bˆJ}‰Š7(wIJ。
‹¾ï/LÚi
jku¤ï’q34ST
[11,41]。
œij’6£,¬
¥!a9/Sw

ï¾5ij’<ðqà³|
’

(wbÃ_`

J}^!ŒXŽ

(OL
g‘LJR

Bɬ’L
[4243]。
ijku’|ÑÒy’Ó£|q>?x>g“
”•w«Qqêë

@A±²lmnop¬@Ai
jkq>?x>ÔՆ)*q–—

æwxÄkq
78%dq@A‹É<˜

™š¬›[œwcd
9tijkq78%R؄…efh:g@A

¬
†ž‹dqŸ 

Ä¡>?9efh:†ij
kqùwlmn&¢q¡£

ˆÐñ¤<=>?Œ
@AˆÍ

¥¦§¨ijk’|g’uqlmn
„UV

‰‘lmnopbijku’|q>
?%©ß

¬cd†ªijkÒ¤y’q“Sg”
ÔÕ78%dqÌp

tÚijkq>?Œ
;L$9:@A®(w[4Ö×+,

«vB¬i
jklmn„q@A

w—Ú:ª¶¸#$/>
z{qlmn„‚ƒg$£”€­®
,¯
01»
2°±glmnŠ~qx>

¥!a9/g"
9/lmn„qKœ

1 ijklmno}@A²³
wxlmn&¢q’

£Ò¤lmno}q?
´µ³¶±1g±2。ˆÍ±²:ijk¥4’ 
w75£,323·lmn&¢¸¹。¤¥,™š›’

 35£),20£wlmn&¢, w86·lmn
¸¹

Šo¬2n=36,lmn,o¬x=18,Šo£
z¬¿~n

ƒo¬M~n

º~ng»~n

ž
Ÿ’
(6£),4£wlmn&¢, 11·lmno}


•¬2n=40,lmn,o¬x=20,•¬¿~
n

œij’
(6£),4£wlmn&¢,lmno}
¬2n=32、64,lmn,o¬x=16,¼†犎.狅犵犻狊狌犻
¼~n6v

¤¢•¬¿~n

ij’

 60£),
5384½ .(/,Z:ÛÜÝ+ijk78%@A„¾
47£wlmn&¢,‡ 222·lmn´o,lm
no}€¿ÀQ

,o€“”¬x=8~16,l
mn~Ñ¿~nÁŠ

æ®wM~n

º~n

Â
ÃÄ~nZ~«»q£z

wlmn&¢q#$
QŠ¬ijkq[4z{

¤¥lmn´oۊ
q34¬àÅގ£

Ʊ†Äkùw’qQ7<
ɶ£z

¬@AÄkq>?„x>g“S”
ÔՆ[4Ö×Wp

2 ijklmn„´µ
ijkq)Š£zp½ÑöǾ¥€Èq¥

,¯
=>

ij

ʝZ

(wËÌÍÎ

žÏÐ
O1 89:PQR;<=>?STUVWXYO
Table1 SummaryofcytologicalreportsinPolygonateae
’È
Genus
£o
Species
wlmn
&¢q£o
Chromosome
number
countedin
specieslevel
wlmn
&¢q
¸¹o
Countof
literatures
reported
lmn
,o
Basic
chromosome
number
lmn
~
Chromosome
ploidy
<Ž“”
Distribution
range
™š›’
犕犪犻犪狀狋犺犲犿狌犿 35 20 86 18 2x
、3x、4x、8x
.%-7

?@A!ÑY.

"GÃ¥G%
7、
s%"7
。EastofAsia,Siberia
andNorthofAmerica,NorthofEurope.
žŸ’
犇犻狊狆狅狉狅狆狊犻狊
6 4 11 20 2x
¥Ò-a.qÒÓ

ԃÕ

Ö

×a
China,Laos,Philippines,Thailand,Vietnam
œij’
犎犲狋犲狉狅狆狅犾狔犵狅狀犪狋狌犿
6 4 4 16 2x、4x
¥!a9/SwEndemicinSouthwestof
China
ij’
犘狅犾狔犵狅狀犪狋狌犿
60 47 222 8-16 2x、3x、4x、6x
ÞØ<ŽÚ"#$9/

f¥<ŽbCDE
F9/ÃBÄà$

Ñ¥q!a9/¬Š
r¥5TemperateregionsoftheNorthof
hemisphere,mainlyfromtheHimalayasto
Japan,diversityinSouthwestofChina
O2 89:Z[、\]^_VW
Table2 ChromosomeinformationofthespeciesinPolygonateae
¥¡È
Chinesename
EÙÈ
Latinname
n 2n x
~
Ploidylevel
Öס£
References
»Q=> 犕.犪狋狉狅狆狌狉狆狌狉犲狌犿 36,72 18 2x,4x [31,4445]
™š› 犕.犫犻犳狅犾犻狌犿 18 36,54 18 2x、3x [31,41,4655]
犕.犮犪狀犪犱犲狀狊犲 36 18 [51,5659]
ÚÛ=> 犕.犱犪犺狌狉犻犮狌犿 36 18 2x [31,41,55,6061]
犕.犱犻犾犪狋犪狋狌犿 18 36 18 2x [6266]
ÜÝ=> 犕.犳狅狉犿狅狊犪狀狌犿 36 18 2x
ÞY=> 犕.犳狅狉狉犲狊狋犻犻 36 18 2x [31]
!a=> 犕.犳狌狊犮狌犿 36,54,72 18 2x,3x,4x [44,6771]
nJ=> 犕.犺犲狀狉狔犻 36 18 2x [31,44,72]
犕.犺狅狀犱狅犲狀狊犲 36 2x [73]
犕.犻狀狋犲狉犿犲犱犻狌犿 36 18 2x [74]
=> 犕.犼犪狆狅狀犻犮狌犿 36 18 2x [31,41,45,65,7273,75]
߄=> 犕.犾犻犮犺犻犪狀犵犲狀狊犲 36 18 2x [31]
pà=> 犕.狅犾犲狉犪犮犲狌犿 36 18 2x [67,69,73]
犕.狆犪狀犻犮狌犾犪狋狌犿 36 18 2x [76]
áJ=> 犕.狆狌狉狆狌狉犲狌犿 36 18 2x [31,7778]
犕.狉犪犮犲犿狅狊狌犿 18 36,72,144 18 2x、4x、8x [51,7982]
犕.狊狋犲犾犾犪狋狌犿 36 18 2x [76,8283]
âã=> 犕.狋犪狋狊犻犲狀犲狀狊犲 36 18 2x [31,44]
Mü=> 犕.狋狉犻犳狅犾犻狌犿 36 18 2x [51,76,81,84]
犕.狔犲狊狅犲狀狊犲 36 2x [76,82]
˜äžŸ 犇.犪狊狆犲狉狊犪 40 20 2x [4041]
žŸ 犇.犳狌狊犮狅狆犻犮狋犪 40 20 2x [40,85]
püžŸ 犇.犾狅狀犵犻犳狅犾犻犪 40 20 2x [40,86]
å]žŸ 犇.狆犲狉狀狔犻 40 20 2x [4041,87]
638 ! " # $ % & 36æ
ç±2 ContinuedTable2
¥¡È
Chinesename
EÙÈ
Latinname
n 2n x
~
Ploidylevel
Öס£
References
犎.狅犵犻狊狌犻 64 16 4x
èYœij 犎.狆犲狀犱狌犾狌犿 32 16 2x [42]
œij 犎.狉狅狊犲狅犾狌犿 32 16 2x [42]
ºéœij 犎.狓狌犻 32 16 2x
aä>

yüz
)Southernlineage(verticilate)
:æij 犘.犪犾狋犲狉狀犻犮犻狉狉犺狅狊狌犿 32 16 2x [88]
o~ij 犘.犮犪狋犺犮犪狉狋犻犻 32 16 2x [44]
æüij 犘.犮犻狉狉犺犻犳狅犾犻狌犿 15 30,52 13,14,15 2x,4x [37,44,67,8890]
èüij 犘.犮狌狉狏犻狊狋狔犾狌犿 28,30,78 14,15,18 2x,4x [44,91]
ê>ij 犘.犳狉犪狀犮犺犲狋犻犻 26 13 2x [9192]
犘.犵犲犿犻狀犻犳犾狅狉狌犿 30 15 2x [77,9394]
Mëij 犘.犵狉犻犳犳犻狋犺犻犻 36 18 2x [71]
‚ìij 犘.犺犻狉狋犲犾犾狌犿 28 14 2x,4x [95]
íJij 犘.犺狅狅犽犲狉犻 30 15 2x [96]
îij 犘.犽犻狀犵犻犪狀狌犿 26,30,64 13,15,16 2x,4x [37,44,88,91,9697]
tüij 犘.狅狆狆狅狊犻狋犻犳狅犾犻狌犿 15 30 15 2x [37,7071,98]
ï§Ê 狆.狆狉犪狋狋犻犻 28 14 2x,4x [37,99]
\Jij 犘.狆狌狀犮狋犪狋狌犿 30,90 15 2x,4x [37,88,99]
¤ðij 犘.狉狅狊犲狌犿 28 14 2x [96]
ñüij 犘.狊狋犲狀狅狆犺狔犾犾狌犿 30 15 2x [6566,99]
!aij 犘.狊狋犲狑犪狉狋犻犪狀狌犿 28 14 2x [37]
òëij 犘.狋犲狊狊犲犾犾犪狋狌犿 30,60 15 2x,4x [37,95]
yüij 犘.狏犲狉狋犻犮犻犾犾犪狋狌犿 15 28,30,60 14,15 2x,4x [37,70,77,93,99102]
`"ij 犘.狕犪狀犾犪狀狊犮犻犪狀犲狀狊犲 28,30 14,15 2x [88,9192,103105]
"ä>

:üz
)Northernlineage(alternate)
óôij 犘.犪犱狀犪狋狌犿 22 11 2x [95,106]
犘.犫犻犳犾狅狉狌犿 20 10 2x [107]
ŠJij 犘.犮狔狉狋狅狀犲犿犪 18,20,22,24 9,10,11,12 2x [88,90,102,104,108111]
p{ij 犘.犱犲狊狅狌犾犪狏狔犻 18,20,22 9,10,11 2x [6566,102,109]
犘.犱狅犿狅狀犲狀狊犲 19(xõn Hybrid) 2x [109]
犘.犳犪犾犮犪狋狌犿 18 9 2x [65,109]
pôij 犘.犳犻犾犻狆犲狊 16,18 8,9 2x [90,102,104,108]
犘.犵犾犪犫犲狉狉犻犿狌犿 20 10 2x [112113]
犘.犵狉犪狀犱犻犮犪狌犾犲 18 9 2x [65]
|ʝ 犘.犺狌犿犻犾犲 20,30 10 2x,3x [41,65,90,109,114119]
ì‰Ê 犘.犻狀犳犾犪狋狌犿 22 11 2x [65,99,109,119]
犘.犻狀犳狌狀犱犻犳犾狅狉狌犿 18 9 2x [65]
¿{ij 犘.犻狀狏狅犾狌犮狉犪狋狌犿 18 9 2x [24,65,90,109,118]
犘.犽犻狅狋犲狀狊犲 18 9 2x [120]
犘.犾犪狊犻犪狀狋犺狌犿 20,30 10,15 2x [65,109,119,121]
犘.犾犪狋犻犳狅犾犻狌犿 20 10 2x [122]
Iöij 犘.犿犪犮狉狅狆狅犱狌犿 22 11 2x [90]
犘.犿犪犮狉犪狀狋犺狌犿 22 11 2x [109]
Q{ij 犘.犿犲犵犪狆犺狔犾犾狌犿 22 11 2x [37,99,123]
犘.犿狌犾狋犻犳犾狅狉狌犿 18,22,36,40 9,10,11 2x,4x [37,93,124130]
ʝ 犘.狅犱狅狉犪狋狌犿 18,20,30,40 9,10 2x,3x,4x
[24,37,41,88,90,101103,
109,114119,123125,131146]
÷øij 犘.狅犿犲犻犲狀狊犲 22 11 2x [63]
犘.狅狉犻犲狀狋犪犾犲 18,27 9 2x,3x [147149]
犘.狆狌犫犲狊犮犲狀狊 10 10 2x [150]
犘.狉狅犫狌狊狋狌犿 20 10 2x [65]
ij 犘.狊犻犫犻狉犻犮狌犿 24 12 2x [65,78,88,90,104,129,151]
7384½ .(/,Z:ÛÜÝ+ijk78%@A„¾
ùZúû
[151152]。
¥%E¶Çb¤>Zdü†
à§q@A

[b¤>?qýD

b8Ng78%@Ad

ijky’|
g’u£|q>?x>àþéb)Qqêë

78
%@AÿѬ:ªijku$£q“Sg”ÔÕ
‹É[4qÖ×+,

lmno}q”g$£„
qx>bL$„d~!x"
[153155],
—˜Š
~g‹h~b£lmn”­®

Š~}#
¬¾#$”g89¬$q34­®

%ŠÌp±
²ùwqwJ#$&ébŠ~“S
[154,156158]。
Š
~ÿÑÊ#$tTU()q*+,-

ί.
/

01

23

ÿÑÅpuÞqLNÆ
[159161]。
¼
†Š~6v

#$b”¬$¥[ð¯†lmn
q]c4Ðövc56à·lmn

7‹h~

[162163]。
bijky’¥

lmno}q€¿
À‹ÉQ

¼†Ñ¿~n¬3nqÂÀ6v

[
È8wŠ~Ñҋh~q¯°ð¯

2.1 `ab[、cde[fg89[U]^_hi
™š›’35£¥wlmn&¢qw20£,‹
Å&¢QŠö)-.g"Gq$£

tÚ<ŽÚ¥
G%I$4/qz{blmnd9¬Ÿ 

9:
;<4„à¦q@A

™š›’qlmn&¢lm
n,og~•«¬¦§
。Tahara[75]ñ=t™š
›’q犕.犼犪狆狅狀犻犮狌犿„…lmn&¢,Œ¯lmn
o}¬2n=36。>?@Z[164]#¬™š›’qlm
n2A,o¾x=9,¨ x=18¾B¬x=9BL¨
ö

æCڙš¬›œwŒ¯w2n=18q&¢,Ñ
Ò×DãĒqEˆ’¯œijgžŸ’Z&¾
»lmn,o

ùÑFG#¬™š›’qlmn,
o+ç¬x=18,QŠo£z¬¿~n2n=36。«
v

™š›’lmnbpÀsKœ«Q
,MengZ[10]
5lmnqpÀsø™š›’<¬àÃ

bÃgM
Ãlmn

#¬ç’qlmnbÂÏdq”¾
5àÃbÃMÃq¼„q。¤H,™š›’
¥wÅlmn[(w8n

bŠ~d

¼†»Q=>

!a=>ð¯³
LqM~n

º~nÁ{

ÑÒ犕.狉犪犮犲犿狅狊狌犿 ð¯
¿~n

º~ng»~nÁ{Ñv

¤ï£¬¿~
n

w&¢#¬Š~wYÚIJ/K

23

LM
Z)*·N

ÅpuÞqLNÆ

»Q=>g!a
=>34<ŽÚ¥!a9/

¤~q€ÿ,
¾*+ç/qwxŠrq989OqˆÍ
。犕.
狉犪犮犲犿狅狊狌犿¬"GÞØ<Žqz{,¿~n、M~
n

º~n

Âû~n&wð¯bç£q"GÁ{
¥

‹ÿ,g"G-!7/qqùP

9:TUKœ
ÑÒ¤QRSTqSwx

UsùV

™š›’
qlmn„34±¯b¿~nÀÁsqpÀ€
œ

‰ÈwƒoqŠ~6N

žŸ’Ç—t6£,¤¥wlmnRØq¬
4£,lmn,o¬x=20,lmno}•¬2n=
40,•¬¿~n,wqlmns[wHWX,lmn
bpÀsqKL¶²

’àÃlmn

ælmnq
YtpÀ²pÚijku¤ï’

œij’Çw6£,•¬¥!aSw,Š’L
ÚZ[cýs

ç’lmno}&¢w4£,2n=
32c64,lmn,o¦§(x=16),QŠ¬¿~n,
Çb<ŽÚºé\4q犎.狅犵犻狊狌犻Œ¯wº~n;
lmnbpÀs!²qbÃ

BÉw5tpl
mng11t]lmne9。
2.2 89[U]^_jiklmnXopUqr
Dn
  ij’¾ijkñQq’,lmno}„‹
Éwx

lmno}€©ª«Q

¯wop±²
璶Çb£uq¶¸Á{|wlmno}€

wqÂÃb¸àÁ{u®wKœ

‰È8wŠ~
ÑÒ^ç‹h~q¯°



ÞØ<ŽÚs.
Q`#$9/qʝ犘.狅犱狅狉犪狋狌犿 (Mil.)Druce,
ö)s%

.%-"7ÑÒ¥!"Òa"9/q
&¢Š¬2n=20[101,114116,118,123124,130131,165],¨ ö)
a-

a¥g!a9/qÁ{lmno}52n=16
g18,bc&¢ÛdQL4qÉج2n=18,¨
b„efڌ¯w3£78Ã2n=18、20g2n=
22[139]。g@hZ[102]®Œ¯Ûdij4ÉØ(2n=
18)gi4ÉØ(2n=16)¬b£78Ã。ʝ­¾
"þÿ#$ގ£

÷¾ijkkdlmn&¢ñ
Šq£z

¶¸%Et¤lmn”g“S&ð¯
†)Šl。
bc
[139]
btʝ8³9/q9³
É؄…†ÂÃ
n#¬oÀ40¾Êl
mno}Kœq
b"o40ÑsʝÑ2n
=20<Ž¬3,—˜ÞQqs.Q`#$9/,¯


rZ

Âë¬2A

"o40Ñs/qÿ,
¾Êq“S¥5

boÀ40ÑÓ/q,¯ ¥q
a-

!a

ʝlmno}b2n=20qsÓ(18
g22)©ª,ÂÃ𯄁UV,boÀ40ÑÓ/
qÿ,¾Êq<¥5
。Park,ÚüsnR
gÂR
s%9/qʝg.t-"9/q
ʝ¬u
‰v;,Úlmn,o
(x=9g
x=10)<9b³<äqˆÍ¾àwq[144]。Park
838 ! " # $ % & 36æ
Z[144]
§¨¤ÿ,¾x¬.sQ`ìyõ

p½}
9:z{

*õb.%-"89¤q€£

#¬lm
nq‹h~bij’ÑÒʝq89<¬$¥
(w[4DZ

ij’lmno}g,oq€

¬Ä’q>
??x>q@AÔՆ‹É*qÖ×g–
—

ίTamura,Úij’qlmno}gÂÃ
;J~qST

îij’<9yüe
(Sect.犞犲狉狋犻
犮犻犾犾犪狋犪)(x=14、15,J~7)gije(Sect.犘狅
犾狔犵狅狀犪狋狌犿),ijeÓ<93³>:Ser.犘狅犾狔犵狅狀犪
狋狌犿 (x=9,J~s7v‚)、Ser.犅狉犪犮狋犲犪狋犪(x=10,
J~¥7v‚c¶v‚

gSer.犐狀犳犾犪狋犪(x=11,J
~Ó7v‚
)[37]。ZhaoZ[106],Úlmn,o、Â
Ã

üRZSTg<Ž/qî7¬MQz{
:x=8、9、10、11(:Lü,<ŽÚ¥Q
7,
¼†01»29/6v
),x=12(yL
ü

<ŽÚ¥-"

a"

a¥ga-9/
),x=
13、14、15、16(yLü,<ŽÚ01»29/)。ñ
¤q<=>?@A®äåij’34<¬MQä

à
ä¬34<Žbaqyüz{

lmn,o34¬
13~16;«àä¬34<Žb"q:üz{,lm
n,o34¬8~11;lmn,o¬x=12[11]q
犘.狊犻犫犻狉犻犮狌犿>?Æè|Ú}dbQz{6|,~ø
ïD¬¤e
(sect.犛犻犫犻狉犻犮狌犿)ö:,€n58Nsö
ï’Ú"Ãqayüz{

æ5>?x>g9:
<Žsö

ï’Ú"ä>

ö)lmn,o®¸
räåï|Úaz{g"z{6|

2.3 89:UstuvwfIx
ijkqlmn„34¬¿~nÀÁqÂÃ
¶t‚ÒpÀ€œ

Š~¯°¶Š
[166167]。
b
œij’gžŸ’¥

QŠo¬¿~n

™š›’
7<£zð¯M~n

º~n

»~n

ij’qŠ
~d

𯆊³º~nÒÄ~nz{

™š
›’¥Š~q£z®¶Š

Hw»Q=>

™š
›

!a=>
、犕.狉犪犮犲犿狅狊狌犿 wŠ~n¯°。bœ
ij’¥Œ¯Çw犎.狅犵犻狊狌犻¬º~n,¤¢•¬¿
~n

bij’¥

Š~¯°®¶Š¶

Q²w
14£ébŠ~¯°,¯ |ʝ、犘.犿狌犾狋犻犳犾狅狉狌犿、
ʝ

ij

èüij

‚ìij

îij

ï§Ê

\Jij

òëij

yüijZ

æ&¬¿~n;
Š~nqwÐn

[œwŒ¯áí¬Š~nq£z

ʝ¾ijkk@Añ¬ÞØq

¾cÿ"#$Þ
Ž£

𯆊³~2x、3x、4x,ÿ,¾bp½q
”¬$¥

;TU·:DZù89q

Š~¶
Çbijkqގ£¥±¯â)²

‰v;9q
<Ž®ébà§qxƒ

_¯<ŽÚCDEF4ë
Ò¤Eˆ/q
(¯
„À"7

…a

!1Z9

qij
k£z

·«Ú¥¤n9/

Š~q¯°u¬†
S

b¸à£z6|

<ŽÚ„À9/qyüij
ð¯¿~ngº~n
,¨
¥Ûd9/qyüij
¬¿~n

<ŽÚ¥Q7<9/qij¬¿~n

æb„À9/‡¬M~n

bijkq¶¸’6
|

®±¯ðzˆqUV

<ŽÚ¥!a9/ÒC
DEF-79/q£zŠ~n𯆉•»Ú¤n
9/

Cÿ¶

CDEFŠ‹q9ŒªÑÒ»
Žq‘ùP·Nÿ,’„†ijk#$qŠ~
„$

2.4 89:UyztuvwfIx
bijkkd

wà³Î«²qS\Ǿ‹
h~¯°Î«“ð

‹34±¯bij’

ij
’u‹h~±¯bb³”H

Pò¾bh³ij
’¶¸£6|

éb²qà³lmn,o€q
•À

58–Aàþã16;«àd¾b%Š£z
u7®éb£u,oq€

ߊJij
(x=9、
10、11、12)、p{ij(x=9、10、11)、pôij(x=
8、9)、ʝ(x=9、10),æüij(x=13、14、15)、è
üij
(x=14、15、18)、îij(x=13、15、16),&
¾(w²q‹h~qƱ



‹h~ÿ
,¾Äkblmn„dqà£Î«íSq‚
ƒ

®—˜†ij’#$lmnΫ™š›6


c]qS。
«vxõ®ÿ,¾MwÄk%Š#
$lmn‹h~qà³[4~œ

ί犘.犱狅
犿狅狀犲狀狊犲lmno}¬2n=19,ÿ,¾B¬犘.犻狀
狏狅犾狌犮狉犪狋狌犿 (2n=18);犘.犾犪狊犻犪狀狋犺狌犿(2n=20)x
õ¨ö

zˆq¯°bij’¥®ÿ,ð¯

~ÿ
,®¾ij’q‹h~qà³[4~œ

2.5 89:U]^_,{mnX@uDn
bijkqlmn,o;>?x>d

ij
’qlmno}€g<=>?Œ·tàw

<
=>?Ìp±²Äk34CbQäe9

àä¾C
Þßq™š›’e9

,lmn,o¦§¬x=18;
«àäC¤¢3³’e9,¤¥2³|’žŸ
’
(x=20)gœij’(x=16)ž9àä;ij’
(x=8~16):¬uãij’éb,oŸ q‚ƒ

‹£‚ƒ±¯bi
j’u7®‹É²

5aqyüz{ã"q
:üz{

lmn,o5Ñ13~16¬3ãÑ9~11
¬3
[11],
ί"<äqij,oñ½
(x=12),ü
9384½ .(/,Z:ÛÜÝ+ijk78%@A„¾
R¬yLü

ÿ,¾"<ä¼a¬¡qà³z
{

«v

lmnop¬<=>?x>ÔՆþ¢
qÌp

ί\Jij
(x=15)gï§Ê(x=14)
•Ñ:Lü¬3

ælmn,oÌp;<=Ìp»
ÀÏÐÊËäåøïG£¤ayüàä

bij
’kCÚüR”qÿ¥

¶,¦§qD¬qWp

ælmnopÔՆ‹Éÿ¦qäå

2.6 89:U|}I~f]^_u
ijkq9:<Žglmn”®ébà§q
‚ƒ

žŸ’gœij’ÑÒij’qaz
{

34<Žb¥ag!a9/

—˜34g
01»2ÑÒCDEF9/
),
lmn,o&Ϋ
»

æbïG¼"§¨õ

ίij’q"ä
>

lmn,o©ªÓ«

4ÐcE›6q¯°‹É
¬­

b$£ÀÁs®w²q9qST

ίÊ
boÀ40ÑsgÑÓ/q&¶¸,®"/q

a"

-"

lmno}¦§

®a/q

a-

!
a

lmno}b2n=20sÓ©ª,¨ vÂÃ[ð
¯„UV

ùÑa/q

oÀ40ÑÓ)ÿ,¬Ê
q<¥5

÷¯ŠJij5!aÁ{qt‚Â
Ããa¥

aa

a-9/q¶t‚ÂÃ

Ñt‚ã
¶t‚ÂÃq”¼¬Wp

ÿѧ¨!a9/
¬¤“S¥5

3 ‡ ˆ
ijku’|ÑÒ£|q>?x>g“”&w
‹ÉQqêë

lmnop¬@Aijkq>?x
>ÔՆ‹É*q–—

B¬cdefg>?h:
ijkqùwlmno}&¢q¡£

ˆÍ±²

(1)ijkqlmn„34¬¿~nÀÁlmn
bpÀsq€œ

Š~¯°¶Š
;(2)ijkq‹
h~¯°Î«“ð

‹h~ÿ,¾Äkblm
n„dqà£Î«íSq‚ƒ

®—˜†çk
lmnΫ™š›6

4Ðc]qS;(3)ij
kqlmn,og>?Œd

ij’qlmn
o}€g<=>?Œ·tàw

<=>?Ìp
±²Äk34CbQäe9

àä¾CÞßq™š
›’e9

,lmn,o¦§¬x=18,«àäC
¤¢3³’e9,¤¥2³|’žŸ’(x=20)g
œij’
(x=16)ž9àä;ij’(x=8~16):
¬u
ˆÐñ¤<=>?Œ@AqˆÍ

¥
¦§¨ijk’|g’uqlmn€UV

‰
‘ijkÒ¤’|q>?%©ß

¬cd†ªij
kÒ¤y’q“Sg”ÔÕ78%dqÌp

‹tÚijkq>?Œ;L$9:@A(w[4
Ö×+,
;(4)bijkq9:<Žglmn”
d

b¥ag!a9/

—˜34g01»2
ÑÒCDEF9/
),
lmn,o&Ϋ»

æbï
G¼"§¨õ

ίij’q"ä>

lmn,
o©ªÓ«

4ÐcE›6q¯°‹É¬­


ijklmn”¥ù±¯ðq¶¸9/|qK
œ

w—Ú:ª¶¸#$/>z{qlmn„‚
ƒg$£”€­®
,¯
01»2°±glmnŠ~
qx>

æ¾ijkuqlmno}€‹Éwx

ï
Gq¯òlmno}glmn”­®•–[¶,
ª°

ÿ,<4ˆÐ<=>?ŒdqˆÍ

„à
¦Zˆ*±„´Ã)w²³ª°

€!K

[1] RUDALLPJ,CONRANJG,CHASE M W.Systematicsof
Ruscaceae/Convalariaceae:acombinedmorphologicalandmo
lecularinvestigation[J].犅狅狋犪狀犻犮犪犾犑狅狌狉狀犪犾狅犳狋犺犲犔犻狀狀犲犪狀犛狅
犮犻犲狋狔,2000,134(12):7392.
[2] BREMERB,BREMERK,CHASEM,犲狋犪犾.Aupdateofthe
AngiospermPhylogenyGroupclassificationfortheordersand
familiesoffloweringplants:APGⅢ[J].犅狅狋犪狀犻犮犪犾犑狅狌狉狀犪犾狅犳
狋犺犲犔犻狀狀犲犪狀犛狅犮犻犲狋狔,2009,161(2):105121.
[3] KIMJH,KIMDK,FORESTF,犲狋犪犾.Molecularphylogenet
icsofRuscaceaesensulatoandrelatedfamilies(Asparagales)
basedonplastidandnuclearDNAsequences[J].犃狀狀犪犾狊狅犳
犅狅狋犪狀狔,2010,106(5):775790.
[4] BENTHAMG,HOOKERDJ.Genera犘犾犪狀狋犪狉狌犿[M].Londi
ni:ReeveandCo..1883:748837.
[5] KRAUSEK.Liliaceae[J].犇犻犲狀犪狋ü狉犾犻犮犺犲狀狆犳犾犪狀狕犲狀犳犪犿犻犾犻犲狀,
1930,15:227386.
[6] DAHLGRENR M T,CLIFFORDHT,YEOPF.Evolution
withinthe monocotyledons[M]// DAHLGREN R M T,
CLIFFORDHT,YEOPF.TheFamiliesofMonocotyledons.
Burger:Springer,1985:91106.
[7] TAKHTAJANA.Systemamagnoliophytorum[M].Moscow:
Nauka.1987:1440.
[8] CHASEM W,DEBRUIJNAY,犲狋犪犾.PhylogeneticsofAs
phodelaceae(Asparagales):ananalysisofplastidrbcLand
trnLFDNAsequences[J].犃狀狀犪犾狊狅犳犅狅狋犪狀狔,2000,86(5):
935951.
[9] FAYM,RUDALLP,SULLIVANS,犲狋犪犾.Phylogeneticstud
iesofAsparagalesbasedonfourplastidDNAregions[C]//
048 ! " # $ % & 36æ
WILSONKL,MORRISONDA.犕狅狀狅犮狅狋狊:狊狔狊狋犲犿犪狋犻犮狊犪狀犱犲
狏狅犾狌狋犻狅狀.犞犻犮狋狅狉犻犪:犆犛犐犚犗,2000:360371.
[10] MENGY,WENJ,NIEZL,犲狋犪犾.Phylogenyandbiogeographic
diversificationof犕犪犻犪狀狋犺犲犿狌犿 (Ruscaceae:Polygonatae)[J].
犕狅犾犲犮狌犾犪狉犘犺狔犾狅犵犲狀犲狋犻犮犪狀犱犈狏狅犾狌狋犻狅狀,2008,49(2):424434.
[11] MENGY,NIEZL,DENGT,犲狋犪犾.Phylogeneticsandevolu
tionofphylotaxyintheSolomon'ssealgenus犘狅犾狔犵狅狀犪狋狌犿
(Asparagaceae:Polygonateae)[J].犅狅狋犪狀犻犮犪犾犑狅狌狉狀犪犾狅犳狋犺犲
犔犻狀狀犲犪狀犛狅犮犻犲狋狔,2014,176(4):435451.
[12] LAFRANKIEJV.TransferofthespeciesofSmilacinato
犕犪犻犪狀狋犺犲犿狌犿(Liliaceae)[J].犜犪狓狅狀,1986a,3:584589.
[13] LAFRANKIEJV.MorphologyandtaxonomyoftheNew
Worldspeciesof犕犪犻犪狀狋犺犲犿狌犿 (Liliaceae)[J].犑狅狌狉狀犪犾狅犳
狋犺犲犃狉狀狅犾犱犃狉犫狅狉犲狋狌犿,1986b,67(4):371439.
[14] LAFRANKIEJV.Anewspeciesof犕犪犻犪狀狋犺犲犿狌犿 (Liliace
ae)fromCostaRicawithanuprightandaerialrhizome[J].
犃犿犲狉犻犮犪狀犑狅狌狉狀犪犾狅犳犅狅狋犪狀狔,1986c,9:12581260.
[15] TAMURAMN,SCHWARZBACHAE,KRUSES,犲狋犪犾.Bi
osystematicstudiesonthegenus犘狅犾狔犵狅狀犪狋狌犿 (Convalari
aceae)IV.Molecularphylogeneticanalysisbasedonrestric
tionsitemappingofthechloroplastgene狋狉狀犓[J].犉犲犱犱犲狊
犚犲狆犲狉狋狅狉犻狌犿,1997b,108(34):159168.
[16] JANGCG,PFOSSER M.PhylogeneticsofRuscaceaesensu
latobasedonplastid狉犫犮犔and狋狉狀犔犉 DNAsequences[J].
犛狋犪狆犳犻犪,2002,80:333348.
[17] f g,´  µ.™š›’ण[J].…a#$@A,1983,5
(1):7778.
LIH,CHENY.Anewspeciesofthegenus犛犿犻犾犪犮犻狀犪[J].
犃犮狋犪犅狅狋犪狀犻犮犪犢狌狀狀犪狀犻犮犪,1983,5(1):7778.
[18] LIH,HUANGJL.Anewspeciesof犕犪犻犪狀狋犺犲犿狌犿:M.nan
chuanense[J].犅狌犾犾犲狋犻狀犅狅狋犪狀狔犚犲狊犲犪狉犮犺.犖狅狉狋犺犈犪狊狋犉狅狉犲狊狋.
犐狀狊狋.,1990,10:5153.
[19] GARCAARVALOA.犕犪犻犪狀狋犺犲犿狌犿犿犲狓犻犮犪狀狌犿 (Liliace
ae)unanuevaespeciedeDurango,Mexico[J].犃犮狋犪犅狅狋á狀犻犮犪
犕犲狓犻犮犪狀犪,1992,17:1921.
[20] LEEY.NewtaxaoftheKoreanflora:5[J].犓狅狉犲犪狀犑狅狌狉狀犪犾
狅犳犘犾犪狀狋犜犪狓狅狀狅犿狔,1993,23(4):263268.
[21] NOLTIEH.NotesrelatingtotheFloraofBhutan:XXII.As
paragaceae:AsparagusfilicinusandConvalariaceae:Maian
themumoleraceum[J].犈犱犻狀犫狌狉犵犺犑狅狌狉狀犪犾狅犳犅狅狋犪狀狔,1993,
50(2):207210.
[22] LIANGSY.Anewcombinationofthegenus犛犿犻犾犪犮犻狀犪of
Liliaceae[J].ActaPhytotaxonomicaSinica,1995,33:229.
[23] TAKAHASHIH.Classifactionanddistributionof犛犿犻犾犪犮犻狀犪
狊狆犲犮犻犲狊(Liliaceae)inJapan(inJapanese)[J].犅狌犾犾犲狋犻狀犖犪
狋犻狅狀犪犾犛犮犻犲狀犮犲犕狌狊犲狌犿犜狅犽狔狅犛犲狉犻犲狊犅,1997,23:103114.
[24] KIMJY,LEENS.AtaxonomicstudyofKoreanSmilacina
[J].犑狅狌狉狀犪犾狅犳犘犾犪狀狋犅犻狅犾狅犵狔,1998,41(1):5058.
[25] CHENXQ,KAWANOS.MaianthemumFH Wiggers[M]//
WUZY,RAVENPH.FloraofChina.BejingandSt.Louis:
SciencePressandMissouriBotanicalGardenPress,2000,24:
219223.
[26] CHENSC,KAWANOS.Twonewcombinationsin犕犪犻犪狀
狋犺犲犿狌犿 (Convalariaceae)[J].犖狅狏狅狀,2000,10(2):113114.
[27] LIR,LIH.Twonewtaxaof犕犪犻犪狀狋犺犲犿狌犿 (Convalariace
ae)fromnorthwesternYunnan,China[J].犖狅狏狅狀,2002,12:
486491.
[28] JUDDWS.ThegeneraofRuscaceaeinthesoutheasternUnited
States.[J].犎犪狉狏犪狉犱犘犪狆犲狉狊犻狀犅狅狋犪狀狔,2003,7(2):93149.
[29] HARAH.NotestowardsarevisionoftheAsiaticspeciesof
thegenus犛犿犻犾犪犮犻狀犪[J].犑犉犪犮犛犮犻犝狀犻狏犜狅犽狔狅,犛犲犮狋.犅狅狋.,
1987,14:137159.
[30] f g.=>’q?[J].…a#$@A,1990,(¶
Ⅲ):112.
LIH.Infragenericsystemofthegenus犕犪犻犪狀狋犺犲犿狌犿[J].犃犮
狋犪犅狅狋犪狀犻犮犪犢狌狀狀犪狀犻犮犪,1990,(SⅢ):112.
[31] MENGY,NIEZL,YANGYP,犲狋犪犾.Karyomorphologyof
犕犪犻犪狀狋犺犲犿狌犿狊犲狀狊狌犾犪狋狅 (Polygonatae,Ruscaceae)[J].
犑狅狌狉狀犪犾狅犳犘犾犪狀狋犚犲狊犲犪狉犮犺,2005,118(2):155162.
[32] EMONSRW.ArevisionoftheCentralAmericanspeciesof
Smilacina[J].犃狀狀犪犾狊狅犳狋犺犲 犕犻狊狊狅狌狉犻犅狅狋犪狀犻犮犪犾犌犪狉犱犲狀,
1945,32(4):395411.
[33] GALWAYDH.TheNorthAmericanspeciesof犛犿犻犾犪犮犻狀犪
[J].犃犿犲狉犻犮犪狀犕犻犱犾犪狀犱犖犪狋狌狉犪犾犻狊狋,1945,33:644666.
[34] CHENX,TAMURA MN.犘狅犾狔犵狅狀犪狋狌犿 Mil[M]//WUZY,
RAVENPH.FloraofChina.BejingandSt.Louis:SciencePress
andMissouriBotanicalGardenPress,2000,24:223232.
[35] BAKERJ.Polygonatum,revisionofthespeciesandgeneraof
Asparagaceae[J].犅犻狅犾狅犵犻犮犪犾犑狅狌狉狀犪犾狅犳狋犺犲犔犻狀狀犲犪狀犛狅犮犻犲
狋狔,1875,15:552561.
[36] ABRAMOVAL.Onthetaxonomicalstructureofthegenus
犘狅犾狔犵狅狀犪狋狌犿 Mil.[J].犅狅狋犪狀犻ˇ犮犲狊犽犻犼犣犺狌狉狀犪犾 (犕狅狊犮狅狑 牔
犔犲狀犻狀犵狉犪犱,1975,60(4):490497.
[37] TAMURAMN.Biosystematicstudiesonthegenus犘狅犾狔犵狅
狀犪狋狌犿 (Liliaceae)III.Morphologyofstaminalfilamentsand
karyologyofelevenEurasianspecies[J].犅狅狋.犑犪犺狉犫.犛狔狊狋.,
1993,115(1):126.
[38] z „.ij’[M]//·Œ¸,z „.¥#$?."¹:+
%ðº»
,1978,15:5580.
[39] LIUZ.Anewspeciesof犇犻狊狆狅狉狅狆狊犻狊(Liliaceae)fromSichuan
[J].犃犮狋犪犘犺狔狋狅狋犪狓狅狀狅犿犻犮犪犛犻狀犻犮犪,1987,25(1):6768.
[40] GUZJ,YANGQE,KONDOK.Akaryomorphologicalstudy
on犇犻狊狆狅狉狅狆狊犻狊HanceinChina[J].犔犪犓狉狅犿狅狊狅犿狅犐犐,1990,
57:19161925.
[41] ¼½¾,¿À….ijk4’6£qÂÃ&¢[J].#$&
,1990,28(3):185198.
HONGDY,ZHUXY.Reportonkaryotypesof6speciesin
4genusofPolygonateaefromChina[J].犃犮狋犪犘犺狔狋狅狋犪狓狅狀狅狀
犿犻犮犪犛犻狀犻犮犪,1990,28(3):185198.
[42] TAMURAM N,OGISU M,XUJM.犎犲狋犲狉狅狆狅犾狔犵狅狀犪狋狌犿,a
newgenusofthetribePolygonateae(Convalariaceae)from
westChina[J].犓犲狑犅狌犾犾犲狋犻狀,1997a,52(4):949956.
[43] CHAOCT,TSENGYH,TZENGHY.犎犲狋犲狉狅狆狅犾狔犵狅狀犪狋狌犿
犪犾狋犲犾狅犫犪狋狌犿(Asparagaceae),comb.nova[J].犃狀狀犪犾犲狊犅狅狋犪狀犻
犮犻犉犲狀狀犻犮犻,2013,50(12):9194.
[44] . ß,>?@,Á Â,Z.íÐ+ĒÃÄ£#$q78
%@A
[J].#$WANGL,GUZJ,GONGX,犲狋犪犾.Acytologicalstudyoffif
1484½ .(/,Z:ÛÜÝ+ijk78%@A„¾
teenspeciesinsixgenusofLiliaceaefromYunnan[J].犃犮狋犪
犘犺狔狋狅狋犪狓狅狀狅狀犿犻犮犪犛犻狀犻犮犪,1993,31(6):549560.
[45] TAGASHIRAN,KAMANOIM,犲狋犪犾.Karyotypestabilityof
犛犿犻犾犪犮犻狀犪犼犪狆狅狀犻犮犪[J].犆犺狉狅犿狅狊狅犿犲犛犮犻犲狀犮犲,1999,3(2):
7982.
[46] MOORERJ.Indextoplantchromosomenumbersfor1969
[J].犚犲犵狀狌犿犞犲犵犲狋犪犫犻犾犲,1971,77:17.
[47] GOLDBLATTP.Indextoplantchromosomenumbers:1982
1983[J].犕狅狀狅犵狉.犛狔狊狋.犅狅狋.,1985,13:1224.
[48] GOLDBLATTP.Indextoplantchromosomenumbers:1975
1978[J].犕狅狀狅犵狉.犛狔狊狋.犅狅狋.,1981,5:1553.
[49] GOLDBLATTP.Indextoplantchromosomenumbers:1979
1981[J].犕狅狀狅犵狉.犛狔狊狋.犅狅狋.,1984,8:1447.
[50] DMITRIEVAS,PARFENOVV.Kariologicheskajakharak
teristikanekotorykhvidovpoleznykhrastenijfloryBelorussi
[J].犐狕狏犲狊狋犻犻犪犃犽犪犱犲犿犻犻犖犪狌犽犅犲犾狅狉狌狊狊犽狅犻犛犛犚:犛犲狉犻犻犪犅犻狅
犾狅犵犻犮犺犲狊犽犻犽犺犖犪狌犽,1985,6:38.
[51] THERMANE.CytotaxonomyofthetribePolygonatae[J].
犃犿犲狉犻犮犪狀犑狅狌狉狀犪犾狅犳犅狅狋犪狀狔,1956,43(2):134142.
[52] KROGULEVICHRE.KariologicheskijanalizvidovfloryVos
tochnogoSajanaV.[M]//MALYSHEVLI,PESHLCOVA G
A.犉犾狅狉犪犘狉犻犫犪犼犽犪犾'犼犪.犖狅狏狅狊犻犫犻狉狊犽:犖犪狌犽犪,1978:1948.
[53] LAANE M M,LIET.Fremstilingavkromosompreparater
medenklemetoder[J].犅犾狔狋狋犻犪,1985:715.
[54] IETSWAARTJH,SCHOORLJW.FructificationinDutch
犕犪犻犪狀狋犺犲犿狌犿犫犻犳狅犾犻狌犿populations[J].犃犮狋犪犅狅狋犪狀犻犮犪犖犲犲狉
犾犪狀犱犻犮犪,1985,34:381391.
[55] HONGDY,SAUERW.CytotaxonomicalstudiesoftheLili
aceae(sl)intheChangbaiMountainsofnortheasternChina
[J].犆犪狋犺犪狔犪,1990,2:151164.
[56] KAWANOS,IHARA M,SUZUKIM,犲狋犪犾.Biosystematic
studieson犕犪犻犪狀狋犺犲犿狌犿 (Liliaceae:Polygonatae)I.somatic
chromosomenumberandmorphology[J].犅狅狋犪狀犻犮犪犾犕犪犵犪
狕犻狀犲犜狅犽狔狅,1967,80(951):345352.
[57] KAWANOS,SUZUKIM.Biosystematicstudieson犕犪犻犪狀
狋犺犲犿狌犿 (LiliaceaePolygonatae)VI.Variationingrossmor
phologyof犕.犫犻犳狅犾犻狌犿and犕.犮犪狀犪犱犲狀狊犲withspecialrefer
encetotheirtaxonomicstatus[J].犅狅狋犪狀犻犮犪犾犕犪犵犪狕犻狀犲犜狅
犽狔狅,1971a,84(996):349361.
[58] KAWANOS,SUZUKIM.Biosystematicstudieson犕犪犻犪狀
狋犺犲犿犻狋犿 (LiliaceaePolygonatae).V.Variationingrossmor
phology,karyologyandecologyofNorthAmericanpopula
tionsof犕.犱犻犾犪狋犪狋狌犿狊犲狀狊狌犾犪狋狅[J].犅狅狋犪狀犻犮犪犾犕犪犵犪狕犻狀犲犜狅
犽狔狅,1971b,84:299318.
[59] WCISLOH.ChromosomenumbersofcertainCanadianplants
[J].犃犮狋犪犅犻狅犾狅犵犻犮犪犆狉犪犮狅狏犻犲狀狊犻犪.犛犲狉犻犲狊犅狅狋犪狀犻犮犪,1987,29:
1930.
[60] GURZENKOV N N.Studiesofchromosomenumbersof
plantsfromthesouthoftheSovietFarEast[J].犓狅犿犪狉狅狏
犔犲犮狋狌狉犲狊,1973,20:4761.
[61] VOLKOVASA,BASARGINDD.Chromosomenumbersof
speciesofChabarovskterritoryflora[J].犅狅狋犪狀犻ˇ犮犲狊犽犻犼犣犺狌狉
狀犪犾(犕狅狊犮狅狑牔犔犲狀犻狀犵狉犪犱),2002,87(4):165167.
[62] NISHIKAWAT.Chromosomecountsoffloweringplantsof
Hokkaido(2)[J].犚犲狆狅狉狋狅犳狋犺犲犜犪犻狊犲狋狊狌狕犪狀犐狀狊狋犻狋狌狋犲狅犳犛犮犻
犲狀犮犲,1979,14:1523.
[63] NISHIKAWAT.Chromosomecountsoffloweringplantsof
Hokkaido(6)[J].犚犲狆狅狉狋狅犳狋犺犲犜犪犻狊犲狋狊狌狕犪狀犐狀狊狋犻狋狌狋犲狅犳犛犮犻
犲狀犮犲,1982,17:916.
[64] SEONBY,SEOLMR,IMJA,犲狋犪犾.Evolutionofendemicvas
cularplantsofUleungdoandDokdoinKoreanfloristicandcytot
axonomiccharacteristicsofvascularfloraofDokdo[J].犓狅狉犲犪狀
犑狅狌狉狀犪犾狅犳犘犾犪狀狋犜犪狓狅狀狅犿狔,2002,32(2):143143.
[65] HANMK,JANGCG,OHBU,犲狋犪犾.Acytotaxonomicstud
yofgenus犘狅犾狔犵狅狀犪狋狌犿inKorea[J].犓狅狉犲犪狀犑狅狌狉狀犪犾狅犳
犘犾犪狀狋犜犪狓狅狀狅犿狔,1998,28(2):187208.
[66] PROBATOVA NS,BARKALOV V Y,RUDYKAEG,犲狋
犪犾.Furtherchromosomestudiesonvascularplantspecies
fromSakhalin,MoneranandKurileIslands[J].犅狌犾犾犲狋犻狀狅犳
狋犺犲犎狅犽犽犪犻犱狅犝狀犻狏犲狉狊犻狋狔犕狌狊犲狌犿,2006,3:93110.
[67] KUMAR.KaryotypeintwoHimalayanspeciesof犘狅犾狔犵狅狀犪
狋狌犿[J].犈狓狆犲狉犻犲狀狋犻犪,1959,15(11):419420.
[68] KUROSAWAS.CytologicalstudiesonsomeeasternHimala
yanplantsandtheirrelatedspecies[J].犜犺犲犝狀犻狏犲狉狊犻狋狔犕狌狊犲
狌犿,狋犺犲犝狀犻狏犲狉狊犻狋狔狅犳犜狅犽狔狅,犅狌犾犾犲狋犻狀,1971,2:355364.
[69] MEHRAPN,SACHDEVASK.Cytologicalobservationson
someEastHimalayan monocots [J].犆狔狋狅犾狅犵犻犪,1979,44:
233240.
[70] ROYSC,GHOSHS,CHATTERJEEA.Acytologicalsur
veyofeastern Himalayanplants.II[J].犆犲犾犾犆犺狉狅犿狅狊狅犿犲
犚犲狊,1988,11:9397.
[71] >?@,Å Æ.!1ÇÈ9/àÅ#$qlmn&¢[J].…
a#$@A
,1998,20(2):8285.
GUZJ,SUNH.Thechromosomereportofsomeplantsfrom
Motuo,Xizang(Tibet)[J].犃犮狋犪犅狅狋犪狀犻犮犪犢狌狀狀犪狀犻犮犪,1998,
20(2):8285.
[72] ¼½¾,¿·….íÐ+7810£qÂÃ&¢[J].#$HONGDY,ZHUXY.CytotaxonomicalstudiesonLiliaceae
(S.L.)(1)Reportonkaryotypesof10speciesof6genus[J].
犃犮狋犪犘犺狔狋狅狋犪狓狅狀狅狀犿犻犮犪犛犻狀犻犮犪,1987,25(4):245254.
[73] HARAH,KUROSAWAS.CytotaxonomicalstudiesonJapa
noHimalayanelements[J].犑狅狌狉狀犲狔狅犳犑犪狆犪狀犲狊犲犅犪狋犪狀狔,
1963,38:7174.
[74] PROBATOVA NS,SOKOLOVSKAYA AP.Chromosome
numbersofthevascularplantsfromthefaeastftheUSSR
[J].犅狅狋犪狀犻ˇ犮犲狊犽犻犼犣犺狌狉狀犪犾犛犛犛犚,1986,71:15721575.
[75] TAHARAM.Chromosomemorphologyin犛犿犻犾犪犮犻狀犪犼犪狆狅狀犻
犮犪[J].犚犲狆狅狉狋狅犳犜狅犺狅犽狌犝狀犻狏犲狉狊犻狋狔(犐犞),1933,8:3337.
[76] KAWANOS,ILTISHH.Cytotaxonomyofthegenus犛犿犻犾犪
犮犻狀犪(Liliaceae)I.KaryotypeanalysisofsomeeasternNorth
Americanspecies[J].犆犺狉狅犿狅狊狅犿犪,1963b,14(3):296309.
[77] MEHRAP,SACHDEVAS.CytologyofsomeW.Himalayan
speciesof犘狅犾狔犵狅狀犪狋狌犿and犇犻狊狆狅狉狌犿[J].犆狔狋狅犾狅犵犻犪,1976,
41(34):573583.
[78] MEHRAP,PATHANIAR.Acytotaxonomicstudyofthe
WestHimalayanPolygonateae[J].犆狔狋狅犾狅犵犻犪,1960,25(2):
179194.
248 ! " # $ % & 36æ
[79] CHINNAPPACC,CHMIELEWSKIJG.Documentedplant
chromosomenumbers1987:1.Miscelaneouscountsfrom
westernNorthAmerica[J].犛犻犱犪犆狅狀狋狉犻犫狌狋犻狅狀狊狋狅犅狅狋犪狀狔,
1987,2(2):409417.
[80] HILLLM.IOPBchromosomedata1[J].犐狀狋.犗狉犵犪狀.犘犾.犅犻
狅狊狔狊狋.犖犲狑狊犾犲狋狋.(犣狌狉犻犮犺),1989,13:1719.
[81] LOVEA,LOVED.InChromosomenumberreportsLXXIII
[J].犜犪狓狅狀,1981,30:845851.
[82] KAWANOS,ILTISHH.Cytotaxonomyofthegenus犛犿犻犾犪
犮犻狀犪(Liliaceae)II[J].犆狔狋狅犾狅犵犻犪,1966,31(1):1228.
[83] STENARH.EmbryologischeundzytologischeBeobachtun
genüber犕犪犼犪狀狋犺犲犿狌犿犫犻犳狅犾犻狌犿und犛犿犻犾犪犮犻狀犪狊狋犲犾犾犪狋犪[J].
犃狉犽.犅狅狋.,1934,26:120.
[84] BELAEVAV,SIPLIVINSKYV.InChromosomenumberre
portsLXXIII[J].犜犪狓狅狀,1981,30:857860.
[85] KUMARVV,BRANDHAMPE.Cytotaxonomyof犇犻狊狆狅
狉狌犿犾狌狕狅狀犻犲狀狊犲 (LiliaceaePolygonatae)[J].犓犲狑犅狌犾犾犲狋犻狀,
1980,35:493497.
[86] LARSENK.StudiesinthefloraofThailand14.Cytological
studiesinvascularplantsofThailand[J].犇犪狀狊犽犅狅狋犪狀犻狊犽
犃狉犽犻狏,1963,20:211275.
[87] SATOD.KaryotypealterationandphylogenyinLiliaceaeandal
liedfamilies[J].犑犪狆犪狀犲狊犲犑狅狌狉狀犪犾犅狅狋犪狀狔,1942,12:57161.
[88] ´ƒÊ.ij’»£#$qlmn@A[J].#$1989,27(1):3948.
CHENSF.KaryotypeanalysisofeightspeciesofPolygontum
Mil.[J].犃犮狋犪犘犺狔狋狅狋犪狓狅狀狅狀犿犻犮犪犛犻狀犻犮犪,1989,27(1):3948.
[89] >?@,. ß,Å Æ,Z.01»2àÅ£=#$qÂÃ
@A
[J].…a#$@A,1993,15(4):377385.
GUZJ,WANGL,SUNH,犲狋犪犾.Acytologicalstudiofsome
plantsfromQinghaiXizangplateau[J].犃犮狋犪犅狅狋犪狀犻犮犪犢狌狀
狀犪狀犻犮犪,1993,15(4):377385.
[90] ·ËÌ,fÍ%,fßÎ.ij’q78³£qÂÃg„
[J].ÌÏ#$%@A,1987,5(1):110,
101102.
WANGJW,LIMX,LILX.Studiesonthecytotaxonomyof
PolygonatumⅡ.Karyotypesandevolutionofeightspeciesof
PolygonatuminChina[J].犑狅狌狉狀犪犾狅犳犠狌犺犪狀犅狅狋犪狀犻犮犪犾犚犲
狊犲犪狉犮犺,1987,5(1):110,101102.
[91] Ð Ñ,·ËÌ,fÍ%.ij’78é-Ò49/ij’#$ÂÃ
[J].ÌÏ#$%@A,1988,6
(4):311314,411412.
YANGJ,WANGJW,LIMX.Cytotaxonomicstudiesonthe
genuspolygonatumⅡ.Karyotypesof4speciesfromSichuan
Jinfoshan[J].犑狅狌狉狀犪犾狅犳犠狌犺犪狀犅狅狋犪狀犻犮犪犾犚犲狊犲犪狉犮犺,1988,6
(4):311314,411412.
[92] g@h,Ó§9,Ô Õ.Ûdij’q78Þ!#$
,1994,14(4):361368.
SHAOJZ,ZHANGDC,QIANF.Studiesoncytotaxonomyof
犘狅犾狔犵狅狀犪狋狌犿fromAnhui[J].犌狌犻犺犪犻犪,1994,14(4):361368.
[93] PANDITATK.Cytologicalinvestigationsofsomemonocots
ofKashmir[D].犆犺犪狀犱犻犵犪狉犺:PanjabUniversity,1979.
[94] PANDITAT,MEHRAP.Karyotypeanalysisoffivetaxaof
犘狅犾狔犵狅狀犪狋狌犿[C].犘狉狅犮.犐狀犱犻犪狀犃犮犪犱.犛犮犻.犘犾.犛犮犻,1982,48:
255263.
[95] DENGXY,WANGQ,HEXJ.Karyotypesof16populations
ofeightspeciesinthegenus犘狅犾狔犵狅狀犪狋狌犿 (Asparagaceae)
fromChina[J].犅狅狋犪狀犻犮犪犾犑狅狌狉狀犪犾狅犳狋犺犲犔犻狀狀犲犪狀犛狅犮犻犲狋狔,
2009,2:245254.
[96] THERMANE.Chromosomalevolutioninthegenus犘狅犾狔犵狅
狀犪狋狌犿[J].犎犲狉犲犱犻狋犪狊,1953a,39(12):277288.
[97] THERMANE.Onthecytologyofthegenus犘狅犾狔犵狅狀犪狋狌犿.
GroupsVerticilataandOppositifolia[J].犃狀狀犪犾犲狊.犅狅狋犪狀狔.
犛狅犮.犣狅狅犾.犫狅狋.犳犲狀狀.犞犪狀犪犿狅,1953b,25(6):126.
[98] MALLASB,BHATTARAIS,GORKHALIM,犲狋犪犾.In
chromosomenumberreportsLXX[J].犜犪狓狅狀,1981,30:75.
[99] Ð Ñ,·ËÌ,fÍ%.ij’78Ö6£ijqlmno}gÂÃ[J].ÌÏ#$%@A,1992,
10(3):201206,303.
YANGJ,WANGJW,LIMX.Cytotaxonomicstudiesonthe
genuspolygonatumⅢ.Chromosomenumbersandkaryotypes
of6speciesfromChina[J].犑狅狌狉狀犪犾狅犳犠狌犺犪狀犅狅狋犪狀犻犮犪犾犚犲
狊犲犪狉犮犺,1992,10(3):201206,303.
[100] KIEHN M,VITEKE,HELLMAYRE,犲狋犪犾.Beitrgezur
Floravon?sterreich:Chromosomenzhlungen [J].犞犲狉犺.
犣狅狅犾.犅狅狋.犌犲狊.?狊狋犲狉狉犲犻犮犺,1991,128:1939.
[101] M?S?ˇCEK,J,JAROLIMOV,V.Listofchromosomenumbers
oftheCzechvascularplants[J].犃犮犪犱犲犿犻犪,犘狉犪犺犪,1992.
[102] g@h,Ó§9,Ð×»,Z.ij’5£#$qÂÃ@A
[J].#$SHAOJZ,ZHANGDC,YANGJG,犲狋犪犾.Karyotypeanal
ysisof5speciesof犘狅犾狔犵狅狀犪狋狌犿 Mil.[J].犃犮狋犪犘犺狔狋狅狋犪狓
狅狀狅狀犿犻犮犪犛犻狀犻犮犪,1993,31(4):353362.
[103] ØÙÚ,fÛÜ,ÝÞ9.ßà9/10£íÐ+#$ÂÃ&
¢
[J].#$SHANGZY,LIRJ,CUITC.Reportonkaryotypesof10
speciesofLiliaceae(s.l.)fromQinlingrange[J].犃犮狋犪犘犺狔
狋狅狋犪狓狅狀狅狀犿犻犮犪犛犻狀犻犮犪,1992,30(5):438450.
[104] bc,ÐáL,sâa.ij’ã³£qlmn@A[J].s
ŽN“%V%&

)*+%º
),1984,(1):6776.
FANGYX,YANGBS,OUSH.Studiesonthechromosomeof
somespeciesof犘狅犾狔犵狅狀犪狋狌犿[J].犑狅狌狉狀犪犾狅犳犛犺犪狀犵犺犪犻犖狅狉犿犪犾
犝狀犻狏犲狉狊犻狋狔(NaturalSciences),1984,(1):6776.
[105] ¿%a,-äå,» æ.íÐ+M£#$qÂÄèé+ê
,2002,22(2):2226.
ZHUXN,JINXF,GAOZ.Karyotypeanalysisonthree
speciesofLiliaceae[J].犑狅狌狉狀犪犾狅犳犣犺犲犼犻犪狀犵犉狅狉犲狊狋狉狔犛犮犻
犲狀犮犲犪狀犱犜犲犮犺狀狅犾狅犵狔,2002,22(2):2226.
[106] ZHAOLH,ZHOUSD,HEXJ,犲狋犪犾.Acytotaxonomica
nalysisofChinese犘狅犾狔犵狅狀犪狋狌犿 (Asparagaceae)species
[J].犖狅狉犱犻犮犑狅狌狉狀犪犾狅犳犅狅狋犪狀狔,2014,32(4):441451.
[107] HILLL.M.IOPBchromosomedata9[J].犐狀狋.犗狉犵犪狀.犘犾.
犅犻狅狊狔狊狋.犖犲狑狊犾犲狋狋.(Zurich),1995,24:1920.
[108] -äå,ër_,Ùìí.îï4ij———ŠJijऀ
£Ò¤;ˆˆ£qΫ@A
[J].ç„Q%%&(ðé;Lñ
+%º
),2002,28(5):537542.
JINXF,QIU Y X,DINGBY.犘狅犾狔犵狅狀犪狋狌犿犮狔狉狋狅狀犲犿犪
var.犵狌狋犻犪狀狊犺犪狀犻犮狌犿———Anewvarietyof犘狅犾狔犵狅狀犪狋狌犿犮狔狉
3484½ .(/,Z:ÛÜÝ+ijk78%@A„¾
tonemaandcomparativestudiesonitandthesimilarspecies
[J].犑狅狌狉狀犪犾狅犳犣犺犲犼犻犪狀犵犝狀犻狏犲狉狊犻狋狔(AgricultureandLife
Sciences),2002,28(5):537542.
[109] TAMURAMN.Biosystematicstudiesonthegenus犘狅犾狔犵
狅狀犪狋狌犿 (Liliaceae)I.Karyotypeanalysisofspeciesindige
noustoJapananditsadjacentregions[J].犆狔狋狅犾狅犵犻犪,1990,
55:443466.
[110] WUS,YANGJ,RAOG.Systematicpositionof犘狅犾狔犵狅狀犪
狋狌犿狊犻犿犻狕狌犻(Convalariaceae)basedonmorphological,cyto
logicalandchloroplastDNAsequencedata[J].犅狅狋犪狀犻犮犪犾
犑狅狌狉狀犪犾狅犳狋犺犲犔犻狀狀犲犪狀犛狅犮犻犲狋狔,2001,137:291296.
[111] ·ËÌ,fÍ%,Ð Ñ.ʝgŠJijq€œÒ¤òó
x>¥
[J].#$WANGJW,LIMX,YANGJ.Variationandphylogeneticrela
tioshipsbetween犘狅犾狔犵狅狀犪狋狌犿狅犱狅狉犪狋狌犿and犘.犮狔狉狋狅狀犲犿犪[J].
犃犮狋犪犘犺狔狋狅狋犪狓狅狀狅狀犿犻犮犪犛犻狀犻犮犪,1991,29(6):511517.
[112] DAVLIANIDZEM T.Chromosomenumbersintherepre
sentativesoftheflorafromGeorgia[J].犅狅狋犪狀犻ˇ犮犲狊犽犻犼犣犺狌狉
狀犪犾犛犛犛犚,1985,70(5):698700.
[113] STRIDA,FRANZENR.InChromosomenumberreports
LXXIII[J].犜犪狓狅狀,1981,30:829842.
[114] SOPOVA M,SEKOVSKIZ.Chromosomeatlasofsome
Macedonianangiosperms.II.Cytologicalandtaxonomicas
pects[J].犌狅犱犻狊犲狀狕犫狅狉狀犻犽:犅犻狅犾狅犵犻犼犪犛犽狅狆犼犲犝狀犻狏犲狉狕犻狋犲狋
犅犻狅犾狅狊犽犻犉犪犽狌犾狋犲狋,1981,4(34):6576.
[115] MALAKHOVA L A.Kariologocheskijanalizprirodnykh
populjacijredkichiischezajushchikhrastenijnajugeToms
kojOblasti[J].犅犻狌犾犾.犌犾犪狏狀.犅狅狋.犛犪犱犪,1990,155:6066.
[116] MALAKHOVALA,MARKOVAGA.Chromosomenum
bersinthefloweringplantsofTomskregion.Monocotyle
dones[J].犅狅狋犪狀犻ˇ犮犲狊犽犻犼犣犺狌狉狀犪犾 (Moscow & Leningrad),
1994,79(7):134135.
[117] AMELCHENKO.Newtaxaofthegenus犘狅犾狔犵狅狀犪狋狌犿
(Convalariaceae)from Siberia [J].犅狅狋犪狀犻ˇ犮犲狊犽犻犼犣犺狌狉狀犪犾
(Moscow&Leningrad),2005,90(5):749752.
[118] KIMYS,KIMJH.Ataxonomicstudyofthegenus犘狅犾狔犵
狅狀犪狋狌犿inKorea[J].犑狅狌狉狀犪犾犓狅狉犲犪狀犘犾犪狀狋犜犪狓狅狀,1979,9:
2741.
[119] YONEZAWAN.犘狅犾狔犵狅狀犪狋狌犿犽犻狅狋犲狀狊犲 (Liliaceae),anew
speciesfrom Kyoto,Japan [J].犃犮狋犪犘犺狔狋狅狋犪狓狅狀狅犿犻犮犪犲狋
犌犲狅犫狅狋犪狀犻犮犪,1998,49:1925.
[120] TANAKA NY.Studiesonchromosomearrangementinsome
higherplants1.Interphasechromosomesinthreeliliaceous
plants[J].犆狔狋狅犾狅犵犻犪,1981,46:343357.
[121] WEISSSCHNEEWEISSH,JANGCG.Cytotaxonomica
nalysisofAustrianPolygonatumspecies(Ruscaceae)[J].
犖犲犻犾狉犲犻犮犺犻犪,2003,2(3):245250.
[122] “|ô,õ|Ê,fNö.=÷àÖ4£íÐ+#$qÂÊ
r@A
[J].!"#$%&,2000,20(5):882888.
FANXF,GUOXQ,LISW.Thestudiesonthekaryotype
diversityof4speciesofLiliaceaeinZiwuMountain[J].犃犮狋犪
犅狅狋犪狀犻犮犪犅狅狉犲犪犾犻犗犮犮犻犱犲狀狋犪犾犻犪犛犻狀犻犮犪,2000,20(5):882888.
[123] DRUSKOVICB,LOVKAM.IOPBchromosomedata9[J].
犐狀狋.犗狉犵犪狀.犘犾.犅犻狅狊狔狊狋.犖犲狑狊犾犲狋狋.(Zurich),1995,24:1519.
[124] LOVKVISTB,HULTGARDU M.Chromosomenumbers
insouthSwedishvascularplants[J].犗狆.犅狅狋犪狀狔,1999,
(137):142.
[125] PARFENOVV,DMITRIEVAS.Kariologicheskayakharak
teristikapredstaviteleiflorysosudistykhrasteni Berezins
kogobiosfernogozapovednika(soobshch.2)[J].犣犪狆狅狏犲犱
犅犲犾狅狉狌狊狊犻犻(Minsk)no,1987,11:6269.
[126] BALTISBERGER M,DAEHLERD,HORATS.Karyolo
gischeUntersuchungenandenschweizerischenArtender
GattungPolygonatum [J].犅狅狋犪狀犻犮犪犎犲犾狏犲狋犻犮犪,2002,112
(2):91101.
[127] ABRAMOVALI.AcytologicalsurveyofeasternHimalay
anplants.II[J].犅狅狋犪狀犻ˇ犮犲狊犽犻犼犣犺狌狉狀犪犾(Moscow & Lenin
grad),1990,75:652658.
[128] MEHRAP,VERMAS.CytotaxonomicObservationsonSome
WestHimalayanPteridaceae:(withPlatesLXXLXXIVand22
textfigures)[J].犆犪狉狔狅犾狅犵犻犪,1960,13(3):619650.
[129] LOVEA,LOVED.In:IOPBchromosomenumberreports
LXXVI[J].犜犪狓狅狀,1982,31:583587.
[130] AROHONKAT.Chromosomecountsofvascularplantsof
theislandSeiliinNauvo,southwesternFinland[J].犜狌狉狌狀
犢犾犻狅狆犻狊狋狅狀犑狌犾犽犪犻狊狌犼犪,犛犪狉.犃2,犅犻狅犾.犌犲狅犵狉,1982,3:112.
[131] SCRUGLIA.NumericromosomiciperlaFloraItaliana[J].
犐狀犳狅狉犿犪狋狅狉犲犅狅狋犪狀犻犮狅犐狋犪犾犻犪狀狅,1982,14:229233.
[132] VITEKE.BeitragezurFloravonOsterreich:weitereChro
mosomenzahlungen.(ContributionstothefloraofAustria:
furtherchromosomecounts.)[J].犞犲狉犺犣狅狅犾犅狅狋.犌犲狊.犗狊
狋犲狉狉,1992,129:215226.
[133] KRASNOBOROVIM,ROSTOVTZEVATS,LIGUSSA.
ChromosomenumbersofsomeplantspeciesofSouthSiberia
andtheFarEast[J].犅狅狋犪狀犻ˇ犮犲狊犽犻犼犣犺狌狉狀犪犾 (Moscow &
Leningrad),1980,65(5):659668.
[134] KRASNIKOVASA,KRASNIKOV A A,犲狋犪犾.Chromo
somenumbersofsomeplantspeciesfromthesouthofSiber
ia[J].犅狅狋犪狀犻ˇ犮犲狊犽犻犼犣犺狌狉狀犪犾犛犛犛犚,1983,68(6):827835.
[135] RASNIKOV A.Chromosomenumbersinsomespeciesof
vascularplantsfrom Novosibirskregion [J].犅狅狋犪狀犻ˇ犮犲狊犽犻犼
犣犺狌狉狀犪犾(Moscow&Leningrad),1991,76:476479.
[136] SOKOLOVSKAYAAP,PROBATOVANS.Chromosome
numbersinthevascularplantsfromthePrimoryeterritory,
Kamchatka region,Amur valey and Sakhalin [J].
犅狅狋犪狀犻ˇ犮犲狊犽犻犼犣犺狌狉狀犪犾(Moscow & Leningrad),1985b,70:
997999.
[137] SOKOLOVSKAYAAP,PROBATOVA NS,RUDYKAE.
ChromosomenumbersinthespeciesofthefamiliesAsteraceae,
Poaceae,RosaceaefromthePrimoryeRegim,KamchatkaandSa
khalin [J].犅狅狋犪狀犻ˇ犮犲狊犽犻犼犣犺狌狉狀犪犾(Moscow & Leningrad),
1985a,70(1):126128.
[138] SHATOKHINA.Chromosomenumbersofsomeplantsof
theAmurRegionflora[J].犅狅狋犪狀犻ˇ犮犲狊犽犻犼犣犺狌狉狀犪犾(Moscow
&Leningrad),2006,91(3):487490.
[139] bc.ʝq789:%@A[J].LN%x?,1989,8
(1):811.
FANGYX.Cytogeographicalstudyon犘狅犾狔犵狅狀狅狋狌犿狅犱狅狉
448 ! " # $ % & 36æ
atum[J].犆犺犻狀犲狊犲犑狅狌狉狀犪犾狅犳犈犮狅犾狅犵狔,1989,8(1):811.
[140] INOUES.Cytogeneticalstudiesofgenus犘狅犾狔犵狅狀犪狋狌犿1.
KaryotypeanalysisofsomeAlternifoliaspeciesinJapan
[J].犔犪犓狉狅犿狅狊狅犿狅,1965,60:19511962.
[141] NISHIKAWAT.Chromosomecountsoffloweringplantsof
Hokkaido(8)[J].犑狅狌狉狀犪犾狅犳犎狅犽犽犪犻犱狅犝狀犻狏犲狉狊犻狋狔(Educa
tionSection),1985,35:97111.
[142] DÚa,DøÊ,f ù.¤ð7<>Z#$qlmnlú
[J].!"#$%&,1990,10(3):203211.
MAXH,MAXQ,LIN.Chromosomeobservationofsome
drugplantsinXinjiang[J].犃犮狋犪犅狅狋犪狀犻犮犪犅狅狉犲犪犾犻犗犮犮犻犱犲狀
狋犪犾犻犪犛犻狀犻犮犪,1990,10(3):203211.
[143] ûüý,þìO.ʝq78%@A[J].ÿÜN“%V%&

)*+%º
),1988,20(2):4153.
LIUCL,GECJ.Astudyonthecytologyof犘狅犾狔犵狅狀犪狋狌犿
狅犱狅狉犪狋狌犿 (Mil.)Druce[J].犑狅狌狉狀犪犾狅犳犔狌犱狅狀犵犝狀犻狏犲狉狊犻狋狔
(NaturalScienceEdition),1988,20(2):4153.
[144] PARKJM,CHUNGKS,犲狋犪犾.Phylogenyofthe犘狅犾狔犵狅狀犪
狋狌犿狅犱狅狉犪狋狌犿 ComplexInferredfrom MultiplecpDNAand
NuclearRNApol2i23SequenceData(Ruscaceae)[J].犓狅
狉犲犪狀犑狅狌狉狀犪犾狅犳犘犾犪狀狋犜犪狓狅狀狅犿狔,2011,41(4):353360.
[145] ! ",.ßa,Ðø#,Z.-"Ö20£#$lmno}
[J].ÌÏ#$%@A,1995,13(2):180182.
SHAW,WANGLH,YANGXJ,犲狋犪犾.Chromosomenum
bersof20speciesfrom NortheastChina[J].犑狅狌狉狀犪犾狅犳
犠狌犺犪狀犅狅狋犪狀犻犮犪犾犚犲狊犲犪狉犮犺,1995,13(2):180182.
[146] POGOSIAN A.Chromosomenumbersinsomespeciesof
monocotyledonsfromtheTranscaucasia [J].犅狅狋犪狀犻ˇ犮犲狊犽犻犼
犣犺狌狉狀犪犾(Moscow&Leningrad),1997,82:117118.
[147] GADNIDZERI,GVINIASHVILITN,DANELIAIM,犲狋
犪犾.ChromosomenumbersofthespeciesoftheGeorgianflo
raesoftheGeorgianflora[J].犅狅狋犪狀犻ˇ犮犲狊犽犻犼犣犺狌狉狀犪犾(Mos
cow&Leningrad),1998,83(10):142147.
[148] DANIELAI.IOPBchromosomedata11[J].犖犲狑狊犾犲狋狋.犐狀狋.
犗狉犵犪狀.犘犾.犅犻狅狊狔狊狋.(Oslo),1997,26(27):1314.
[149] GERVAISC.Listeannoteedenombreschromosomiquesde
laflorevasculairedunordestdel'AmeriqueII.[J].犖犪狋狌
狉犪犾犻狊狋犲犆犪狀犪犱,1981,108:143152.
[150] þìO,fZ$.4-9/>Z#$lmno}qlú(II)
[J].¥›>,1989,20(6):3435.
GECJ,LIYK.Observationonthechromosomenumbersof
medicinalplantsofShandongProvince (II)[J].犆犺犻狀犲狊犲
犜狉犪犱犻狋犻狅狀犪犾犪狀犱犎犲狉犫犪犾犇狉狌犵狊,1989,20(6):3435.
[151] %&è,¯Î,()*,Z.ʝ>:úûÒ¤–ŒYZ@
A„¾
[J].SÖ@A,2012,34(3):7376.
JIANGZL,BAIRX,GUIFR,犲狋犪犾.ResearchProgresson
PharmacologicalEffectandUtilizationofPolygonatumOdo
ratum [J].犛狆犲犮犻犪犾犠犻犾犱犈犮狅狀狅犿犻犮犃狀犻犿犪犾犪狀犱犘犾犪狀狋犚犲
狊犲犪狉犮犺,2012,34(3):7376.
[152] ´ +,,--,Å./,Z.3£>Zijq%9<Ò>
:Š@A„¾
[J].¥›>,2015,46(15):23292338.
CHENH,FENGSS,SUNYJ,犲狋犪犾.Advancesinstudieson
chemicalconstituentsofthreemedicinalplantsfrom犘狅犾狔犵狅
狀犪狋狌犿 Mil.andtheirpharmacologicalactivities[J].犆犺犻狀犲狊犲
犜狉犪犱犻狋犻狅狀犪犾犪狀犱犎犲狉犫犪犾犇狉狌犵狊,2015,46(15):23292338.
[153] STEBBINSGL.CytogeneticstudiesinpaeoniaII.theCytol
ogyofthediploidspeciesandhybrids[J].犌犲狀犲狋犻犮狊,1938,23
(1):83110.
[154] RAMSEYJ,SCHEMSKEDW.Pathways,mechanisms,and
ratesofpolyploidformationinfloweringplants[J].犃狀狀狌犪犾
犚犲狏犻犲狑狅犳犈犮狅犾狅犵狔犪狀犱犛狔狊狋犲犿犪狋犻犮狊,1998,29:467501.
[155] ADAMSKL,WENDELJF.Polyploidyandgenomeevolu
tioninplants[J].犆狌狉狉犲狀狋犗狆犻狀犻狅狀犻狀犘犾犪狀狋犅犻狅犾狅犵狔,2005,8
(2):135141.
[156] LEVINDA.Polyploidyandnoveltyinfloweringplants[J].
犃犿犲狉犻犮犪狀犖犪狋狌狉犪犾犻狊狋,1983,122(1):125.
[157] SOLTISDE,SOLTISPS.Polyploidy:recurrentformation
andgenomeevolution[J].犜狉犲狀犱狊犻狀犈犮狅犾狅犵狔牔犈狏狅犾狌狋犻狅狀,
1999,14(9):348352.
[158] BEBBETT,MICHAEL.Perspectivesonpolyploidyinplants
ancientandneo[J].犅犻狅犾狅犵犻犮犪犾犑狅狌狉狀犪犾狅犳狋犺犲犔犻狀狀犲犪狀犛狅犮犻犲狋狔,
2004,82(4):411423.
[159] STEBBINSGL.Chromosomalevolutioninhigherplants
[M].London:EdwardAmoldLtd.1971.
[160] STEBBINSGL.Polyploidyinplants:unsolvedproblemsand
prospects[M]//Lewis W H.Polyploidy:biologicalrevel
ance.NewYork:PleumPress,1980:495520.
[161] TATEJA,SOLTISDE,SOLTISPS.Polyploidyinplants
[M]//GREGORYRT.Theevolutionofthegenome.Am
sterdam:Elsevier/AcademicPress,2005:371426.
[162] OLSON K,GORELICK R.Chromosomalfissionaccounts
forsmalscaleradiationsinZamia(Zamiaceae:Cycadales)
[J].犅狅狋犪狀犻犮犪犾犑狅狌狉狀犪犾狅犳狋犺犲犔犻狀狀犲犪狀犛狅犮犻犲狋狔,2011,165
(2):168185.
[163] ROBERTSON W.Chromosomesstudies.I.Taxonomicrela
tionshipsshowninthechromosomesofTettegidaeandAcri
didiae,LocustidaeandGrilidae:chromosomesandvariations
[J].犑狅狌狉狀犪犾狅犳犕狅狉狆犺狅犾狅犵狔,1916,27:179331.
[164] >?@,. ß,f g.í0„9/7<á=ü#$q7
88N%@A
[J].…a#$@A,1992,14(S5):13.
GUZJ,WANG L,LIH.Karyomorphologicalstudiesof
somemonocotsinDulongjiangarea[J].犃犮狋犪犅狅狋犪狀犻犮犪犢狌狀
狀犪狀犻犮犪,1992,14(S5):13.
[165] LEEYN.ChromosomenumbersoffloweringplantsinKorea
(3)[J].犑.犓狅狉犲犪狀犚犲狊.犐狀狊狋.犅犲狋犲狉犔犻狏犻狀犵,1970,5:127129.
[166] 120,Å Æ,>?@.34/}=#$lmn@A²
³
[J].…a#$@A,2004,26(1):3557.
NIEZL,SUNH,GUZJ.Asurveyofchromosomenumbers
fromAngiospermsoftheHengduanMountains,S.W.China
[J].犃犮狋犪犅狅狋犪狀犻犮犪犢狌狀狀犪狀犻犮犪,2004,26(1):3557.
[167] NIEZL,WENJ,GUZJ,犲狋犪犾.Polyploidyinthefloraofthe
HengduanMountainshotspot,southwesternChina[J].犃狀
狀犪犾狊狅犳狋犺犲 犕犻狊狊狅狌狉犻 犅狅狋犪狀犻犮犪犾犌犪狉犱犲狀,2005,92(2):
275306.

!"

#$%

5484½ .(/,Z:ÛÜÝ+ijk78%@A„¾