免费文献传递   相关文献

Taxonomic Revision of Rhododendron tsoi (Ericaceae) and Its Related Taxa

中国特有植物景烈杜鹃(杜鹃花科)及其近缘类群的分类修订*


基于文献考证、标本查阅和野外调查,对景烈杜鹃Rhododendron tsoi(杜鹃花科)及其近缘类群主要形态性状的分类学意义进行分析,进而作分类学修订。结果表明: 叶片的大小、形状、叶片先端的形态和叶背的毛被特征可以区分景烈杜鹃及其近缘类群,可作为划分种下等级的依据,而雄蕊长度及其与花柱的长度比较在种内变异很大,不能作为种及种下等级划分的依据。从而将景烈杜鹃及其近缘类群划分为1个种及4个变种,提出4个新组合变种名,即: 背绒杜鹃R. tsoi var. hypoblematosum 10(Tam) X. F. Jin & B. Y. Ding、细石榴花R. tsoi var.nudistylum(Tam) X. F. Jin & B. Y. Ding、惠阳杜鹃R. tsoi var Huiyangense (Fang & M. Y. He) X. F. Jin & B. Y. Ding和千针叶杜鹃R. tsoi var.polyraphidoideum(Tam) X. F. Jin & B. Y. Ding。将棒柱杜鹃R. crassimedium和粗柱杜鹃R. crassistylum作为R. tsoi var.hypoblematosum的异名,将垂钩杜鹃R. unciferum和鼎湖杜鹃R. tingwuense归并入R. tsoi var. nudistylum。描绘各个分类群的地理分布。

A taxonomic revision of Rhododendron tsoi (Ericaceae) was made in this paper, based on literature survey, field investigation, specimen examination, and evaluation of taxonomical significance of the major morphological characters. The size and shape of leaves, leaf apex and indumentum on lower surfaces were found to be reliable diagnostic characters for the circumscription of R. tsoi and its related taxa. However stamen length and its relative length in comparison with the style were so variable that they could not be used to circumscribe R. tsoi at species and sub-species level. Based on the leaf characters the previous R. tsoi and its related taxa were circumscribed as one species, that is R. tsoi,and four varieties, I.e. R. tsoi var. hypoblematosum (Tam) X. F. Jin & B. Y. Ding,R. tsoi var. nudistylum (Tam) X. F. Jin & B. Y. Ding,R. tsoi var. huiyangense (Fang & M. Y. He) X. F. Jin & B. Y. Ding , and R. tsoi var. polyraphidoideum (Tam) X. F. Jin & B. Y. Ding. In addition, R. crassimedium and R. crassistylum were reduced to the synonyms of R. tsoi var. hypoblematosum, and R. tingwuense and R. unciferum were synonymized into R. tsoi var. nudistylum. The geographic distribution of R. tsoi and these intraspecific taxa were described as well.


全 文 :! !"" ! ##
# %
& ’ ( )
’()*+,)- ’)./-* ’)+)(-*
/012!"*+02#
341 5*
!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!!
\]^7 _‘‘6
662 =Fo-F)/6UKe())D =F 76%%78@2 SA&)D&’U/0[9)D >? 7667%%@ 72 HFF)/6UKe())D HF 7"%#X ! "# /{Fxm>Gz0E@xRSDE*bN2’: !"##’#& ()#*6’:2(X23u;!lNpOJ
33#z!)qrmnzl*m!lz!)l4[czN2’:23u;!l*4lu^zv?F8#8o*!BCM423U2b#M4?@$v?)9
sF*|2luv2v?F8^z#8oox!AN2’:23u;!l^zu 6Iv2 !I)v*z9 !I6q
5)v<*0tAD’: ! 5 ()#* 9:; 5 "+,#-.%/0(#)1/6,:5 3?@ A B5 C5 D?@EG-VE2 ! 5 ()#* 9:; 5 &1$*)(+.1/
6,:5 3?@ A B5 C5 D?@EGF[’: ! 5 ()#* 9:; 5 "1*+0&2%&)%6>:@E A F5 C5 GHX=5 >5 3?@ A B5 C5 D?@ExGP
[’: ! 5 ()#* 9:; 5 ,#.+’0,"*#*%1/6,:5 3?@ A B5 C5 D?@EoAHb’: ! 5 3’0))*/%*1/ x}b’: ! 5 3’0))*)(+.1/lu ! 5 ()#* 9:; 5 "+,#-.%/0(#)1/ #9<*AHI’: ! 5 1&3*4%’1/ x^N’: ! 5 (*&251%&)% ?}& ! 5 ()#* 9:; 5 &1*)(+.1/oyP6Iz!l#h6zao
%%&4%# J %%7J%%I J %& J %6o
/STUt7QG_()/SghTU67%7#%6%8Xo
"fJKuYklmo
!"#&’( )*+’,’- !"##%&’#& ()#*6./’("(*"*X"%0 12, )*3"2*0 !"#" 3?@ =?:0KH@E6 3?@ ’L4?L4 D?@E B?@EM:@E7 NL40 =?:0O?:06
662 63"##. #4 7*4% 0&8&9*’#&/%&( 63*%&3%)*:0&2;"#1 <#’/0. =&*9%’)*(+ :0&2;"#1 76%%78@2 63"##. #4 >#’%)(’+ 0&?*#P@%3"&#.#2+*A"%B*0&2 >#’%)(’+ C#..%2% 7*&~0& 7667%%@ 72 63"##. #4 7*4% 0& 8&9*’#&/%&( 63*%&3%)*D%&;"#1 =&*9%’)*(+ D%&;"#1 7"#X
45,2/"(2t - Q:O0@0?@9HSQ?E:Q?0@*SVHR?:@U SL:VH 0K 1H:9HS*1H:K :VHO :@U ?@U4R?;R4S0 9:;?:W1H QL:Q QLHM R041U @0Q WH 4SHU Q0 R?;R4V;H9?04S ! 5 ()#* :@U ?QS ;H1:QHU Q:O: TH;H R?;R4"+,#-.%/0(#)1/6,:5 3?@ A B5 C5 D?@E*! 5 ()#* 9:; 5 &1$*)(+.1/6,:5 3?@ A B5 C5 D?@E*! 5 ()#* 9:; 5
"1*+0&2%&)%6>:@E A F5 C5 GHX=5 >5 3?@ A B5 C5 D?@E*:@U ! 5 ()#* 9:; 5 ,#.+’0,"*#*%1/6,:5 3?@ A B5 C5
D?@E 5 )@ :UU?Q?0@*! 5 3’0))*/%$*1/ :@U ! 5 3’0))*)(+.1/ TH;H ;HU4RHU Q0 QLH SM@0@M! 5 (*&251%&)% :@U ! 5 1&3*4%’1/ TH;H SM@0@MQLHSH ?@Q;:SVHR?K?R Q:O: TH;H UHSR;?WHU :S TH11 5
6*7 8/0,t !"##’#& ()#*@SQ:Q?SQ?R:1 :@:1MS?S@<0;VL010E?R:1 9:;?:Q?0@ 0K 1H:K@Q:O0@0@HT SM@0@M<
FH;;?1166&7!XVWZ7Q)u0#vA*[C Gwh6:=VRuZXEt{#z0*%rPN2’ :6!"##’#& ()#*X*0v[C:Z*4fZ{*=V9QZ|L*R.ovSMuXN2N26/
!kuVN2’:Zo\-*GHZ7z0Cdxz
!VW|}*&*-%XLLJ#!lL^%:}
yk%r**+ntOP5F6!"#XbQ)’:
2B*Du0g.VWv*%rPsi6v*3N2’:LuJ#v%^N’: ! % "#)(@G
\*OP56!"&’@!"&(@!")XOPVW.Eh
c’:2Bu0v*-%rP%ILJ#!l*z{
ntAD’: ! % *+,-./(01"-)’0GHb’: ! %
231))#0(4#’0GGP[’: ! % ,-/+31,*#4-#4(’0G-V
E2 ! % )’.(35(+’4#)"+/’0 xHI’: ! %
2#6(3’0@lFdF6!")X%rPF[’: ! %
*’#+1)(@Ts06!"#X%rP}b’: ! % 231))#)"+/’0o[1#=L!l*[qLbH*23Q 3RmO*M|2 !,- ./*BC -S*FMN+|F M*2TU2bN+*UX‘"+|uo 01’/(2+3’45F6!""6X7Ab’:2B4{RC B !*-4-4($43-$ 78(925% 7)’")’)# l9PEqo<brVvPHI’:xN2’:*}A^N’:x-V E2luN2’:#9<@Hv*I+vuGP[ ’:nXIWz|uN2’:#v*}AHb’: xAD’:luGP[’:#9<@F[’:lu XIW]#vz6*!}b’:;f%z6oTs 06!"":XRXY<7u0YZ’:2BZg<*A
N2’:GGP[’:GHI’:G^ N’:GHb’
:G}b’:GAD’:;luW]#v!S[P
,*}AF[’:z6u^N’:#9<*!’P,
-VE2 ! % )’.(35(+’4#)"+/’0o;2F6&66-X
$Y<3=+’ => 0145’ZEX~o[&6&66:XVW<7’:2Bu0
v*]XAA-VE2luN2’:#9<*s&_
\f%m 01’/(2+3’45F6!""6X#TYo &_*b{N2’:23u;v*[C#z!< !qsi_‘xsFz]on^_%6#Koz
0n|2b8‘y).#p7#o0FxKoz0
9%*p5Ko6h#al_‘*bN2’:23u
;v#OJ33mnP-~#TU*m!l9z!
! ~3Ula
!"! HIDE
7?S6Iz0b<*E@N2’:23L S!l#z06NpnKoz0X*.oKoz01 Q,#)**Z7Ec{hmnQR#z0Cd*
#P$RSf%Cd8F[’:x}b’:[S*
3I!l;%XK^1#z0Cd6‘0^1(r
!XoCdv[G+_‘uX;*[X/42f4v
#)9g4*}bz0S[,6xKob6ILS
!lz0mnDXOE*Q,=Lv#OJ33o
!"# iN4:
D ?@?? !!,-wJIJHX3z0[Mx[d
#:;>xzO8*[f4|Hv[qFH#)9
>x[d#:;>xzO8*bU1#z0mnP9G
zlx?@oIJ[MB[d-f4x?@|Hv
#[O)9o
& pqUzl
#"! 8a<9L_V
N2’:23LS!l#OJ33e(r !o
$al_‘xTU/J1%:*N2’:#[qu
rONurO*Md?u!,)-" : C 6,6&! #*U-V
E2GHI’:GGP[’:GHb’:G^ N’:G
AD’:;%I&78968 D 6,6!X*!3I%v
’:2f&73OL*;f%89o
x[q6J-:*4[s&zu ) I!Lt!X
rO}5H;*1N2’: !v@&Xf;NQ;*-
VE2GHI’:GHb’:G^ N’:GAD’:G
}b’:xF[’:@)XE;*1GP[’: !vo
9 ! N2’:23u;!l[qFH#)9
<49%! E2’>F74G2 *’+4’A4=5 => ! % ")-# ’5H
A12 +23’A2H A’I’6/2’5 C ?JX
x[A#+B-:*F_v!N+N5gh#
+*t%#B+m*:o
4(*[q6J#OJx[qAq#+B3<*
al?[K*al‘%)9o ))! ## Si7Ft<7k%u0N2’:6’:2(X23u;!l#z!<br!"! c:;_L_V N2’:23u;!l#[qFH)9e(9 !ox9 !s&4[:8*N2’:#[qFHs& ?3Iv!H*Npf4[q#M41*4I{I
bU-VE2G^ N’:|uoF[’:61 "IK
oz0X[qH?3I#vF*!3Iv!x[qF
H1:*f%s&#c{o4(*YZ[qFH4[
c{F[’:x3I#!lo
5 # <=>?XY@A*/LBC
%&’ # ()*+%,-.)/ )0 ! ’ "#% %/1 234 ,45%241 2%6%
z!l
#%&’ TU‘0 ^1 ()*+,)-
’+./), &0
1’-121-+3* 33 45,67), BCM4 87.)’ 3)’975
[M:[d
;)0 3)’975: 3)0 <1-75
[q6J
;)0 =)%
[A+B
>’-+.)’7+. &’
3&<), 3)0 *+,06)
N2’:
! ? "#% !@ A B !@ ..*|FMNuFM*?2 bMN?2bQ A B !@ .. 3&’9*+’)C+3 &, )C+3 1’ 3)’975*3&’9), &, *5&,7 75’ *7D3)*
!E@FA G H IEI"! J
rO*H; K&+’-)-*.+6,&’7)
N+Nh+
L3/,&+* &, *=,*)3D =+/)*6)’7
-VE2
! ? #&’()(*+( 2, ? )&,%#"-.&/ !I !! B !@ ..*|FM*?2bQN uFM !! B !@ .. 3&’9*+’)C+3 1’ 3)’975*
*5&,7), &, )C+3 7& *7D3)* !EM@G N H IEIN" M /6 &rO*f2Q; O331=7163*&/7+*)
’- .+6,&’7)
N+Nh+
L3/,&+* &, *=,*)3D =+/)*6)’7
HI’:
! ? &)0%1(*&/ A
!" B !G ..*|FM*?2bMN
?2bQ
!" B !G .. 3&’9*+’)C+3 1’ 3)’975* 3&’9), &, *5&,7 75’ *7D3)*
!EAIM I H IEI@M J
/6-
&rO*Q;
O331=7163*.+6,&’7)
h+
8=,*)3D =+/)*6)’7 GP[’: ! ? 2.-*324%,%,(&/ A !@ B !G ..*|FMNuFM*? 2bQNuFM !@ B !G .. 3&’9*+’)C+3 &, )C+3 1’ 3)’975**5&,7), &, )C+3 7& *7D3)*
!EM"@ J H IEIAI M
/6-)
j3&rO*E;
P27)Q)331=7163*
6+.1’7)
Igh#+
R),D *=,*)3D =+/)*6)’7 Hb’: ! ? 0*3##%/(,%&/ J !" B !G ..*|FM*?2bM !" B !G .. 3&’9*+’)C+3 1’ 3)’975*
3&’9), 75’ *7D3)* !EMGJ F H IEI"@ F /6-)0 M&rO*Q; ;&’9Q)331=7163*
.+6,&’7) #B+ S)’*)3D =+/)*6)’7 ^N’: ! ? "%)56&()#( "" !I B !G ..*|FM*?2bMN uFM !I B !G .. 3&’9*+’)C+3 1’ 3)’975*
3&’9), &, )C+3 7& *7D3)* !EMGI M H IEIFG I /6-)09 &rO*f2Q; O331=7163*&/7+*)
’- .+6,&’7)
N+Nh+
L3/,&+* &, *=,*)3D =+/)*6)’7
AD’:
! ? 4-2’.(/3"#&/ "@
!I B !@ ..*|FM*?2bMN
uFM
!I B !@ .. 3&’9*+’)C+3 1’ 3)’975* 3&’9), &, )C+3 7& *7D3)*
!EA!J ! H IEI"" M
/6-)09
M&rO*Q;
;&’9Q)331=7163* .+6,&’7)
#B+
S)’*)3D =+/)*6)’7
}b’:
! ? 0*3##%#"-.&/ "
A B !" ..*uFMN|FM*?2
bMNuFM
A B !" .. 3&’9*)C+3 &, +’)C+3 1’
3)’975*3&’9), &, )C+3 7& *7D3)* !EA"@ @ H IEIGF N /6-)095 M&rO*Q; ;&’9Q)331=7163*
.+6,&’7) #B+ S)’*)3D =+/)*6)’7 F[’: ! ? 4&%-3)5()#( " !@ B !G ..*uFM*U2buF M !@ B !G .. 3&’9*)C+3 1’ 3)’975*
)C+3 7& *7D3)* !EAF" G H IEI@M ! /6-)095 j3&rO*f2Q ; P27)Q)331=7163* &/7+*) ’- .+6,&’7) h+ 8=,*)3D =+/)*6’)7
!"7 DEQL_V BCntFMG2bntM{NQ{BC[C tBEu"+op#z!33*4YZ4_‘0B C#U1%:*N2’:xu;!l#BCM4; %sF)96r !Xo3%v4?BCM*%vUb FM*%v@4@bQ*=L33al?/jHX
3z01*Q%sF#)9o=U T1’F6"IIJXk K’: ! ? 1&04#%%1.%&/ #z0zl~#o
@ p7
YZbLSOJ33#j5zl*4[%:N
2’:U3u;!l#c{1{[q#T%I3 3*p5h6za*bN2’:23u;!l*l9 G@ & ’ ( ) GF" Pz!*[CE[VvZ8$,ICB<I|:<br*=/CDV)vZ,u5I)v*Ko)vU6)v#c{kle(O
dro
!"# )_(.F5
"# [qrONurO*6Jr!5H; "% N2’: ! % "#% &’ % "#%lllllllllllll
"# [qM&rOG&rOjj3&rOo
(# [qAq#BRs_mn+ ")% AD’: ! % "#% &’ % &’()*+,-"#.,lllllllllll
(# [qAqN+NBghmn+o
*# [q6Jf;NQ;o
!# [q@H*MY+|2 ( +,*dY+|2 " +, "+% -VE2 ! % "#% &’ % /.0%#"’*.,ll
!# [q@F*M - ( +,*d - " +, ".% F[’: ! % "#% &’ % &.%’-/1+/#+llllllll
*# [q6JE; "/% GP[’: ! % "#% &’ % (*’2-(&%0%0+.,llllllllllllll
!" (*GH N2’: !&00+/02/ "#% 0/’’ % 12 312422 5+1 % 6 % "*t
"!(# "7*!@2829,8:; 12 <+82% =8’,% 512 % *t"!>*? % !(@A# "7B!@C% =% D,*5:’&/9 E/2% FG8.8./2.’82 5%
=G12tH!# "7>*@=G,)/’I12 J F/ 12 K.12):’4G 6 %
L8? % !Bt"*H# "77A@0% M% N/ 12 OI % F/1P % C8P% 512 %
@B6(Xt!"B# "77!@0% M% N/ J =G,)/’I12 12 OI %
=G12 "!t!!7# (AA@# DMCKt=G12% E:24.8246Q )X*0?% Q2R:2;G26Et{X*"7*( S A! S (@*T% D% D;246op7X(A**(6G8I8?9P/*K*UVQq*UMWq*Cq*CKqX%
+tde*< A#@ X ( ,o>$/v#B:r#
mn+o[+d/{HvXJ*s,Nus,*&r
OGj3&rONtdjO*M A#@ X (#H +,*d
A#! X "#* +,*6Jrf*+5H;u*/sdvON
rO*1qh/mn+NN+*?q//mn+*w
&@[RM " X * ,,*hBmn+ozO2UX/*
%2 * X H{*2|M * X H ,,*#Bmn+o2} @
%*%q|s&*Bmn+@23Q3Rm3*:R
_Nqr_*M 7 X "! ,,*%q @*3~r~3*M *
X @ ,,*9qN+*%qMrO@BC @*|FM*2
TM 7 X "! ,,*N+Nu/sBB!+@s8jI
O*#Bmn+@2bM B X "* ,,*N+o"qMj
IO*M ! X @ ,,*hBmn+o2# !2@%o
<7k%v*za{Q)GQBGIcGABxN
Eo
" N2’:6Ko)vX !&00+/02/ "#% 0/’’ % &’ % "#% 0)v[qrONurO*6Jfr!5H;o 1({Q)RuEt{6Ko6hXo E:24.8246Q)X*3824,/26wh*:=R
uX*0?% Q2R:2;G26Et{X*Q2R:2;G2 KYP/.%
6Et{DE#XB"!"@67X (@*B76 6A"H*QU5 S A"B*QU5 S A">*QU5 S A"7*QU5 S A(!
6ND=X%
") AD’:66q5)vX
!&00+/02/ "#% 0/’’ % &’ % &’()*+,-"#., 6D,XZ% O% 612 J L% M% [124*+8,)% /? ;?? % 28& % L;1829,% !&00+/02/ &’()*+,-"#., C% =% D, 12 L:II % L8? % F/; %6N’)12X(6"Xt7A*\14 % (# "7>( /? 12 5:’&/9 E/2% FG8.8./2.’82 5% =G12t!>*PI %
(Ht" S H# "7>*# DMCKt=G12% 6124Y16ABX*
5:1+G:26&WX*0?% 6124424;G26‘a{X*5% 5 % 316’%(XHHAA@@6G8I8?9P/*3LEqX%
222!&00+/02/ 32-##%,+0%., C% =% D, 12
L:II % L8? % F/; %6N’)12X(6"Xt7H# "7>( /? 12 5:’&/9 E/2% FG8.8./2.’82 5% =G12tH!*PI % Ht" S H# "7>*#
;92% 28&% DMCKt=G12% 6124Y16ABX*5:1+G:26& WX*]1I1246)cX*L2^1860*X*"7@7 S A! S (B* 4 % 4 % 5-%6’%(X(B76G8I8?9P/*3LEqX% 222 !&00+/02/ 32-##%#"’*., 0% M% N/ 12 6 % 51+G:2 V21& %6Q? % 5+1 % /. %X(!6"Xt>**\14 % "# "7>B# ;92%
28&% DMCKt=G12% 6124Y16ABX*5:1+G:26&WX* _:8262?X*"7BA S A! S (A*(*H S 67(+0 %6(*H~
+qX*AH6G8I8?9P/*CKq@1;8?9P/*CKqX%
0)vUKo)v#c{{[qM&rO* Aq#BRs_mn+o za{ABBs‘a{XEo 6124Y16 A BX* 5:1+G:2 6 & WX* 0?% 6124424;G26‘a{X*5% 5 % 316’%(X*>H@
6*!>"@*@*H"6"B@*7H***H7@*HHA@"(63LEX@6 % Z18246,-X
(*7A*(!(!63LEX*(!((**A*(63LE*CKX*(!*"
6CKX@T% N% T2 J I %6sF.FX"(AA!6@*! B# Si7Ft<7k%u0N2’:6’:2(X23u;!l#z!!"#"%#&’ ()*+,)6-./X@!"#"%#&’ ,0+*,*12++**
13+,116456X@0)7 8 9:;<=>60)7 ~+qX*(3
6?9X@/> -> @&!"A6S7&X’ > " >**+2*,,,)*
,,,36@BCX@D> E> FG"6Si7X,0*26@BCX>
,H -VE266q5)vX
!"##%&$’#& ()#* I K!J > &+$*)(,-+.6B!%X
D> E> FG" L .> M> NG"A*H#%O>

"#K >
.!’G#"%> !"##%&’#& )+/%&%’0% ?> C> B!% K!J >
&+*)(,-+. ?> C> B!%*Q&JK< /<"> RS#=#=<"=J#" Q>
CSG"!t7+ L ,+(T ,2()T BM?9tCSG"!> /&!"A=#"A6Q
)X*@&G!"A6F[X*IP> -G!"S&!’S!"6/2{X* ,2)* 8 +( 80* 1 > 2 > 2)3&46op7X 0**(7 6S#U#P;<*V.QCq@G’#P;<*WX> 222!"##%&’#& (*&45+%&)% ?> C> B!% G" I<=>
I!P > /&!"A=#"A ,21(63Xt)7*XGA > 7T ,21(T ’"> "#K> BM?9tCSG"!> /&!"A=#"A6Q)X*/!#!#6IP> NG"AS&’S!"6^N{X*?> C> B!%6OP5X1),1
6S#U#P;<* V.QCqX> 222!"##%&’#& +&6*7%’+. ?>
C> B!% G" /&GS!G! 060Xt1)*XGA > *T ,2(0T ’"> "#K> BM?9tCSG"!> /&!"A:G6QBX*@<"A:G!"6U>X* IP> I!!"’S!"6g1{X*Y> Y> CS<"6mM2X*+)++ 6S#U#P;<*V.4q@G’#P;<*456qX> 0)vUKo)vc{[q&rO*6Jf
;NQ;o
za{Q)GQBxNEo
/&!"A=#"A6Q )X* /!#!#6< pX* IP> NG"AS&’S!"6^N{X*?> C> B!%6OP5X1))2 6V.QCX@/> -> QSG6V7tX,*37(6V.QCX@/> Z> NG"A L /> -> QSG6fQbxV7tX07+6V.QCX@ W> [!"A6n3jX)+,7*1),16CBBIX@0)7 8 9:;<=>60)7~+qX022(6?9X@Q> @> FG" L F> D& 6SwDx%XB@Q 8 ++0*B@Q 8 ++)*B@Q 8 ++7*
B@Q 8 ++2*B@Q 8 +,+*B@Q 8 +,,*B@Q 8 +,)*B@Q 8
+,(*B@Q 8 +,26@BCX@F > D& L @> .> -G6%xi .5X3(*32**+**,**0*****7**16@BCX> FG!#UG"A64cX*-> B<"A6;tX3(706 V.QCX> /&!"A:G6QBX*@<"A:G!"6U>X*IP> I!!"’S!"6g 1{X*Y> Y> CS<"6mM2X*+02(6V.QC*?9X@.> M> NG"A L M> ?> CS<"6fJKxm[:X1*3(* 1*326@BCX> @&"!"6NEX*/&G=#"A6_)X*.> /> -G L Q> .> [!"6i%kxs56X*3(36 V.QC* @665X> ,= F[’:66q5)vX !"##%&’#& ()#* I K!J> "+*,3&4%&)%6E!"A L
I> M> @ E> FG" L .> M> NG"A*H#%O>

"#K>
.!’G#"%> !"##%&’#& "+*,3&4%&)% [> ?> E!"A
L I> M> @< G" .&UU > .#P > R<’ >6@!JOG"X)6,Xt3*
XGA > 3T ,2()T BM?9tCSG"!> /&!"A=#"A6Q)X*
@&G!"A6F[X*.!G&"\S!"A60*7X*,2)0 8 +)
80* B> I> 2)+* ,026 S#U#P;<* ?9q@ G’#P;<*
456qX>
0)vUKo)vc{[qj3&rO*@ F*6Jf;NQ;o 1({Q)F[o /&!"A=#"A6Q)X*@&G!"A6F[X*[> B>
B’!"A6op7X,026V.QCX>
,< GP[’:66q5)vX
!"##%&$’#& ()#* I K!J > 8#-,’38"*#*%+.
6B!%XD> E> FG" L .> M> NG"A*H#%O>

"#K >
.!’G#"%> !"##%&’#& 8#-,’38"*#*%+. ?> C>
B!% G" .&UU > .#P > R<’ >6@!JOG"X063Xt(3*XGA > 3T
,2(0

RS#=#=<"=J#" Q> CSG"!t*1 L
,+7*;U > 07t1 8 ,0T ,2()T BM?9tCSG"!> E&]G!"6I
cX*CS#"A~!"68?X*DG"AH&"6cWX*,21* 8 +3 8
+)*-> 4> -<<6ityX1*,+)6 S#U#P;<*EI?@ G’#P;<*?9qX>
0)vUKo)vc{$[qj3&rO*6
JE;o
za{IcbJ{o
9 0 N2’:23v?!l#h6za
EGA>0 BS< A<#AJ!;SGH =G’PJGO&PG#" #X ! > ()#*
!"= GP’ G"PJ!’;! N2’: ! > ()#* K!J > P’#G@"-VE2 ! > ()#* K!J >
&+*)(,-+.@#GP[’: ! > ()#* K!J > 8#-,’38"*#*%+.@AD
’: ! > ()#* K!J > ",8#/-%.3(#)+.@%F[’: ! > ()#* K!J >
"+*,3&4%&)%
E&]G!"6IcX*CS#"A~!"68?X*[&G’S!" 9:;<=>6bJ{mU#X03,)6V.QC*E65X@WH!=> QHG > 9:;<=> 6jm#X 12++7+* 12+,0** 12+311 7) & ’ ( ) 3*" 6!"#*!%X*&’())*6!"#X@+, -, +./06&-yX
1()2)6!"#X@+, -, +336ityX&41()*&41(&
6!%X@5, 6, 7.8/06AYX9 , / ,6!%X@, 7,
:8/0 ; 8< ,6k*"?@X*142)*
1’)26"?@X@A, 5, B./0 ; C, @, 6D8/06fJKx
DR9XE122*E124*E12&*E12E*E12’6CFGX,
!"! 7:)xh6za:*N2’:23v?!lF_1
6{3Ko6h*{*AD’:16{AB#‘a{XE*GP[’:
6{IcbJ{*F[’:6{Q)F[o!-V
E2zaLb@Q*$Q)s*QB)sx
NE)Es*6Iz!l#zae(9 =o
s t , -
lFd*Ts0 ,1’E) ,’:2B#VW ,u0VW*)61Xt1 H EI
[& , =((2 ,<7’:2B#%I69< ,u0z!))*2=6*Xt 4** H 4&(I Ts0 , 1’E& ,<7’:2B&6v ,dWF)))tG_()^*=2 61XtE) H E&I Ts0 , 1’’2 ,4{RCBJJ’:;*ls< ,<7u0Y ,>?t() 9^_*)’= H 21&, OP5 , 1’E=8,’:HY ,u0VW*=62Xt&& H ’EI OP5 , 1’E=K,QB’:2VW6XX,QBu0*=6=Xt*’ H &*I OP5 , 1’E) ,QB’:2VW6&X,QBu0*)6)Xt1&& H 1E&I OP5*n3j*r=r*F ,1’&E ,Q)’:B*Du0VW ,Q)> *g3*62Xt)) H 21I GD8LK3M<8./ B !*N83 @ 7, 1’’(I ? M3O.9.P/ PQ !"##%&’#& RS 9TK03/T9
()*+)*,), , >U./KTM0D 7PTM/8< PQ APV8/W*2&tE’ H =((I
C3 5*GD8LK3M<8./ B G, =((4I !"##%&’#& 9TK03/, ()*+)*), JJ:T G
5*N8O3/ % C, !.99PTM. APV8/.Y8< Z8MU3/ %M399, 7./ [ !*B./0 A 5*7./ @ C*%+ -. , =((&I N3O.9.P/ PQ 9PL3 \MPK<3L8V.Y V8]8 PQ !"##%&’#& 93YV , ()*)+)*),6>M.Y8Y383X QMPL GD./8, ?//8<39
APV8/.Y. !3//.Y.*22t1E H =2I
3MM.<< > B, 1’)2I #/M3YPMU3U \<8/V9 QMPL -^8/0VT/0 %MPO./Y3 RRR , +./0/8/ @Y.3/Y3 7PTM/8<*1)t14 H 4=I !"#% @AB)
&)! &# Si7Ft<7k%u0N2’:6’:2(X23u;!l#z!

  鄂ICP备06018747号 Copyright @ 2017
  植物通 All Rights Reserved
  Email:23198511@qq.com
回顶部
51La