免费文献传递   相关文献

Differential Expression of SA and H2O2 between Susceptible and Resistant Poplars Challenged by Botryosphaeria dothidea

SA及H2O2在杨树与溃疡病菌互作中的表达差异



全 文 :!!"
" !
#
#
$ % # &
$
#
%
& ( )
()*+,)- )./-* )+)(-*
/012!"!+02#
3456!$ % # &
708"#%6##9%9:;6#%%#<9!==6$%#&%##D
*+,#
" $%## >#$ >$%#
-.,#
" $%#$ >%= >#%$
/012
"
34&ŸmyQ(lØ112
%$%#$%!?%#&#
34(5®/@yTOØ1
%$%%"PB$#%#%%&$
!“p"LMNOP
$
-
6
d
$
e
$
0&|ghй­Og_‰</
!
²
E
%
#
E
,³´
$
E
µ
E
‚
#
E

E
·
#
E
q5Y
#
E
$¸¹
#
%#2
g3&()lmdü&jÚÕÖ|Üòlmn
E
34&ü&jÚ)|ü&Üò)”ú~sÕ
E
Ö×
#%%%"##
$2
àpbc)ÍkÜò)d
E
£f
$9#%#=&
!
E
"
! E
ÏBµ0zÖ×0Lrsz{
!
lm€0&RQghй¡‰Û¯Ø´0x
%-&
zoQ5õ
%d
$
e
$
&
_u5/
$
‡ˆ‰Š
"
Bµ0RQ¡
-
|
d
$
e
$
§š¨©}F
!9$ M
–IrA×
-
§š<’dä
%#"#2!#
!
H(H
>#
PN&!
iñœ
d
$
e
$
§š_Ÿªz
(-,
RVy_‘‹
#
Ö×0RQ¡IrA×
-
§šu5€¨©
%
däîL
92=&
!
H(H
>#
PN&!
€ñœ
(-,
Vy_V‘
!

d
$
e
$
§š|Bµ0»*<’dä_–Eª
!
dä‹
%
¯#L
9&92?$
z
&%D2"" 5G01(H
>#
PN&!
5Š€0&|ghй­O0¿g
-
|
d
$
e
$
/¨©
#
–
!
ðké6/ÂBµ0´0
x‡ß>µ
YV-@Y
zÖ×0
YZ-@Y
>µ_
$
Ç/û
!
Á/xo~*Z¯uŠ
!?*,3:?
z
?C,3:?
/Â_Á/x
|ýõ
%+V-@Y&
Á/x¹yL
"%c$ m,¯u‰Š
"
RQ¡Bµ0
,3:?
/û_‰<
D q$! M
–‘’•–
!
©åü–
-
|
-@Y
‡ß
!
•–
-@Y
_
(-,
Vy
!
e#
$! M
¡
d
$
e
$
§š_¨©F
$
ÐrsŠ
"
€0&
RQghй¡
-
|
d
$
e
$
u5/¨©
!
0&ZghÐ_˜y|²0I
-
|
d
$
e
$
_|\-ú½6
$
#$%
" E
0&
#
ghй
#
´0x
#
oQ5õ
&()*
! 9D&2#$ 9D!EEE
+,-./
! -EEE
+01*
! #%%# >9!=="$%#&#%# >%#%9 >%9
N/"#+#,%/6.!H4+#&&/,"*56,-d
A
A
)#%8##,*E&$#4%/).#6,-L#&/&%6,%
(4.6+&D26.#,0#-)3 F0(+&0)4$1+.$ -0(4.-1$
f43845
#
E.8O AMR5\O
$
E.{ oO45
#
E.845H3O5
#
EB45 C05HMO8
#
EAM45HK85H\40
#
%#2A$)$1(./ 92)*-*0*$"&"($)*<."#"%+! <25-("2;$2*128 ?("*$.*-"2! !3EB$+@1C"&"($)*<."#"%+
128 "($)*?("*$.*-"2 "&,*1*$"($)*(+38;-2-)*(1*-"2E:$->-2% #%%%"## $2!"#$%$"&?#12*?("*$.*-"2! ,/128"2% 3%(-.0#*0(1#
42-5$()-*+ED1-)12 $9#%#=&
5)&%+6$%" E/4U84V805[85 [418X\18X4X87 % -& 457 M\7U0HR5 TRU0S87R%d
$
e
$
& R^UR4541\QR7 85 78WRUR5VUR[8[V45XR
T0T14U[%?"=0#0)*";$2*")1 457 ?"=0#0)pC$->-2%$2)-)& 850XO14VR7 8^VM :"*(+")=/1$(-1 8"*/-8$16,MRUR[O1V[[M0^R7 VM4V
VMR-X05VR5V85XUR4[R7 85 ?7*";$2*")1! VMRX05XR5VU4V805 0WR570HR50O[WURR-TR4IR7 %#"#2!#
!
H(H
>#
PN& 4V9$
M! M^8XM 4^[4XX0GT458R7 8^VM VMRUR7OXV805 0W(-,4XV8]8V\457 85XUR4[R85 d
$
e
$
X05VR5V6d0^R]RU!85 ?7KC$->-2%$2)-)
VMR-X05VR5V787 50V[8H58W8X45V1\4U0[R%96=&
!
H(H
>#
PN&! 457 4^[50V4XX0GT458R7 Z\7RXUR4[R[85 (-,4XV8]8V\457
d
$
e
$
X05VR5V6,MRd
$
e
$
TR4I 0XXOUR7 R4U18RU^ 8VM 10^RU]41OR[X0GT4UR7 8^VM ?7*";$2*")1 %&%D2"" 5G01(H
>#
PN5)
9&96?$ 5G01(H
>#
PN&6,MR4Z0]RUR[O1V[8578X4VR7 VMRUR^ RUR[8H58W8X45V78WRUR5XR[85 -457 d
$
e
$
85 78WRUR5VT0T14U[
85VRU4XVR7 8^VM :78"*/-8$16fR45 M^81R! R^8[014VR7 457 87R5V8W8R7 WO1<1R5HVM X0785H[RLOR5XR0W?*,3:?457 ?C,3:?
WU0G?7*";$2*")1 457 ?7pC$->-2%$2)-)[VRGXC+-[6,MRTU0VR85[0WYV-@Y457 YZ-@YZR105HV0VMR(4V414[Rw18IR
[OTRUW4G81\! 457 VMR8U[RLOR5XR[4UR"%c 87R5V8X41V0VM4V0WV0Z4XX0-@Y4VVMR4G8504X87 1R]R1Z4[R7 05 +(@)Z14[V
457 TM\10HR5RV8XVURR4541\[8[6m,850XO14VR7 8^VM :78"*/-8$1! [OHHR[V85HVM4V- 4^[Z0O57 8^VM -@Y4VVM8[V8GR! 85M8Z8V85H(-,4XV8]8V\! VMO[
TU0G0V85H[8H58W8X45V85XUR4[R85 d
$
e
$
X05VR5V4V$! M 4WVRU850XO14V8056,MRRSTRU8GR5V[7RVRUG85R7 VM4VVMR78WRUR5XR0W
X05VR5V0W-457 d
$
e
$
85 T0T14U[^ 4[7ORV0850XO14VR7 8^VM :78"*/-8$1! ,3:? 4^[85]01]R7 85 VMR78[R4[RUR[8[V45XR0W
T0T14U! 457 [OHHR[VR7 VM4VVMRUR[8[V45XR0WT0T14U[V0:78"*/-8$1 0^O17 ZRUR1R]45VV0VMR4XXOGO14V805 0W-457 d
$
e
$
6
7#3 8+-&" E?"=0#0)[TT6# :"*(+")=/1$(-1 8"*/-8$1# [418X\18X4X87%-&# d
$
e
$
& ( )
!"
"
E
EE
0&
%?"=0#0)&
ghÐ
%:"*(+")=/1$(-1 8"*/-8$1&
$)30&_ó7êWÐj
%
J×/å
! $%%&#
ù
!Kå
! $%%&#
ž,¹å
! $%##&$
0&ghЭO
0¿_lmc̅g¹´µW¹ùÚV_­
O£žlm
!
Z°0&ghИЉÛ1ð¿Û_
O?£–_û_;€Ì
$
œ3lm5Î|ÉÊ_Ž
a
!
3;)PZýõ
%L-."*-121 *1C1.0;&
|Ù·
%3(1C-8"=)-)*/1#-121&
åõÐÍk|Ðþ¹­O_
‰Û1ð¿ÛŽQ_™Ìlm
!
iM!_P8Ík
ΫÐþ¹_O?áU
%/10V$*1#7! $%%=&!
P8l
mŠ´0x
%-&
|oQ5õ
%d
$
e
$
&
$Ík˜Ð
Š>g_”7‰Û¯Ø
!-
‘0>µ
% -Z85785H
TU0VR85! -@Y&
|oQ5õR
%X4V414[R! (-,&
¼½
ƒ_ry
!
f½lmŠýõåÍk0I
-@Y
¼½
(-,
Vy
! -@Y
|
-
‡ß¡²
(-,
Vyd•–
!
evÍk0I
d
$
e
$
´µ
!
‰V˜
л6/û_‰<
!
Ž‰·¸˜y
%(MR5 $*1#!
#""## -]RU\450]! $%%"&$
-
z
d
$
e
$
Íkۊ>gO?_lmcÌ
s°bOkÉç
!
Z&"|Ðþ¹­Og_lm€
Ì
$ C4]8[
å
%$%%$&
lm‰Š
!-
ŒÏªð±&
%?-20)[TT6&
å"ÐÍkgΫ/û_‰<
!

-
Œg&"Ϋ¿Ûg=i†3”7Vw
$

!
l
mÌ6Á0&
-
‰Û1ðè³
!
‡ˆ‰Š;Ò
-
ŒÏªð0&˜Ð/û_‰<
%F0XM $*1#6! $%%%&!
P
|ýõg_lm‡ˆÜݹñ
%kI5R[$*1#6!#""&&$
m;
!
lm‰Š0&Ir
-
§šC°ýõz|
Ù·g_§š
%F0XM $*1#6! $%%%&$
ûü
!
½k7Z
0&˜Ð‰Û1ð¿ÛŽQ>?lm
!
ÐNë]B
µ0
% ?"=0#0)*";$2*")1 &
z Ö × 0
% ?7 p
C$->-2%$2)-)&
Lrsz{
!
Molm²
-
|
d
$
e
$
§
š_u5
-@Y
/û_ðk6/û‰<
(-,
Vy
u5€0&g_‰</
!

-!d
$
e
$
‰Û
¯Ø6
-@Y
$yð°0&|ghй­O¿Û
g
!
LŽ¹aŠx0&
>
ghй­Og_‰ÛÔ
ð£–_lmvwxy
$
#E
z{|ÉÊ
9:9;
V2¾˜
&
¿…]À:
wr0&LBµ0
%
›ªùÚV”Ð-ÀL
% qD2%&
Ö×0›ªùÚV”Ð-À
!?2#
Ï}
!
%
0¼$å
! #""%#
¤må
! #""D&
_
#
$jpq—
S0&^
$
wr¹ÊLÐ~sÕeÖ×0ghÐM}¯×
iÜð_0&ghÐÐþ¹
!
nԍ;Ã@º
!
ñÐ
Dgå¹Ê
$
¹Ê
$9 b
&V°
YC-
µ/}a

9
™
!
ˆ¹&V?¸¹›4˜¹ßý§ì
è³L
? GG
_¹kOLRQ0÷?
$
k¶¹kRQ
"
¥®RQ®ÿ?
9?c
=ßlm
¶·
!
¿?ˆ¹´lm
$
ï
!
?¸¹ûRkÙ¶~·
UkÍ
!
»æÙ
& GG!
ª¡l½¹b¹ç…bkÍ
n1
!
;dƒ¸¹ö;
!
?Üíd¯oHºRQú
$
½&QÑ
$
ǝž
"
k¶RQ¸¹
YC-
;Ã/
%(F&#
k¶RQghй
$
~ҝž¡
% %
ˆk

&!D!#$!$!!!=!9$
z
"D M
¯#Z×ÐM
%2? XG
&KŽQì¨
!
½ž½–Eú”Å
&
B
$
ì¡
„%p?µ¦g
!
iÜð°
>=% b
:òg÷?
$
9:A;
½¾”
"*5#
º\=¯°
ë?
/RUZRU5R
_ÉÊZ¨©g_
-
§šŽQ
åº
%/RUZRU5R$*1#6! $%%$&$
âì
# H
¨©
!
ª?˜
Q5zޚ_
=%c
Ðß
!
‹Vl¬h°q
$
Ò
°
% q! b
lv
!#$ %%% U(G85
>#
×®ì}ºµ
$
l
r?
=%c
Ðß¿Bvì
$
B
!
ßi}ºµ
$
l}º
µµ¶¯h
$
ê
!
¯#?°A×´0x
%PURR
[418X\18X4X87! P-&
z›´0x
%,0V41[418X\18X4X87!
,-&
_åº
$
}ºµ‚fó´»
!
ŠÅ¸²
!
×®
!
ª?
Y/Y
Íéwù
!

$ #
ꚵª?å0|_
$ GG01(.
>#
d(1!
¯°¡1?rÑ
!

!=% b
Õ
# M!
#›´0xg‡ßÚ_´0xxûLA×´0
x
!
Ï帲g›´0x_§š
$
ª¡
$
ꚵ
g¯#ª?
$ G.?c
¥s=x
!
?
& G1
=x=¾
%/&g
ÕtÑ
%/& h#g#
lv
&
B
!
ßi½£»
!
‡ƒ

$
Ï
#2? G.9%c
Ðß´ûlr
!? G.9%c
Ð
ßµ`
(#=
£Ò¡
!
¨©oÒ
!
?
& G.9%c
Ðß»
Y£Ò
!
*…»Yµ
$
»Yµ‡ƒWã
!$?% O.
E
Ø»ÓZµ´ûlr
!
o
%2!?
!
G
4šd
!
ª¡
?Æµ»w·
%dY.(&
)娩g
-
§š
$
dY.(
ð˜UÛL
N4VRU[$D"?![O5W8UR(#=
Ò
!
E%
%2= G.(G85
>#
!
ܾ–E
#%2"!& G85$
9:<;d
A
A
º\=¯°
å‘
Y4VRU[05
å
%#"=!&
_ÉÊ
!
¿½tØ
$
ì
# H
¨©ª
& G.
C!¿¬h°q
!#$ %%% U(G85
>#
×
®
!
}ºµºk
!
Š>µg§
%2# G.$%c %\I\&
,8(1!
_‚«x
%2$ G.
‚Á´z
# G.
}ºµjh
_oQ5k
>,8
Åßk
!
?!¿»
?
B
!
!¿ú“
¡´°
& G.# G01(.
>#
¡xg
!!#% GG
Våÿ±*
ä
!
¢¨Ýo–
d
$
e
$
.xž‘
$
9:B;
Á˜AÂÃ
"D51!D9:99:9:S#
>B¯°
RVyåºÉÊå‘g3()d}qÍkjž
lmn
%#"""&
_oQ5õRåºÉÊ
!
¿½tØ
$
=%#
E!
#
# ¦
E

" -
6
d
$
e
$
0&|ghй­Og_‰</
¯#â쀁ž_z{
%# H&!(-,
v쵪?
%2$ G01(.
>#
§xñ{µ
%Td92=&$
¿¯#ª?f
™N¾¨z*=ÍuvÑ
%Y/YY&!
:Õl¬h°
q
!! b #= %%% U(G85
>#
!
×®
$? G85!
ì}ºµ
!
ý
Rvìµ
$
Ï
# G85

#
Ç
eC
äL
(-,
RVD
Ôÿ
$
9:R;
>~ÄÅ6
L#6.C%/?#(DL

#2?2#E
¿
m+-
2‚h
XC+-
ÀÁÂ2{ª
E
m+-
vìrw
%@80VRIR
Ÿ(
&
v쨩_›
m+-!,4F4m4
_Š1qrwŠ1qh
XC+-!
ÉÊ
¢•5ŠžŽQ
$
lné6_
m+-
z
XC+-
¯#
Üð°
>=% b
z
>$% b
÷?
$
#2?2$E
ÃK
,3:?
~ÄÅ
E
Ðrs¤y
*f@.<
@45I
}f“‰_Bˆ0
%?7*(-./".1(=1& ,3:?

o~
%-XXR[[805 5OGZRU[" (f%%%&&=&
Ñ8/û
emP
o~_qk
!
}Aqk
# %ll-,(((,-(--
l,-((l,((-,&
zVAqk
$ %(-(l,,l-l-
(l-l-,l(,-l(,&!
ϊ1qUkLá/
!
ŽQ_
Y(m

$ Y(m
Š>0¿L
$?
!
.!
¯§
#% pZOWRU
$2?
!
.!7+,Y%$2? GG01(
!
.
>#
?
!
.!
}

VAq
k
%#%
!
G01(
!
.
>#
&
—
#
!
.!,4L
*ßR
%? k(
!
.
>#
&
#
!
.!
›
XC+-%#
!
H(
!
.
>#
&#
!
.!77d
$
e
ó
$?
!
.$
Š>ÝoL
"! b
muy
? G85!"! b
uy
&% [!
?= b

!% [!9$ b
JK
?% [!&%
ǒա
9$ b
J
K
#% G85$
Y(m
U k > ?
T*-B<,& (10585H /RXV0U
%,U45[lR5 @80VRXM! @R8;85H! (M854&!
ÁR
!
15é6
o~
!
o~åo
%)5]8VU0HR5&6
#2?2&Em,Æ:ÇÈWÉÊ
E
0&
,3:?
_o
~
%lR5RZ45I 50" 3o$!=?9#&
z
+(@)
}/_0&
%
>D0C0#-2 %-XXR[[805 5OGZRU[" (f%%%&&92#&
OL

!
¯#Ù?,
YU8GRU?2%
Ñ8ÿºš
Y(m
qk
!
¢‰
#$
9
9;
pqƶX[
16):9;(+/?#+&#TE#,$#&E&#-/,%2/&&%E-3
qkNâ
YU8GRUX07R[
qko~
%?) >&)&YU8GRU[RLOR5XR[%?) >&)&
,OZ185 ?) >l(-((--(,,l,,l-l--,l(>&) ?) >l,(,(-ll(,l-,---,(,,,l>&)
,3:? ?) >,,,(--(,l-((-lll--((>&) ?) >l,,,(,l(--(,(ll--,-,,l>&)
#2?2!Em,ϊ1qé6_
XC+-
ÛÜ
$%
|Oá/
!
¢V~gê7–
Y(m
Š>µ
%
‰
$&$

mR4HR5V
#?š
-TT18X4V805 4G0O5V:
!
.
B*mTURG8S*S,4L
,f
%$ p&
#$2?
Y(mW0U^4U7 TU8GRU%#%
!
f& %2?
Y(mUR]RU[RTU8GRU%#%
!
f& %2?
meKURWRU5XR7\R%?% p& %2?
C+-
á/
$2%
Cd
$
e
"2%
,0V41 $?2%
!
.
ÐBrs#?
,-F-m-
Ÿ(
B@m
+
?($;-Q

$ mR41<,8GRY(m
Š>ÝoL
" "? b
muy
&% [#
RVS$
!%
ǒ
Õ
!"? b
uy
? [!D% b

&! [#
½Ç¨©
&

Å
!Y(m
Š>‡%¡ŽQ€ûž‘¯u
$
9:S;
Ç=>~=ÈÉ(÷
Ù?
C+-[V4U!YU8GRUYURG8RU?2%
,Z2_

XC+-
/ÂÁ/xo~ŽQ¯u
#
?
+(@)
À
y›g_
@14[V
,l2_/û/Â_Á/xo~
|
+(@)
g_o~ŽQ*Zz»>yK˜
#
ŽÙ
?
(1O[V41K
z
lR5RC0X
ŽQÌo~ry*Zz*
T¯u
$
$E
‡ˆ|¯u
A:9;
VW¾˜À:
F<-0(4.-1$
´
*5
 
d
A
A
º
\xA
$2#2#E
ËDÌ
-
›1̈́
E
RQ
:78"*/-8$1
¡
!
Mo
dY.(
庇ßÚ|A×Ú
-
§šu5
‡ˆ¡$
#!$$
Bµ0zÖ×0RQ¡A×Ú´0
x§š¶F
!
˜ÐBµ0A×Ú
-
e
D M
9Z~
F
!9$ M
<’dä
%?i%2%%% #&!
ª¡
"D M
9
ZV‘
#
Ö×0A×Ú
-
RQ¡
$! M
9Zñò}
F
!
<’¬
!
}Fvƒ*Bµ0£
$!2?!
|
#

ãRQ
:78"*/-8$1
_Z•g
!
Bµ0zÖ×0_I
rA×Ú
-
§šu5i€c
$
P5Š
-
§š_
u5$4°RQ
:78"*/-8$1
q@_
!
VA×Ú´0
x§š>
!
Bµ0*Ö×0}Fvƒc–Eª
!
P¹ú|Z•½Š¨c#
$
RQ
:78"*/-8$1
¡Bµ0zÖ×0g‡ßÚ
-
§š¶½}F_9:
!
²gBµ0RQ
$! M
¡
´0x§<’¬ä
%?i%2%%#&!
Ö×0RQ¡u
5€Š¨
!
Z•gBµ0zÖ×0_‡ßÚ´0
x§šu5€¨©
%
$
$&$
"%#
& ( )
!"
"
E
$
#E
A×Ú´0x§šu5
P8H2#E(M45HR[05 WURR[418X\18X4X87 X05VR5V
!‰í¹–EI
!
Ö×0zBµ0ÉE_/
!
!L
?i%2%?!
!!L
?i%2%%#!
!!!L
?i%2%%% #$
-[VRU8[I[8578X4VR[V4V8[V8X41[8H58W8X45V78WRUR5XR[%
!
h?i%2%?!
!!
h?i%2%%#!
!!!
h
?i%2%%% # VO7R5V)[*VR[V& ZRV^RR5 1R]R1[85 ?2p
C$->-2%$2)-)457 ?7*";$2*")1 W0UR4XM V8GRT085V6

$ ,MR[4GRZR10^6
$
$E
‡ßÚ´0x§šu5
P8H2$E(M45HR[05 X0GZ85R7 [418X\18X4X87 X05VR5V
$2#2$E
ËD
:28"*/-8$1
Ì>?ÃK¶
d
$
e
$
›12
ÎÏ
E
RQ
:28"*/-8$1
¡
!$
Q0&
d
$
e
$
§š }
F¡V‘
!
í«PÉE_u5_vðŠ¨_
/
!
Bµ0¨©°Ö×0
$
Bµ0
9$ M
–<’
d
%?i%2%%% #&!
Ö×0
!= M
–<’d
!d
$
e
$
§
š¯#L
&%D2""
z
9&92?$ G01(H
>#
!
*»>
(F
¯
#ٻ
&2%=
z
#%2%&
|
!
Z•Bµ0zÖ×0
d
$
e
$
§šu5€c
%
$
&&$
A:A;
¾˜½¾”ur†j>~
"?5F/C*6./$3./$
6$/-C)/,-/,0 4+%#/,#
Ê>”ÈÉ(÷
lðk_0&
,3:?
/û
+(@)
}ÁŠ
!
lR5@45I
/qÛ¯#$
3o$!=?9# %?2*";$2*")1
YV-@Y&
z
3o$!=?9$ %?2pC$->-2%$2)-)YZ-@Y&$
Z/û©ð_Á/xo~ŽQ
YU0VT4U4G
Ýom
å¯u
!
/û
emP
o~Ù
# !D$ ZT!
Æ/Â
!=9
Ç
$
&E
oQ5õ§š_u5
P8H2&E(M45HR[05 d
$
e
$
X05VR5V
Á/x
!YV-@Y
z
YV-@Y
»Z¯Øuš¯#L
?D ###2$=
z
?D %D?2$#!
ž G å ¯ ú L
D2?D$
+V-@Y% L-."*-121 *1C1.0; -@Y lR5@45I 506
--(!="#=2#&
»Z¯ØušL
?D &DD29$!
žGå¯
úL
D2?%$
#?
lR5RC0X
,¯uýõ_
-@Y
>
µÁ/x|nðk_Bµ0zÖ×0o~©å_Á
/xŽQ*¾
%
$
!&!
ÍwcFLƒÜÀ_»
Á/xo~
!
µwcFL®¯»Á/xo~
!
µw
cFL/_Á/xo~
$
Bµ0zÖ×0
,3:?
/û/Â_Á/x|ýõ½ÿê_ry
!

õÁ/x¹yL
"%c!
È°
(4V414[Rw 18IR
[OTRUW4G81\$
A:<;F:-0(4.-1$
Ë̾˜´
?5F/
>~9Í6
D51
>BxA-
$2&2#E ,3:?
~¦-ÐÑ1†‡
E
RQ
:7
8"*/-8$1
¡
!
Bµ0
,3:?
‰<š»Z°Ö×0u5
¨©
$ ,3:?
Bµ0RQghй¡‰<ši
°}w9:
!D M
–‰<š}F
%*hD29&!!?h
%2%&&!
LZ•_
&2=%
|
!
œ¡
#$!$! M
‰<šV
‘
!
í
!= M
–‰<š„%}Fi<’¾ä
%*h
##2"$&!?h%2%%% #&!
LZ•_
##2#%
|
!
¡‰<
‘’•–
!
‰<š~‘‹
#
Ö×0g
,3:?


9$ M
Çu5€Š¨
!"D M
–Ö×0
,3:?
‰<
šT½‘’•–
!
qª-·}F9:
%*hD2&$!!?h
%2%%&&!
LZ•_
&2=%
|
%
$
?&$
$2&2$E
ÒӄÔÕ
!(-,"
_g2̈́
E

#
Œ
2
"
ãRQghй_Bµ0zÖ×0
(-,
Vy
n庖EIˆ¨©u5
!
Z+°Ý¹Ç¾‹
µŸ_´µ
$
RQghй¡
$
Q0&
(-,
Vyu5Š¨
$
²g
!
Bµ0
(-,
Vy- }F¡V‘_9:
!
R

!= M!
Bµ0
(-,
VyŸ3i<’dä
!
V
%##
E!
#
# ¦
E

" -
6
d
$
e
$
0&|ghй­Og_‰</
EEE
$
!E
Ö×0|Bµ0
-@Y
Á/xo~_*Z¯u
P8H2!EfO1V8T1R[RLOR5XR418H5GR5V0W-@Y0W?7*";$2*")1 457 ?2pC$->-2%$2)-)
Bµ0
%?7*";$2*")1 YV-@Y&
Ö×0
%?7pC$->-2%$2)-)YZ-@Y&
|ýõ
%L-."*-121 *1C1.0;+V-@Y&-@Y
Á/x_o~*Z
¯u
!
ÍwcFLƒÜÀ_»Á/xo~
!
µwcFL®¯»Á/xo~
$ fO1V8T1R[RLOR5XR
418H5GR5V0W7R7OXR7 4G8504X87 [RLOR5XR[0WYV-@Y%?7*";$2*")1 -@Y& 457 YZ-@Y%?7pC$->-2%$2)-)-@Y& 457 VMR
TU0VR85 [RLOR5XR0WL-."*-121 *1C1.0;-@Y%+V-@Y&6)7R5V8X41UR[87OR[4UR[M47R7 85 Z14XI 457 [8G814UUR[87OR[85 HU4\6
$
?E
RQghй¡€˜y©Qg
,3:?
/û‰</
P8H2?E*STUR[[805 0W,3:?0WT0T14UXO1V8]4U[[O[XRTV8Z1R%?2pC$->-2%$2)-)&
457 V01RU45V%?7*";$2*")1& 4WVRU:78"*/-8$1 850XO14V805
yL
!92&&
!
G01(GH
>#
G85
>#
!
$Z•_
#&2D%
|
%?i%6%%% #&!
œ¡„%V‘
!
Ïó‹°Z•
#
Ö
×0RQ¡
(-,
Vy-}F9:
!"D M
<’dä
!
L
#"2=D
!
G01(GH
>#
G85
>#
!
$Z•_
&2"$
|
%?i%6%%% #&#
Bµ0|Ö×0»*¾
!(-,
Vy
RQ¡
#$ q!= M
‰·¬¨©y
%?i%2%%% #&!
!= M
¡Bµ0
(-,
Vy‘‹
!
Ö×04Ý,}
F
$
5ŠoQ5õR0&˜ghйoÝgð
¹º_/
$
&E
‡G|HG
0&|ghй_­O£–$¹ÇƯÅô_

!
€î¯§0ó8Ú

«

jžj5z¯Ø
###
& ( )
!"
"
E
EE
9
9;
VWÀ:Îý´ÏÐÑ¿NVW‘B¾˜&Á˜AÂÃ>B=TU
!
16):9;!"#$%"F<-0(4.-1$ ,D51#,K3?#&6$%/M/%3 "44.6+6%-/"#+#,%%/?#6"%#+/,$E.6%/,
RQ
)50XO14V805
–E
,8GR:M
Bµ0
?7*";$2*")1
Ö×0
?2pC$->-2%$2)-) (-,
Vy*ä
m4V800W(-,
*V
TRX8W8X4XV8]8V\:
%
!
G01(GH
>#
G85
>#
&

m4V80
*V
TRX8W8X4XV8]8V\:
%
!
G01(GH
>#
G85
>#
&

m4V80
Bµ0
:
Ö×0
?7*";$2*")1I
?7pC$->-2%$2)-)
Bµ0
(F:
Ö×0
(F
?7*";$2*")1%(F&
?7C$->-2%$2)-)%(F&
RQ
)50XO14V805
%
&2#&" s%2?$&
#2%&!4
&2!"? s%2$9%
%2"$?4 %2="94 %2=%&4
(F &2%&? s%2%$9 &299? s%29&"
RQ
)50XO14V805
D
!2%?# s%29&&
%2"==4
&2%=" s%2D%%
%2=$%4 #2$"D4 #2%9?4
(F !2%%D s%29&D &29DD s%2!$$
RQ
)50XO14V805
#$
$#29== s%2?!$
!!!
!2"?$Z
&2!D! s%2&&=
#2#=#4 D2$="Z #2?%%4
(F !2&"" s%2=#" $2"&# s%2=!=
RQ
)50XO14V805
$!
$92$=# s%2#&%
!!!
?2==DX
"2?$? s%2!#9
!!!
&2%"$Z
$2=D&X #2?%!4
(F !2D&! s%2D"& &2%=% s%2?9D
RQ
)50XO14V805
!=
!92&&% s%2D$=
!!!
#&2?"#7
##2"#" s%2$9$
!!!
!2&"#X
&2"9%7 #2$=$4
(F &2!=$ s%2!#% $29#! s%2&%!
RQ
)50XO14V805
9$
##2%"$ s%2%?D
!!!
&2&$9R
#&2?D& s%2="=
!!!
&2?#%Z %2=#94 %2=D$4
(F &2&&& s%2$#$ &2=D& s%2&&?
RQ
)50XO14V805
"D
#2?!! s%2D=D
%2D$#4
#"2=?" s%2$?D
!!!
!2==9X %2%99R %2D994
(F $2!=! s%2D!% !2%D& s%2&&#
EE
%‰gÀyLµ¶ä
s
.x¼
#

h
RQR*VD
:
»>Z•R*VD
#
,‰í¹–EI
!
ž|Z•ÉE_/
!
!
?i%2%?!
!!
?i%2%%#!
!!!
?i%2%%% ##
~€£hiډ퀁–EžÉE/¨©
%?i%2%?&$ ,MR74V44URH8]R5 4[GR45[s*# U4V80h
850XO14V805:(F# -[VRU8[I[85 ,4Z2# 8578X4VR[V4V8[V8X41\[8H58W8X45V78WRUR5XR[%
!
?i%2%?!
!!
?i%2%%#!
!!!
?i%2%%% # VO7R5V)[*VR[V& X0GT4UR7
8^VM VMRX05VU014VVMR[4GRTU0XR[[85HV8GR# 78WRUR5V1RVRU[85 VMR[4GRX01OG5 8578X4VR[8H58W8X45V78WRUR5XR[4V78WRUR5VTU0XR[[85HV8GR%?i%2%?&6
å´µ_u5oÝ
!
;¯§3—Q‰Û_ԏoÝ
$

!
0&|ghй­O_lmó7…ggh
О)

Оjž)Ï6О5)}
%
bå
!
$%%?&!
Z²˜Ð‰Û1ð£–_lm¾f
$
0
ó|Ðþ¹­Og
!
´0x|oQ5õ‰Ûè³g
dÐþ¹n‰V
!
ŒÏOL‰ÛÔð¯Øªð¹¿
~/Âۊ>>µ/û_‰<
!
e‰·¸˜y
%(MR5 $*1#6! #""&# AM45H$*1#6! $%### N4U$*1#6!
$%##&$
Zýõ

|Ù·åõÐÍklm‰Š
"

k0I>gð¹&½Æ_
-
‰Û”2
:
‰Û1
ð£–
%AM40$*1#6! $%%"&!
\ÍÊdÐþ¹”Ö
!
-
jkßh¿Ûdªð‰V
!-
´µ¨Û}F
!
Ž
ÍÊ0I¼½6hš_
-
‰V‰Û1ðè³
!
¬+ðñÍʘÐgDŸ3
$
›ªùÚVP8Í
Ê0I
-
´µÿê‹
!
0&Ir
-
´µC°
ýõ

|Ù·åõÐÍk_
-
§š
!-
|
d
$
e
$
‰
Ûè³$y0&ۊ>g@3”7O?–7>
?lm
%l14QRZU00I! $%%#&$
Ðrs‡ˆ‰Š
"
RQghй¡
!
Bµ0z
Ö×0&Kg
-
z
d
$
e
$
§š¶F
%
$
# q&&$
Bµ0RQ¡
9$ M
–A×
-
§š<’dä
!
ü–
(-,
Vy‘‹
!
–ñœ3
d
$
e
$
§š_Ÿª
!
P|
ýõg_u5»¹ñ
%-]RU\450]! $%%"&$ (MR5
å
%#""&&
Šýõg_
-
‡ß>µ
% - Z85785H
TU0VR85!-@Y&
|
(-,
ƒr
!-@Y
|
-
‡ß
¡_¢_²
(-,
Vy
!

d
$
e
$
´µ
!d
$
e
$
W†_
ñjkVyQŒ‰V˜ÐŠ>è³g_˜Ð»6/
û_‰<
$
Ðlmg
!
MoZ
?*,3:?
z
?C,3:?
$
Ç/û
m,¯u‰Š
"
RQ¡Bµ0
,3:?
/
û_‰<
#$ q$! M
–‘’•–
!
#
(-,
§š‘
‹
!
–4°
-
|f½_
-@Y
‡ß•–_²
(-,
Vy
!
e#
$! M
¡
d
$
e
$
§š_¨©F
$

Ö×0RQghй¡
-
§šu5€¨©
!

0RQ¡
-
§šŸªÜ%
!
Œg$Bµ0*Ö×
0¼½“”_‰Û·Ø£–
$
ãRQghй_B
µ0zÖ×0
-!d
$
e
$
!(-,
n庖EIˆ¨©
u5
!
Z+°Ý¾‹µŸ_´µ
!
©å
,3:?
å
|0&_˜ÐoÝ
!
‰Š0&ghÐ_˜y>g
|0&0I
-
|
d
$
e
$
_|\-ú½6
!
Bµ0»Z
°Ö×0u5¨©
!
ÐrsERŠ
-
|
d
$
e
$

ÛO?Bµ0g6’)¯‰·
!
Ö×04g€
‰·PQÆ>
!
5ŠRQghÐg‰VBµ0_
-
|
d
$
e
$
‰Û1ðè³
$
©åBµ0d
:78"*/-8$1
J
á
!-
dªð‰V
!
#0&0I¼½6hš_
-
‰V‰Û1ðè³
!

d
$
e
$
OLI_gE‰Ûû
Ø
!
Mo´0x_þӑ0
,3:?
d
$
e
$
_§š
!
#;/‰ÛԏoÝg~dÄc
!
Ž
1q´µ}·Øzwˆ¹8†kÎv/û_‰<
!
¬+ðñBµ0˜ÐgDŸ3
$
í$˜ÐO?£–:œ0ó|Ðþ¹_€
ð3YÔz¯S
!
ÏähÅô_‰Û8x
$
ÀP
_rs“·
"
Bµ0RQ¡A×
-
§šŸ
!
‡ß
$##
E!
#
# ¦
E

" -
6
d
$
e
$
0&|ghй­Og_‰</
Ú
-
§š=F
!$! M
–<’dä
!
nÏA×
-
ªð&V_§šF€vS›Õ!цÉè³
z‡ßÚ
-
_15
!
½Œg$Ík0I|
-
ß
h»6_/ûd·Ø
!

-
‰ÛÔðè³_}A
Œg;ð8£–@O?
!

d
$
e
$
0&
>
g
hй˜Ð‰Û¿ÛgV€îî$@’!lj#_
O?
!
Œg0&‘’ghйØÃ_Q5y“Ó# 
‡
!(-,
¯û
d
$
e
$
!
jhQ¯Ø
!
¡“_ÕÐxjh
-!
ðñ‡ßÚ´0xŸª
#
m¹Éç
!
Œg$
-
‰
“
,3KD%
´0xо1ÈR
&
/û‰<
%d8TT4OW$*
1#6! $%#%&!
#
-
15L´0xо
%fR-!
¹Q‡
ßÚ´0x
&!
e#‡ßÚ
-
§šŸª
!
´0x
‡ß>µ
$%-@Y$&
/û_‰<—ŒÏ#
fR-
b
-
_15
%Y4UI $*1#6! $%%9# ,U8T4VM8$*1#6! $%#%&$
ûü
!
0&
>
ghЭO0¿g
d
$
e
$
$yv$i†
!lj#_VwÏ6$yð
fR-

-
_o
Ý
!
;½–Ž¹a>?lm
$
f g + ,
ù!K
!
J:(
!
/ 
!
å
6$%%&6
)3&"/ûTÝlmŽ¡z6
oÉF
6
&()
! &" %?&" ### >##=2
ž,¹
!
J×/
!
“p"
!
å
6$%##6
0&RQ
@0VU\0[TM4RU8470VM87R4
ghй¡>µu‰</¯u
6
Ù×&c))Š
"
ݻ()
y
!$%##!&?%!&" # >D2
b
E

!
żº
!
ž:?
!
å
6$%%?6
)30&|ghй­O_О
)lm
6
&()lm
!#=%$&" $#! >$$#2
¤
E
m
!
‡ˆ@
!
r
E
É
!
å
6#""D6
0&ghÐ0óªð˜Ðy_l
m
6
ÞÖ&)d)Š
!##%#&" # >!2
0¼$
!
+y.
!
ëz 
6
å
6#""%6
˜ghÐ0&QT_w&lm
6
Þ
Ö&)d)Š
!?%!&" # >#%2
J×/
!
{½‘
6$%%&2
g3ü&”cjk|j
6
Ö×
"
g3&¸y
û
6
g3()d}qÍkjžlmn
6#"""6
·†Íkjž)~s-Ù
6
Ö×
"
()¸yû
6
-]RU\450]-6$%%"6eS874V8]RZOU[V457 T145V78[R4[RUR[8[V45XR6PU05V
@80[X8%*18VR*7&! #" #!$ >#?$2
(MR5 A! F1R[[8HCP6#""#6)7R5V8W8X4V805 0W4[01OZ1R[418X\18X4X87<
Z85785HTU0VR85 VM4VG4\WO5XV805 85 [8H541VU45[7OXV805 85 VMRT145V
78[R4[R
=#=&2
(MR5 A! 81]4d! F1R[[8HCP6#""&6-XV8]R0S\HR5 [TRX8R[85 VMR
857OXV805 0WT145V[\[VRG8X4XLO8UR7 UR[8[V45XRZ\[418X\18X4X876
X8R5XR!$D$%?#!#&" #==& >#==D2
C4]8[3f! NO dl! (00IR3*F! $*1#7$%%$6Y4VM0HR5 XM41R5HR!
[418X\18X4X87! 457 ;4[G058X4X87 URHO14VRRSTUR[[805 0WXM8V854[R
HR5RM0G010H[85 T85R6f016Y145Vf8XU0ZR)5VRU4XV!#?%!&! &=% >
&=92
l14QRZU00I 36$%%#6lR5R[X05VU0185HRSTUR[[805 0W7RWR5[RUR[T05[R[85
-U4Z870T[8[.$%%# [V4VO[6(OUeT85 Y145V@801! !%!&" &%# >&%=2
d8TT4OWP! f8XM41[I\*! dO45Hm! $*1#7$%#%6*5Q\G4V8X! RSTUR[[805
457 [VUOXVOU4178]RUHR5XR[4G05HX4UZ0S\1eHR5R7OT18X4V805 457 [TRX84V805 85 +8X0V84546Y145Vf01@801! 9$
%&&" &## >&&%2
F0XM 3m! (URR1G45 m-! *[M8V4 f! $*1#7$%%%2eQ05R[R5[8V8]8V\85
M\ZU87 T0T14UX0UR14VR[^ 8VM 85[R5[8V8]8V\V0Z0VM [418X\18X4X87 457
;4[G058X4X876,MRU01R0WTU0HU4GGR7 XR1 7R4VM 85 1R[805
W0UG4V8056Y145VYM\[801! #$&%$&! !=9 >!"D2
Y4UI N! F48G0\0*! FOG4UC! $*1#7$%%96fRVM\1[418X\14VR8[4
XU8V8X41G0Z81R[8H541W0UT145V[\[VRG8X4XLO8UR7 UR[8[V45XR6
X8R5XR! &#=%?=!9&" ##& >##D2
Y4VRU[05 @C!f4XUR4*-!PRUHO[05 )@6#"=!6*[V8G4V805 0WM\7U0HR5
TRU0S87R85 T145VRSVU4XV[O[85HV8V458OG % )/&6-541\V8X41
@80XMRG8[VU\! #&" " !=9 >!"$2
,U8T4VM8C! 3845HB.! FOG4UC6$%#%6-@Y$! 4GRVM\1[418X\14VR
R[VRU4[R8[URLO8UR7 W0UVMR[\[VRG8X4XLO8UR7 UR[8[V45XR857OXR7
Z\4X8ZR5Q014U<
&!D&2
kI5R[! C85XMRU! PU8R7U8XM .! $*1#7 #""&2 mRHO14V805 0W
T4VM0HR5R[8[#4HR5RRSTUR[[805 85 V0Z4XX06Y145V
(R1%$&! ?! #?" >#D"2
/10V-(! F1R[[85HCP! Y4UI N6$%%=6\[VRG8X4XLO8UR7 UR[8[V45XR"
VMRR1O[8]R[8H541[6(OUeT85 Y145V@801! ##"!&D >!!$6
/RUZRU5Rf(! @U0O R^U+! CR1Z845X0P! $*1#7$%%$6fRVM07 W0UVMR
RSVU4XV805 0WVMR]014V81RX0GT0O57 [418X\18X4X87 WU0GV0Z4XX01R4W
G4VRU8416YM\V0XMRG-541! #&%#&" !? >?%2
N4U-m! Y4O1U4;fl! N4UfB! $*1#7$%##6m01R0W[418X\18X4X87 85
857OXV805 0WT145V7RWR5[R[\[VRG85 XM8XITR4%(8XRU4U8RV85OG.6&6
Y145V8H541@RM4]!D%##&"#9=" >#9"$2
AM45HNY! 3845H@! .0O .+! $*1#7$%##6)GT4XV0W[418X\18X4X87 05
VMR45V80S8745VR5Q\GR[\[VRG457 M\7U0HR5 TRU0S87RTU07OXV805 85
(OXOG8[[4V8]O[O57RUXM8185H[VUR[[6A +4VOUW0U[XM (! DD"
!#& >!$$2
AM40+! lO45 3! P0U0OM4UP,! $*1#7$%%"6,^0T0T14UGRVM\1
[418X\14VRR[VRU4[R[78[T14\X0GT4U4Z1RZ80XMRG8X41TU0TRUV8R[ZOV
78]RUHR5VRSTUR[[805 T4VRU5[6YM\V0XMRG8[VU\! 9%%#&" &$ >&"6
!
!"#$
E
 ¡¢
"
&##