概述竹类植物基因组学研究进展,包括RAPD, RFLP, AFLP, ISSR等分子标记的开发与应用,开花、细胞壁生物合成、光合作用、抗逆等相关的重要功能基因研究,基因组测序与比较基因组学等; 在分析和综述的基础上,提出竹类植物分子生物学研究基础薄弱、功能基因鉴定困难等不足,并明确遗传转化体系的建立、比较基因组学与功能基因组学研究、分子标记辅助育种是今后竹类植物基因组学研究的重点。
The progress of genomics research on bamboo is summarized in this review. I focus on development of molecular markers (RAPD, RFLP, AFLP and ISSR) and their application on bamboo studies, including bamboo genes related to flowering, cell wall synthesis, photosynthesis and stress tolerance, the genome sequencing of bamboo and comparative genomics with other species. The basic research on bamboo genomics is still missing and the bamboo gene functions are hard identified. Therefore, bamboo genetic transformation system should be established, and the research on comparative and functional genomics and the molecular marker-assisted breeding should be emphasized in the future.
全 文 :!!"
" !
#
#
$ % # $
$
#
%
& ( )
&()*+(, &(-.,) &(*(,)
./01!"!*/1#
2345!$ % # $
KW§¨Çî)PQö¶
HñF
"
Ké&RS &Ké()YDOCÇÈÉ
7
B
#%%#%$$
!
7
"
!7
8ºKW§¨Çî)PQö¶
!
ðñ
E,?P!EB-?!,B-?!(&&E
¼[Óå>µYÞ¹
!
`
(
¾Å
2ꬁ[
(
JŤ¹
(
Õu¼vw>¸Ê#ÇPQ
!
ÇîNYÖdÇî)¼
#
|[\º>
!
-ÝKW§¨[Óͨ)PQ~þ
(
Ê#Ç6¼n¿
!
)`±Èö¡:ú>
(
ÖdÇî)Y
Ê#Çî)PQ
(
[Óåñ§¬«ÑÉKW§¨Çî)PQ>
%
#$%
&7
KW§¨
#
Çî)
#
[Óå
#
Ê#Ç
#
N
#
ÖdÇî)
&()*
! &9#"1!>777
+,-./
!,777
+01*
!#%%# 69!"""$%#$#%# 6%#:; 6%"
#
& $%## 6%" 68%#
#
& $%## 6## 6#:%
&
&;(P
"$%%9%!%%#$%
9"*%"#00)&Q8<** W#&*8)30$#0#"34
2I34JRFLHI
"O&,F8A$"8.$",$(C8+A$$8*= 98.8* ?:5&*:&8*= >&:/*$%$,$(?.8.&3$"&1.",)=+5*51."8.5$*
X*.&"*8.5$*8%#&*.&"($"C8+A$$8*= 98.8*7C&5I5* #%%#%$$
;<0("3(&7+LF@S/JSFVV/XJF4/
SF03TFU T/X0/NFSI4J! YF0N30VW4TLFVIV! @L/T/VW4TLFVIV34U VTSFVVT/0FS34YF! TLFJF4/
¬ÄrrͨDO7ªvw(D>Q¬µ¶
!
*Qͺ>Ó¼Ð[f
$#
]^>()T
%
#;">
$,Çî)>8jñ
!
ªvwPQnó
°#)ô²ÎTÀ
!
¹Çî)Ð[fͺ()
ØSÌ
(
S{?@>¹Â)(
%
YÞ
!
Ç
îX
YÊ#ÇîPQ1WÁßÀ¬µ¶
!
Ç
î)>PQа#Áͺ()>¢Ø
!
|°
53?@Äͺ()$ñ>µ¶
"
GQ
!
$%%9$%
Ù§¨ñ=U
")"8A5=$E151./8%58*8$(
Qé
"J",G8 18.568$
â[ÇîNjñ
!
{MÛ
¬§¨vÓ¶ßÇîN
!
ªRP&(
"?/3YF3F$
§ ¨ E Ê
"K$"=&;+ 6;%8"&$(
t Q
" ?$"/;+ A5:$%$"$(
¿ K Ø â u
"C"8:/,E$=5;+
=51.8:/,$*$(
eÊ
">"5.5:;+ 8&1.56;+$(
x}
"T&8
+8,1$
¼vÓâ[
!
hK (
"D3<[HV/IUF3F$
§¨>
ÇîNÍÛW
%
KW§¨ÛuP&(K (
!
«w¸>¦
&FG
!
|»¦&NE!>û(X
!
KÓ
jªÍÎÀ¬
(
½ÆÍ;4>ç
!
|Ü~4Î.^
RS
(
öÍáLÃ;<êµB߸¤
¹
!
ªµx¹|»çès|}ßtcwX
%
Ç
Þ
!
¤f;PQKW§¨Çî)NÀÁ
Æ
%
¹
!
KW§¨Çî)PQk¸þR|[Ó
å>µYÞ¹
!
Ê#Ç>[²Yçè[!
ut¢N>
)&+(
Î
YP*,(
Çî>[\Y
ÖdÇî)¼ê
%
#7
[Óå>µYÞ¹
[Óå«jÈö¨>ÈöåP/4JOF4tH34! OH \I3/U34! -I4 CH34! &.8%5$%##5)Z30H3TI/4 /X#:
!
#âýͨ¢ú¨¬Çî
R¬lm>çm;
P*,
^ß
"
?¼
!$%%#$%
[ÓåDOÐô²ÎÞ¹u২7¼Í¨>È
öß
(
Çí
(
ÈöÛK;[(
äîw
«
(
Çî]ÛN
(
ÇMâ¼ê>PQ
%
=
uEÛÚjô;q¯>KW§¨ñ3
!
[ÓåD
Ok¸Þ¹uÈöÛK;
(
ãö:[(
¬<
& ( )
!"
"
7
¼ê>PQ
%
BABC
YZ[2M(¾
KW§¨uvÛ½
!
]¯{
9%
Û¢Û
# $%%
Û¬
!
|Íáã
(
¨¬
(
ÈöÛK;êi·{%
&;
!
8òÞ¹
E,?P!EB-?!,B-?!(&&E
¼[Óå
DOñàß¿OKW§¨¢ú¨úlm>ç
m;
%
E,?P
DOôSU¹uKW§¨ÈöÛK;P
Q
%
=x+K
"7;1/8*58 *5.8M8,8+&*151$
>PQ]
^oO
!
«3,²{clm7éßú[:
!
ª
ÛK;«°uÍÃlmy-[>e3,[:
"AVI3/
&.8%D! #;;!# #;;;$#
=vK
"!/,%$1.8:/,1&=;%51$
¬
X>
9
¢YX¨·¦X7ªÛ>
$
¢ªî¬ö
Èöw>PQ
"
^¨¼
!$%%$$!
j7=ñ
>
¢ú>
8%
v·UK
"C8+A;18 E&"68"58A5%51$
¢ú
>ÈöYmPQ
"
@6V¼
!$%%8$
öÊ_2Çß
E,?P
[Óå=KW§¨¬sÈöÛK;PQ>
{ ;
#
= b K
"
: æ K
$ " !/,%$1.8:/,1
65$%81:&*1$"
¼
!$%%#$(
ªK
"C8+A;18 $%=/8+5$
"
!)û¼
!$%%:$
>PQ_`
!E,?P
[Óå=K
W§¨¬jX¼ö[W{Ê>½;
%
K
!E,?P
[Óå1½j¹uÛw>PQ
"
G
JC¼
!$%%#$%
,B-?
[ÓåDO]Åß
EB-?
E,?P
D
O>ç
!
Ð[f§¨äîw
(
¬<«7ÈöÛ
K ; P Q > { ß É Ê
%
= 0 K Û
"!/,%$1.8:/,1$ $8
¢¬
(
WKÛ
"!%&5$A%81.;1$ $
¢
¬
(
ÏKÛ
"?5*$A8+A;18$#
¢¬>
,B-?
ÛK;[
\>Ú1-O
!
|ã)PQR¿âÚ>
,B-?
|¨îÅ«:Wæ±òWw>
"
ºXL¼
!
$%%"$%
|ÛNYKÛ
"X*=$:8%8+;1$#>
¢¬
:
¢
+áv¸>ÎGK¬>Çî
P*,
Ûá;µr
!
§{>YKÛ§¨ò[ÊW
!
ªR
:
¬ÎGKW§
¨ò[ÊW
#
,B-?
[\-.>[Ó21¶
ßZÑ[Wã
!
Ä6YK
"X*=$:8%8+;1+5$5$
y#«YK
" X*=$:8%8+;1%8.5($%5;1$
R
"
[!¨
¼
!$%%;$%
=\K
"#/5+$*$A8+A;18 .;+5=515*$=8$
$
¢¬>ÈöÛK;>[\]^_`
!
\K¬
s{dt>ÈöÛK;
!
{xu\K¬
Ë
"
]²Ç¼
!$%#%$%
¬ÄKW§¨PQM¤>öÊ_¶
!
K
W§¨>
(+&"A/U^I4V/4 &.8%5! $%%%$!,K\"
´
^¼
! $%%9 $!(&&E"
we_¼
! $%%"# -I4 &.8%5!
$%#%[$!&&E"+34J&.8%5!$%##$!)&+_&&E"
G^¼
!
$%##$
¼[ÓåvÓôµ
!
¹uÈöÛK;[
%
fß-t[Óå>{;
!
éÝrß
$
¬
8
¬[ÓåÅ@¹>
!
)Wß7(>
^
%
=fgn3×>l+`K
"
:E`
$
"\&*="$:8%8+;1(8"5*$1;1$"
«a¼
!$%%"$(
ÜéæK
"C8+A;18 "55=8$"
«a¼
!$%%;$
>PQ]^_`
!
Ò¹
E,?P
(&&E$
¬[ÓåÅ[!
½j{
3¬sÈöÛK;
#
=u¬s¬<Ò¹Å
«[\>
½j{-t¬<«
!
=
#%
S
vK¬<Ò¹
8
¬å>Ú1Å+«[\]^o
O
&
ª^çf
",B-?j(&&E j&E,?$ m
",B-?j&E,?$ m"&&Ej&E,?$ m",B-?j(&&E$
"
o}J¼
!$%#%$%
BAEC
UV%K(¾
°uKW§¨¬WqÛ
(
+á;%&ÛY
(
Û
$Íʸ;`¼>Çy[f㵧)PQ6
%
KÓöã>+á)[Wk¸0Ûu`jÎ>5
P9ß
!
y+á)[W
&b:Ë|Ñ;
%
[
ÓåDO«u
?E
DOö
P*,
5>ËÌ
;°®
!
°®.¨>Ûá;#â¹u+ón¨¬
É>ãö:w
%
ÇÞ
!
x¹Èö¨<
P*,
ñ
öKW§¨>ãö:PQ{Ä`o>÷ø
%
Ò¹
E,?P
[Óå#â{3[\KW§¨
ªî¬>äîw
(
+ÈöJ²
"
G:¼
!$%%!$!
=WK
"!%&5$A%81.;18+8";1$
7ªîK
"
#9
¢K
¬
$
> ä î w > P Q ] ^ _ `
!
E ` K
"!%&5$A%81.;1"8+5*&;1$
YeæWK
"J%5$1.8:/,;+
(;I5*&*151$
>ÈöJ²SE
!
f
%1>8: >!
U´WK
"!%&5$A%81.;1I;@58*&*151$
YÜéWK
"
:bcK
$
" ?5*$A8+A;18 5*.&"+&=58 $
> È ö J ² S e
!
f
%1$!; %%
°uªúÇî>;¦;
!
ª
P*,
dæ3Þ
¹uKW§¨ãö:PQ
!
o
*=/3"03S^&.8%5!
#;;:$!"E1a"*3U/T&.8%5! #;;:$
j7
"E%bc
>sÓ
"rF0YL4FS&.8%5! #;;9$
¼
%
)Î
!
Þ¹?Ç
P*,
¶ãö:[\PQé{?ìú
P*,
s¡
Ó×
"KH/&.8%5! $%%$# &H4 &.8%5! $%%:$(
s
$úÇ
"E1%a"h4J&.8%5! $%%>$
<ú
E1A,_."*A!
"E%deB."*-
"E1bc
sÓ5
"C34JAM&.8%5!
$%%"# C34J2D&.8%5! $%#%$
¼
%
!
?Ço
`Ç
3>"C/^/&.8%5! $%%"$(P)\?b(>Cb
[d
Ç
PJ#b"
mx·
!$%%9$
¼1ô¹uPQKW§
¨>ãö:
!
é{¢YÊ$Î
Öd¨¬É>ãö:
"
`%¿¼
!$%%>$!
[«
Ãn¼c|nK¬çès*QßKW§¨>ö
:w7ª|P&(ãö:R>3í
%
õÇ
%>#
7!
#
# ¿X
&
KW§¨Çî)PQö¶
"@VFHU/JF4F$
«úƨ¬ö:>ÉÊ
"KFSVTFI4
&.8%5! $%%>$!
y¡ýÓYõÇ>Ýre{v¸É
A
!
ÇÞª$ñ¡ýÓ|KW§¨â:R>Èöç
;1ÀÁßPQ¥>wX
%
=¡ýÓQÅ
"?/4J_
0I^FF0F
7¡ýÓýQÅ
"+W#_Y/@I3SFTS/F0F
ç;>[\oO
!
ò·{d(>Ûá;
!
h¡ýÓ
QÅ>ã)[\]^YKW§¨>ãö:)n
Ë|µT
%
ÇÞ
!
F
f
KW§¨>ãö
:^¸ÃÇî>Qòñö
%
BA@C
wN~
KW§¨{u&K
(
4&K
!
{vÍK
(
±ÍK
(
1ÍK
!
ª¬<>«k¸uK¬>+áçè
!
y
2K¬>+áçènâùªÆÖd6
!
©
ª«¬&¬
!
ªFÇ;½6Ã+á¼cYä&v
×[
%
[Óå
(¥ß+á)«>n¿
!
#
âÃÈö¨<
P*,
QòñÃM
(
æ±3ÛNnK
W§¨¬<>lm
% BSI3S
¼
"#;;#$
íUÞ¹
EB-?
=
$>
¬KÓ>
!$
¢nÇXöß
PQ
!
~1»¬äîwÝ
$:
¬f±ÍK
%
&&E
|Ç>ní5i{[ä
!
(·{n
>Ê#
!:1
gò×>
&&E
·{¡TAÊ#
!
s
Ós>
&&E
TöÇ>_"
!
ÎoÓR>
&&E
L
?@Ç>¡
(
"-I&.8%5!
$%%!$% &&E
½#Yª_X{Äbw
!
¤fIË
å5f{
%
ÇÞ¤f
P*,
[ÓåR>lX
!
&&E
вÎÞ¹uFõ&¬>
" UFh0IZFIS3&.
8%5! $%%$$%
¬ÄKW§¨Çî5
)&+
5
>nó®Ï
!
f
&&E
å>IË-.ßE>Ú1
ñG
!
¢Yͨmn)[\½½£Ý{
&&E
[Ó
å
%
x¹
*D(
Ú1¬RvKÇî
K&&
"JF4/
&&E
å
=&2K¬>«]^_`
!
|
8
¢
&&E
í
"?D\%#!!?D\%#"
?D\%$:$
°®.¨>Dt
Äõ[«fªh\&Äõ>îÅ
!
N]^_`&
¬ä&É>v=Þ>5·{G;
!
2`ß
&2K¬>FÇ;
"
¿p¼
!$%%;$%
x¹
)&+
5µÝñ>
)&+_&&E
[Óå
!
Ǫñ_"
5_rfo;å
!
|ä&YÓrÉ>½¡
;dt
"D/HY^ &.8%5! $%%9$!
§j=u&¬>
5f{
%
x¹ÃøªK
"C8+A;18 $=81/5+8&$
ªK
YP*,
u¬Rµ>
)&+_&&EV!
=`K
"C8+A;18 .&@.5%51$
Y_K
"\&*="$:8%8+;1%8.5(%$";1$(
vÅÁ%K
"\&*="$:8%8+;1/8+5%.$*5$
Y_K
(
·U
KYªK>¬&¬FÇ;>«iWß7(>
^
!
=uó$o;å
!
v=Þ>&2K¬°®
.¨>Dt]^[«_rfªh\&Äõ>îÅ
!
N]^öÊ_È2ß&¬ä&É>5·{
G;
!
2Çß
)&+_&&E
[Óå>{;
"
i
*¼
!$%%;# P/4J&.8%5! $%##$%
ßx¹KW§¨bµÝñ>[Óåñ
ö¬<«Î
!
ªR§¨>¢¹;[Óå
!
©ª
«KW§¨ªî¬Qé
(
x}
(
eÊ
(
Eʼ>[Ó
å1½¤f½(>
%
&§Ú¼
"$%#%$
Ï
EjÖ E)
"kD$
>
(&&E
|¨5
!
ÃR
IËÝ
"
¢Ûá;dt>|¨¹uõvK
"!/,%$1.8:/,1M08*15&*151$
vK>
ý2¯¸&
¬
A#
A$
>ÈöJ²[!
]^_`&¬
A#!A$
|ÇîQòY»äÉË|`olm
!
h&>
ÈömnÐí#Ór
!
2Çß
$
¢¯¸&¬>FÇ
;
# $
¢&¬É
(
&¬Yh\&ÉiË|uv
>ÈöYm
!
)(
$
¢&¬|äîwïuõ
vK
%
$7
Ê#ÇPQ
EABC
MO#AÈÉ
nK¬>`
(
?#Âm
!
ÇÞ`ͨ
)ͯͨ)ʵ«KW§¨PQR>6
!
U
ENc3ÑÒßKW§¨&2§¬M¤>¶
%
[
Óͨ)DO>Q¬µ¶fÃ[ÓQòñPQKW
§¨>`µ§-.ß7(>
%
\,P&_[/Q
Ç«`µ§TARÊW¸
>¡ÇÓ
!
Y9ßµ§
(
`>AÒ
%
[
¼
"$%%:$
Ò¹
E,)
DOÃvÍK_KMâÁß
Ê¢
\,P&
ñÇ
\%P)\?bf!
)¢Y|ñ=UR
mí_">_X[!
p_2`æǽ½#Y
_K`>TA
%
tøÕ¼
" $%#%[$
Ò¹
E+_?E
E,)
DO
!
ÃvKR[²ß
\,P&_[/Q
Ç
!&P)\?b!
Þ¹CR`7Ä>Ç¡#ñ
=URmí_"
!
¡Çñ=U·{`-æ
(
çý
"
(
µ§1ܼ_X
!
_`
!&P)\?b
½½#
YvK°5PÍΡͯÍÎ>µ§TA
%
!
é{Ê$YKW§¨`vw>Ç
!
o
C$P)\?b"
tøÕ¼
!$%%9[$(93>b"AIV3$%%9$(>Cb"
G6X
!$%%9$(#J-?>)-?"
ö^C¼
!
$%#%$
¼vÓôMâ
!
)ößvÞ>Ê#ÃNY
_"[%
EAEC
ïC5Ö×í:O#AÈÉ
¾Å2«§¨¾Å>¯Ñô
!
«[§¨>Ç
#>#
& ( )
!"
"
7
ú
!
>Ê#ÌY§¨>ÊͺÌàõ{w
!
ªÍ¨Å[>PQʵnÀwX
%
KW§¨¾Å2
µT?@KÎ>Î;
!
8òÐ4èÃ[ÓQò
(
Ð
WQò=ªÍ¨Å[>vwÇ7ªïû>ÐWö
PQ
%
¾Å2¸î[4<È
(
0\ȼÛì73ì
>ͨÅ[TAʵ«¾Å2PQ>T
%
ß¾Í
Ë~Íw
" @LF4W03034I4F3<4I3_0W3VF! ?,-$
«ß
¾_Wrµ>wOw
!
«¾Å2¸î[4<ÈÍ
¨Å[>!
#
¢w
!
PQ_`êKN´R[²>
?,-
·{
?,-g+,-
>»Ì;
!
|ªKs{
!
¢
GÇ
"AVIFL &.8%5! $%#%3# $%##$#
úÎ_"PQ
2Ç
!
ªR
C$!)Fe!C$!)Fa
_">ÐW·{
?,-g
+,-
»Ì;
"AVIFL &.8%5! $%#%[$!
y
C$!)Fb!
C$!)Fd
_">ÐWê{
?,-
Ì;
!
óâQÄ|ª
Kús>n
?,-
½#ïû
?,-
>n
!
@
Â>Ê#
%
klË
6h6
^¡w
"h\+$
«
4<ÈͨÅ[>)ÊwOw
!
¤¥§|PQî=
¡ýãªK
#JP>
Ǭu
"-5:$.58*8 .8A8:;+$
>
4<ÈN]^oO
!
¡Çg4<È
õ{nNc>Y:
!
ªRXPSÛ>4<È
Ö=KXPß
$"19i!
3`MÒ§¨sG
#JP>
>
_"{3P߬uR4<È
%
÷ìÅ[w
Ç«SæÃKÓR[²Ýñ>YìWrµ>
Ç
!
|ªKRb!{
!
¢GÇ
!
ª_"ucYª
ÍάvÊé
!
h|n²í>_"ucDË|
ÄÊ>lm
!
úÎ_">ÐWÌ;ÛNoO
!
î
ÐW
SD/&H! SD/&H&$! SD/&H&8
SD/&H&!
=֓
ü¨ > Ì ; [ « f
%1##:! $18;9! %1"%!
%18%> k
!
óâQÄ
!
¬Çïû>ÐW|K
Óús{ÄÂn>Ê#
!
=KÓ¾Å2>µ§
j7¬ÍµBĸ¤¹
"LIH &.8%5! $%%>$%
2
!
Ð4[²>YKW§¨¾Å2ͨÅ[>
Çé{
#$#JP>"
ºxò¼
!$%%"$!3gK"
G)u¼
!
$%%;$!!&!)Fb"
tøÕ¼
!$%%;$!!/!Bb"
G)u
¼
!$%#%$!#aK"
çx¼
!$%#%$!C$?4>e"
tøÕ
¼
!$%#%3$
!E#&1)b"
ë··¼
!$%#%$
¼
%
EA@C
pí¶£O#AÈÉ
KW§¨¬Í>çâQĪ½#·{d4>
:#
!
yª:#«¢YJŤ¹>[ÓT
AñÇr>
%
ª$ñ
!
Ð{Ã[ÓQòñ*)KÓ
JŤ¹vwsÍ>"#
!
o=ªKR
%/:Ab"
tø
Õ¼
!$%%93$(%/:8a
Ç
"
ºxò¼
!$%#%$
>[Y_"PQ
!
=vK
(
í´K
"!/,%$1.8:/,1";.5%8$
¼K
"!/,%$1.8:/,1(5+A"5%5;%8$
R
%/:8
Ç>Mâ
Y["
ÏuC¼
!$%%"3# $%%"[$%
¤¥§|PQ
î
"
î^
!$%%"$
ÃvKR[²Á
#$
¢@J×È
]ÅÐWÇ
!
[«Ûu
?&
%>
%/:Ab!%/:Ae!%/:Ad!
%/:Aa!%/:Ag
%/:Ac
WÇ
!
ªR
%/:Ab!%/:Ae
%/:Ad
ïû>ÐWÛuE@J6s
!%/:Aa!%/:Ag
%/:Ac
ïû>ÐWÛu¼@J6s
%
ÐW]Ã
N_`
!%/:Ab!%/:Ae!%/:Ad
WÇïû>ÐWið
7m
!
([QÐW>¿]ið
!
¢ àD]
(8
¢WnÈ]Åíj7vÞ>ªÈ
3![
]Åí
%
îçm;_"ÛN_`
!>
WÇ|
^
(
ûq{Ri{_"
!
(|^RSt
!
y
|~R$ÛNÁ_"
%
|4JK8
"# :%%
#
0
<
6$
V
6#
$
ÄÅX
!>
WÇ>_"irXT>N
ø
%
úÎ_"vK
%/:Ab!%/:Ae!%/:Ad
ïû[QÐW
)YW.ú×Èî+[%Åú>çèoO
!-LY[#
-LY[$
i#+[3úrú
!
y
-LY[8
n#
#
n
î3úrú>ÐW5çè
(
×È[
(
J
çèT
6
J;[\]^_`
!-LY[8
>Â
êçè`onu
-LY[#
-LY[$!-LY[8
·{d
>ö#dt>T;
!
|3úR
-LY[$
S
%
óâQÄ|½Y>LÃRKÓn>
-LY[
Ð
W½#|J#@:YÄ#>±RµBÄn
>Ê#
%
EAFC
67O#AÈÉ
¬Ä»LÃ>Y:
!
ÃÚ
(
tR
(
R
(
È}
(
ۼͨVWÐ4[f?@§¨
wÍε
§>¸ÑÒÇÓ
!
¶KW§¨ÕuvwÇP
Q=u¢YÇMN¦§Õu6»¬·{¸Ñ
%
*ÜÐW
"]I4YXI4JFS@S/TFI4$
Ç«Ê¢
oE>¡ÇÓ
!
=ÃÚ
(
tÈR¼Í¨
VW·{Ê>Tö¤¹
%
îø¼
"$%#%$
Ãv
KR[²ß
!&T3!
Ç
!
ªïûÐWYªR*Ü]
ÐW{dt>G;
!
5
V·{lX>
,*#
WX>*Ü]
Q$_!Q;_#$YQ$Q!YQ$AQ:AQ!
|
*
V·{lX>
,$%
WX*ÜÐW]
!
LNÞ
!&T3!
f§¨Õu
J1X)!b
W¸Ç
!
|Ê#Yv
KÕu;vw
%
¤¥§|PQîêKR[²Á
#
¢
D_D/Q
X*ÜÐWÇ
C$CT3"KF4D34^
pÍ
&
)k>%>%$:$!
ªïû>ÐW·{
$
¢
D_D/Q
]Ø
!
Ûu
D_D/Q
X*ÜÐW
#
îçm;_"oO
C$CT3
fî[X_"
!
|^
(
û
(
q{~Ri
{_"
!
ªR|^>_"ucdt
#
|ñ=UR
mí_"
!
¡
C$CT3
ǧgHÚ;`o-t
!
â
QÄ
C$CT3
Y§¨>ÕÚ#{w
%
)Î
!
Õuv
wÇog1q
68 6
"Ëé
Gâ¼
!
$>#
7!
#
# ¿X
&
KW§¨Çî)PQö¶
$%#%$(
(
6#!8 6
|òìwÇ
"
bk¼
!$%#%$(
$
óô
*3
j
gA
j
u¯ÀÐWÇ
"
bR9¼
!$%##$
¼1vÓÃvKR[²
!
=u$ñ>KW§¨[Ó
§¬-.ß{>
%
ßjÌWÊ#ÇPQÎ
!
é{¢YûE
MÒ&2«vKö
¬²Î>_"5år
"RL/H &.8%5! $%##3$(
ÏK
"!/,%$1.8:/,1*5"8$
Ûa
|:wÇ
"
¤s¼
!$%#%$(
ÀÂÈ>mÍö7v
wÇ
C6#X_9"RL/H &.8%5! $%##[$
¼Ê$Ê#Ç
MâY[\>vw"#
!
hiRS°#>Ê#PQ
%
87
NPQ
@ABC
8
B
9$öÏÐE£
|Çî)PQR
!P*,
>5[\«öÊ_
PQ ->Ç>
%
x¹MNöK
W§¨>PQÍ$%"#
!
¬ÄàN[f
P*,
5[\>kK
!
KW§¨
P*,
5PQ"#1¬
É®Ï
!
o¹§º>ÂWÇõ«¢YàND
O:W}5ñöͨmn)[\YÊ#p_
È2>
%
b
$%%"
$
#
%
*D(
RK (>Çî
5
$ :">
Ä
!
t^¼
"$%%"$
=ªvwmnöß
ã¡[!
)=ª|
K&&
5
(
[Óå
(
ÇM
â
(
ãö:¼ê>Þ¹öß/º
%
¬ÄNDO>nóµ¶
!
!
#
rN=>Û
¢#DO½(3-tßN
%
x¹
,D(
>
89%%
N=[«ÃªK>`KN
YP*,
u¬
R:Wß
! !9%
8 "9"
Ä
)&+"-I4 &.8%5! $%#%3$!
Þ¹
,D(
>
898%
N=[«:Wß_KªK
ªúÇîÚ1
#8; 8:% [@
#8; 8>: [@"OH &.8%5!
$%%;$!
)=:WÚ1ößvÞ>[%
¤¥§
|PQîÞ¹
,D(
>
898%G-
N==vKnÎ
HWX>Î
YP*,
u¬öN
!
:Wß
89 9;9
Ä
:b
V5
!
ðß
#% >%"
¢Î
YP*,
5
!
òi
Îcf
# %;$ [@!
ªR
"$i
>5{
#%%
¢Í
Ëj>
hEB!
)í¸|TªÇî>Qò=K
W§¨]Ççöß[\PQ
"?F4J&.8%5!
$%#%$#
)Î
!
é¢Y=_K^iÊ:u¬>N
!
:Wß
; :9!
¢t<>
)&+!
4î*WÁ
: 8#9
¢
H4IJF4FV"K3/&.8%5! $%## $%
x¹
+(KE?034T
+S34VYSI@T,VVF<[0IFV
Ú1¬
!
ÄK (>vKYé
(
"hSW]/IUF3F$
R>Qé
!
æQP (
"?//IUF3F$
R>eÊ
(
EÊ¿KØâu
!
u (
"?34IY/IUF3F$
R>x}
(
tQ
(
g÷
" ?8::/8";+$(5:5*8";+$
~
v
"!8*5:;+65"8.;+$
¼ö5Ö=
!
µr|P
&(§¨RvK5YQév¸;St
!
"
!;1$i#
|
;
¢P&(¨¬
#
¢>¨¬ñ=U
R¡Áß
!8
ÄG5
!
ÿ¡
8" !#" [@!
x¹SE
½#;
D3WFVI4XFSF4YF
WÁ>ö:_
`K (æQP (äîw5ª
!
y°
*FIJL[/S2/I4I4J
WÁ>ö:ÿoOK (
é (äîw5ª
"?F4J&.8%5! $%#%$%
@AEC
8
E
9$öÏÐE£
¹
!
!
$
rt¢NDOвÎÞ¹uÍ
¨ÇîN
!
|KW§¨>Þ¹1WÁßd
µ¶
%
x¹Ñ>>
!:! K& B-G
N=
,D(
>>
898%Q0
N=v]Å>ß=vK
##
¢
D,
MâößPQ
!
:Wß
$
¢âù>
D,
5
"KM$:$"">
KM$:$""9$!
§WÇî5YQ
é
(
tQÇîÉ_rfdt>s;
!
óâQÄ
Qé
(
tQ>Ð75=u~bKW§¨Çî·
{¸Ñ
"KHI&.8%5! $%#%$%
x¹
(0H
>
&/0FQ3
N=Nß
#
¬TõKÓ
:
¬RõK
Ó>ªúÇî5
!
)YÐ7>
$
¬KÓ
ªúÇî5
"OH &.8%5! $%%;$
Ö=µr
!
K (
§¨>ªúÇîw;¦
!
·{7(>s;
d>[Óö:¬
!
x¹r{>P&(§¨
ªúÇ5>P&(㵧oO
!
K
(æQP (fwxw
!
|K ([¶R
Tõ[¶Rõ[¶[«f
$
¢½(>3
!
2`
Ò¹ªú㵧Çî)ß=RõKÓö
㵧«ÖdÉ{>
"RL34J&.8%5!
$%##$%
y
!
=uKW§¨ÇîN>"#ÍÛ
W
!
fϬR&4Èö§¬öN
!
@R&|
¼MØ{§¡E
!$%%9
$¤¥Uy7ßR&
!
#
¢¨¬>ÇîN¡È
)))
vKÇî
NPQ
!
)Ås÷ø3íþÇÔ
%
æ
x¹
&/0FQ3
NY
D,
½VNv]Å>Ú¯ñ
ö
!
¹Ð4î*:WßTª
$ K[
>Çî5
ÿ
!
v2uvKÇî5ÿ>
;%i
j
!
·úÚ1ëÄä
%
t¢NDOϬ߿KW§¨ÈöÓ¼
>öN
!
¢YNê½jß~3¢Ç>]
!
ö
Ê#ÃN
(
¶Ê#PQ
!
:½jß~ù¢Çî]
Yî[
!
¶ÖdÇî)PQ
!
¢Yͨmn)
¶KW§¨ãµ§)ö:PQ
!
*)KW§
¨|P&(¨¥ù¢§¨¯>ãâ:w
(
ö:
¬¼
%
!
½jµÝE{>[Óå
!
¹
uKW§¨>[Óñ§¬
%
8>#
& ( )
!"
"
7
!7
PQYµ¶Nø
KW§¨FG«Ò3¦&Íáã>¸î[
²[
!
·{7(>Íá
(
4Ñ
L
!
|zÐ
[f&µ¶R>Ê¢6>4Ñ®Î
!
ª;
PQÀÁÆ
%
´MKW§¨Çî)PQö
¶
!
|[Óå>µYÞ¹
!
Ê#Ç>[²
(
N
Y[!
ÖdÇî)¼êiWß½g>ö
¶
!
z=KW§¨Çî)>PQ1|ß]¯
¹5
%
h°uKW§¨b>çä;
!
7éª]
Çî)
(
Ê#Çî)
(
ÖdÇî)¼;P
QÍË|MÛn¿
!
©ª«=KW§¨[Ó§¬>
vwPQ
!
`o´ÉuªRÙ§¨YC¤¨
!
óâ
QÄKW§¨>Çî)PQ{Ä^E>µ¶
%
FABC
A
B
9$öÏÐEÃH£
¬Ä¬rr(Dß>À¬µ¶
!
_"ËN
(
Î7úN
"E/TL[FSJ&.8%5! $%##$
DOnóâ
!
:ÝrßÊ$6>NDO
!
o1J_"Ë
N
"&I
ß3]X[\DO
"LF4 &.8%5! $%##$
¼
!
ófKW
§¨>Çî)PQ-.ß4{>¶·
!
ìÄ
t¢NDO|KW§¨>Þ¹
!
KW§¨>
Çî)lÄWÁê>µ¶
%
FAEC
AYZL|:E;H=
|j4lÈö),/fÜ7>ÇR
!
Ýrß
MÛô
¹öãÈö,/ñ~
>ͺrH
!
ì
PQ>nó°#
!
6>Èö),/
)))P*,
zß,
/Ðô-Ý
"OH &.8%5! $%%>$!
æ,/Ïfà ͨ
>Çîõ«°
!%
¬QÅ[
!
ªRðñ
>
¬]
QÅ
"K
(
Ç
(
¡ýÓ
(
ý¡ýÓ
(
Î3%
5]%
$(8
¬ö:vw>QÅ
"
î
(
}ã¢}ó#YRK
$
8#
WÊ#
Ç
!
æ,/>
±;nóWÁ2Ç
"L34J&.8%5!
$%##$%
ófKW§¨>PQYµõñß6>³
¬
!
{ÂuµTAKW§¨¸;>Ê#Ç
!
)¹uKW§¨>òÅ[Ó§¬
%
FA@C
ü"ÈÉh<
¸ÇrKW§¨Çî)>굶Í^|j
XÌ¢ê°#¶PQ
& #$
Èö¡:ú>
Y÷:
# $$
Çî>°cNYÖdÇî)P
Q
# 8$
E^ßÇî¡:YÊ#Çî)PQ
# !$
Û¬[Óå>µYñ§¬Þ¹PQ
%
ìÄ
KW§¨Çî)PQM¤>ê¶
!
ªP
QÄL{Q;>µ¶
!
¦§·{k7Ö.>
KW6»¬ÄÅÆYfrÇ
%
X Y + ,
ö^C
!
G{
!
Gp`
5$%#%5
vÅ%K
h*&+,*&
GÇ>M
âY5[5
&()PQ
!$8 "#$& # 6:5
´^
!
[
!
Âa
!
¼
5$%%95,Kå|KÓR>¢¹;
5
?B&E))"
!8:"#$& ! 6>5
ë··
!
|
7
!
À
5$%#%5
bK0\ÈÅ[wÇ
YP*,
Mâ
Y_"[5
¿ACE))"
!8$ "8$ & :8: 6:!%5
`%¿
!
o¤
!
¹
5$%%>5
KW§¨YQ鼪P&(¤¨
>ãö:w7Ç5î[>Öd
5
&()PQ
!0;
"$$& #>: 6#>;5
7
!
}-
!
æ
!
¼
5$%%#5
bKn·¦WX
E,?P
[Óå
>PQ
5
é¿&)F)"
!#""#$& # 6:5
?
!
f8
!
Ï^7
5$%%#5
¤¨
P*,
åñ§¬
5
B
&
(
)Ýv
5
tøÕ
!
ºxò
!
$¨
!
¼
5$%%935
KÓ@JªÈ
3g[
]ÅÐW
ÇÎ>Mâ5[5
&()
!!8"8$& 8! 68"5
tøÕ
!
î^
!
ºxò
!
¼
5$%%9[5
梻K
\,P&_[/Q
Ç>M
âY5[5
[Ó§¨§¬
!:">$& ">> 6"9%5
tøÕ
!
$¨
!
ºxò
!
¼
5$%%;5
vKß¾ÍË~ÍwÇ
YP*,
Î>Mâ7î_"[5
&()PQ
!$$ "8$& !!;
6!:85
tøÕ
!
G)u
!
$¨
!
¼
5$%#%35
ªK
C$?4>e
Ç>Mâ
(
[Y ¾ÅíPQ
5
&()
!!>"$$& !: 6!;5
tøÕ
!
4
7
!
$¨
5$%#%[5
vK
\,P&_[/Q
Ç>Mâ7Ê#
p_PQ
5
&()
!!>"#%$& 89 6!#5
o}J
!
àe
!
[!¨
!
¼
5$%#%5
vK¬<[Ó«
&E,?(,B-?(
(&&E
Å[5
¿ACE))"
!8$":$& ;"$ 6;">5
t^
!
b
7
]
!
\
5$%%"5
KÓÇî5PQ7Þ¹
5
KÓ
PQ®l
! $9"$$& # 695
«
7
a
!
Í-
!
/ª;
!
¼
5$%%"5
fgJn3×l+`K
E,?P
Y
(&&E
ÈöÛK;PQ
5
£&)F)"
!$""8$& $9> 6$"%5
«
7
a
!
Í-
!
/ª;
!
¼
5$%%;5
æ{¸
5
fgn3×ÜéæK
E,?P
(&&E
[5
KÓPQLl
!$""#$& > 6##5
çx
!
]F
!
t
7
5$%#%5
vK4<ÈÅ[vwÇ
#aK
>M
â7î_"[5
&()PQ
!$8"8$& 8#; 68$:5
ºXL
!
oeû
!
$¨
!
¼
5$%%"5,B-?
|¨îÅÚ=æ±PQ
KÓãw>?@
5
§¨)¢"
!$:"!$& !!; 6!:!5
ºxò
!
tøÕ
!
$¨
!
¼
5$%%"5
ªKklÎñw
,_h_
^¡
wÇ
"#$#JP>$
>MâY[5
[Ó§¨§¬
!> "8$& :"9
6:;$5
ºxò
!
$¨
!
tøÕ
!
¼
5$%#%5
ªKJã$@JªÈ
3g[
]
ÅÐWÇ>
YP*,
ÎMâ7[5
ECE))"
!89
"!$& >!8 6>!"5
î^
5$%%"5
vKJã%@J×È]ÅÐWÇ>[²7ª_
"PQ
5
R&()PQF®¯)í/u
5
îø
!
bR9
!
G
7
^
5$%#%5
vKÕu*ÜÐWÇ
YP*,
Mâ
Y5[5
ͨDO¢"
!"#$& "9 6;$5
&§Ú
!
7
!
~Í
!
¼
5$%#%5
õvKvK
ý2&¬>
(&&E
5
[Ó§¨§¬
!"":$& ;:" 6;>$5
¿p
!
}§{ñ
!
7
!
¼
5$%%;58
¬KW§¨&¬>[Ó
5
&()
!!:"8$& $; 68!5
!>#
7!
#
# ¿X
&
KW§¨Çî)PQö¶
[!¨
!
$¨
!
qÉ-
!
¼
5$%%;5
²[YKÛ§¨>1J
,B-?
[
5
&()
!!:"##$& 8$ 68:5
]²Ç
!
´À
!
b
7
!
¼
5$%#%5
¯¹
,B-?
DO[\\K¬È
öÛK;
5
&()PQ
!$8">$& ":% 6"::5
we_
!
&6
!
&§Ú
!
¼
5$%%"5
vK¬GÈöÛK;>
,B-?
(&&E
[5
é¿&(D
!$""$$& $; 6885
mx·
5$%%95
uÊ#Ç
P)\?b(PJ#b
>Cb
>KW§¨[Ó
ã)PQ
5
é¿E)®¯)í/u
5
^¨
!
GJC
!
&6
!
¼
5$%%$5
vK¬X¼>
E,?P
PQ
5
=
&E))"
&
()v
!$>"8$& >: 6>"5
ÏuC
!
$¨
!
t
7
5$%%"35
0KÛ
8
¬¸±ÍKJã$
Ç
"F/:8b$
>Mâ
(
5[\ÐW]ÃN
5
B&E))
"
!8% "!$& #%; 6##:5
ÏuC
!
$¨
!
t
7
5$%%"[5
vK
"!/,%$1.8:/,1&=;%51$
Jã$
Ç
F/:8!&he
Î>MâY5
5
ECE))"
!8:"$$&
#:8 6#:"5
[
7
!
/§i
!
N³
!
¼
5$%%:5
Ê¢KW§¨
\,P&
ñÇ>M
â7ª|ñ=UR>_"
5
()¢"
!:%"$$& #!: 6#:#5
¤
7
s
!
oeg
!
&§Ú
!
¼
5$%#%5
ÏKÛa|:wÇMâ7ª
_"
5
é¿&)F)"
!$9":$& 9"# 69":5
i*
!
´u=
!
\
5$%%;5!
¢vÍ&¬K>
&&E
[Ó
5
[Ó§¨§¬
!9":$& ;:; 6;>:5
G6X
5$%%95
æK
>Cb
ÇMâ7ñ=U¡Çú>*+
5
é
¿&)FB¯)í/u
5
@6V
!
úk
!
º)
!
¼
5$%%85
·UKÈöYm>
E,?P
[5
&()PQ
!#>">$& >:: 6>>%5
GJC
!
l{
5$%%#5
¹
E,?P
[Óå*)K>Ûw
5
KÓPQ®l
!$%"$$& ;> 6;;5
GQ
5$%%95
Çî)
5
B
&
t¼±§Ýv
5
G
7
^
!
6
7
B
!
bR9
5$%##5
KW§¨
)&+_&&EV
[äçè7Þ
¹
5
CͨDO)"
!#;"#$& :9 6>$5
G
7
:
!
7
!
&6
5$%%!5
x¹
E,?P
[Óå=WKYªî
Käîw>PQ
5
Ù=E))"
&
()v
!$>">,$& >8
6>95
G)u
!
$¨
!
tøÕ
!
¼
5$%%;5
vKn³Ë
_8_
Ü:wÇ>
Mâ Y _ " P Q
5
E C ( )
! 89 " $; $& #!%:# 6
#!%:8!#!#;95
G)u
!
$¨
5$%#%5
Ê¢vK¾Å×È
?!:%
Ç>MâY_"
PQ
5
ECE))"
!89"#$& ##> 6#$#5
G
7
â
!
bR9
!
¨:
5$%#%5
vKg1q
_8_
"Ëé
âY5[5
4Ñ&PQ
!$""8$& 9 6#8!$!5
!)û
!
b
!
7
!
¼
5$%%:5
ªKn·¦WX
E,?P
[Óå
>PQ
5
A=&)F)"
!$:"!$& ;" 6#%#5
b
7
k
!
t
7
!
{`
5$%#%5
vK(
_#!8_
|òìwÇ>Mâ7
5[5
[Ó§¨§¬
!""8$& :88 6:!#5
bR9
!
G
7
â
!
¨:
!
¼
5$%##5
vK$óô
*3
j
gA
j
u¯À
ÐWÇMâ7_"[5
CͨDO)"
!#; "#$& >;
69>5
D/HY^ ,! .IVI/4 +5$%%95+LF0FYH03SFY/0/JIVT1VJHIUFT/FQ@SFVVFU
VFuHF4YFT3JV5\/0FYH03S)Y/0/JW! #>":$& ;%9 6;$!5
L34JC-!LH34JAO!\F^VF
34U [/<[3SU
LF4 ?! ?34 P! B34 ! &.8%D$%##5K/0U 434/@3STIY0FVX/SLIJL_
TLS/HJL@HT JF4/TW@I4J /X 0/4J_S34JF L3@0/TW@FV5 *3THSF
*34/TFYL4/0/JW! >& >8; 6>!!5
LIH OD!-I4 A!L34J2!&.8%D$%%>5\/0FYH03SYL3S3YTFSI]3TI/4
34U FQ@SFVVI/4 /XX/HSYP*,VF4Y/UI4JVHYS/VFVW4TL3VFXS/
03S^-K! RL34JO! OF4UF02B5#;;:5,@LW0/JF4W/XTLFJS3VV
X3
UFh0IZFIS3, ! */Z3YK3SYI3,! SIVT/X34I\! &.8%D $%%$5
(UF4TIXIY3TI/4 /XYITSHVLW[SIUVTLS/HJL TLFY/<[I43TI/4 /X0F3X3@FQ
Y3FV@IT/VF [3<[// )&+_&&E <3S^FSVX/SYS/VV_V@FYIFVgJF4FS3
TS34VXFS3[I0ITW34U 3[I0ITWT/IUF4TIXWI4TFSV@FYIFVLW[SIUV5?034T
DSFFUI4J!#8%":$& :;> 6>%%5
BSI3S)!r/YLFSTK5#;;#5D3<[//JFS<@03V
K3/R\!-I-!?F4JRA5$%##5KF4FS3TI/4 34U 3430WVIV/XFQ@SFVVFU
VFuHF4YFT3JVXS/<34/S<30I]FU YP*,0I[S3SW/XW/H4J0F3XXS/<\3
[3<[//"\&*="$:8%8+;1%8.5(%$";1\H4S/$5?034TF0EF@/STV! 8%
"##$& $%!: 6$%:95
KFSVTFI4 \! RLF4JP5$%%>5+LFSF300IXF/X@VFHU/JF4FV5&YIF4TIXIY
,
JF4/
KH/RA! LF4 CC! -IPR5$%%$5?LW0/JF4FTIYVTHUIFV/4 TLF
>/8+*$:8%8+;1JS/H@ 34U ITV30IFV"KS3
)Z/0HTI/4! $$ "#$& $% 68%5
AIV3XS3J
X0/NFSI4J[FL3ZI/HS5rFNDH0! >$& !>8 6!985
A/U^I4V/4 +E! EF4Z/I]F& ,! L/4JL3I0FK*! &.8%D$%%%1,
/<@3SIV/4 /X(+& 4HY0F3SSP*,VFuHF4YFU3T334U ,B-?<3S^FSV
X/S @LW0/JF4FTIY VTHUIFV I4 !/,%$1.8:/,1 " D3<[HV/IUF3F!
?/3YF3F$52?034TEFV! ##8"8$& $:; 6$>;5
AVI3/2C! -FF& \5#;;;5KF4FTIYUIZFSVITW34U
I4 +3IN345\/0FYH03S)Y/0/JW! ""$$& $>8 6$9%5
AVI3/2C! EIFVFLFSJ-A5#;;!5?/@H03TI/4 JF4FTIYVTSHYTHSF/X
7;1/8*58 *5.8M8,8+&*151"D3<[HV/IUF3F! ?/3YF3F$ I4 +3IN345
\/0FYH03S)Y/0/JW! 8"8$& $%# 6$%"5
AVIFL -&!AVIFL C-!CFL &!&.8%D$%##1\/0FYH03SYL3S3YTFSI]3TI/4
/X3@LF4W03034I4F3<4I3_0W3VFJF4F"D/?,-#$ XS/
AVIFL -&! \3K2!C34J!&.8%D$%#%[50/4I4J! FQ@SFVVI/4! VITF_
UISFYTFU
& ( )
!"
"
7
#;;; 6$%%;5
AVIFL -&! CFL &! ?34 A! &.8%D$%#%350/4I4J34U FQ@SFVVI/4 /X
3@LF4W03034I4F3<4I3_0W3VFJF4F"C$!)Fe $ XS/< C8+A;18
$%=/8+5I4 <1:/&"5:/58 :$%534U !5:/58 E81.$"51D?S/TFI4 )Q@SFVVI/4
34U ?HSIXIY3TI/4! 9#"$$& $$! 6$8%5
rF0YL4FS& ,!03S^ -K5#;;95\/0FYH03SFZ/0HTI/4 34U @LW0/JF4FTIY
HTI0ITW/XTLFYL0/S/@03VTS@-0> I4TS/4 I4 #/;1H;&8 34U TLF
D3<[HV/IUF3F"?/3YF3F$5\/0FYH03S?LW0/JF4FTIYV34U )Z/0HTI/4!
""8$& 8": 68;95
-IC!r/S/0,D!B3LI<3+!&.8%5$%%!5\IYS/V3TF0ITFVNITLI4 JF4FV&
VTSHYTHSF! XH4YTI/4 34U FZ/0HTI/45\/0DI/0)Z/0! $# " > $& ;;#
6#%%95
-I4 G! L/N+C! LF4 AA! &.8%D$%#%35k4UFSVT34UI4J[3<[//
X0/NFSI4J[3VFU /4 03SJF_VY30F3430WVIV/XFQ@SFVVFU VFuHF4YFT3JV5
KF4FTIYV34U \/0FYH03SEFVF3SYL! ; "$$& #%": 6#%;85
-I4 G! -/H CB! -IH 2! &.8%D$%#%[5S/VV[SFFUI4J/X!/,%$1.8:/,1
V@FYIFV"?/3YF3F$ 34U IUF4TIXIY3TI/4 /XTLFISLW[SIUVHVI4J(&&E
<3S^FSV5KF4FTIYV34U \/0FYH03SEFVF3SYL! ; "8$& #8;" 6#!%!5
*3U/T&! DIT3SK! 3STFS-! &.8%D#;;:5,@LW0/JF4FTIY3430WVIV/X
4/Y/TW0FU/4V[3VFU /4 TLFYL0/S/@03VTJF4F"E1a! HVI4J@3SVI4W
34U 34FN4H
h4JA ! ?30
?F4JRA! -H ++! -I-D! &.8%D$%#%5KF4/
YP*,VFuHF4YFV5D\?034TDI/0/JW! #%& ##>5
E/TL[FSJ2\! AI4]O! EF3SIY^ +\! &.8%D$%##5,4 I4TFJS3TFU
VF
&I
"?/3YF3F& D3<[HV/IUF3F$ [3VFU /4 I4TFS430TS34VYSI[FU V@3YFS
VFuHF4YFV/X4HY0F3SSI[/V/<30P*,5DI/YLF
+34JPM! -H 22! B34JO! &.8%D$%##1PFZF0/@
OH ! O34J& 2! RL34JAD5$%%>5(4TFS3YTI/4V34JJF4/
3430WVIV/XTLF)"8A5=$E151./8%58*8 JF4/
OH BA!r34 P?!-FF& D!&.8%D$%%;1/<@0FTF4HY0F/TIUFVFuHF4YF
/X\&*="$:8%8+;1%8.5(%$";18*= C8+A;18 $%=/8+5 YL0/S/@03VT
JF4/
C34JAM! C34J2D! ?F4JRA! &.8%D$%%"5,0FYH03S@LW0/JF4FTIY
34U XSHITFZ/0HTI/43SW 3430WVIV/XTLF <3t/SJS/H@V /XTLF
@30F/TS/@IY30N//UW[3<[//V"KS3
\/0FYH03S?LW0/JF4FTIYV34U )Z/0HTI/4! !" "8$& "%; 6"$!5
C34J2D! C34JAM! -IPR! &.8%D$%#%1?LW0/JF4W/XC8+A;18 34U
ITV30IFV"?/3YF3F& D3<[HV/IUF3F$ I4XFSFU XS/< 4HY0F3SKD&&(
JF4F34U @03VTIU @V[,_TS4A! S@08$_TS4-34U S@V#> I4TS/4 P*,
VFuHF4YFV5+3Q/4! :; "!$& ##%$ 6###%5
C/^/A! A3SHTVHJH r! \I^I/r5$%%"1 kVF/XX0/NFSI4JJF4F
B-hO)E(*K-hk&+"B+$ L/0/JVI4 TLF@LW0/JF4FTIY3430WVIV
/X[3<[HV/IU 34U F3S0WUIZFSJI4JJS3VVFV52?034TEFV! #$#& !:# 6
!>#5
RL34JC2! \3?B! -IPR5$%##5AIJL_TLS/HJL@HTVFuHF4YI4J/XVIQ
[3<[//YL0/S/@03VTJF4/
#%1#89#gt/HS4305@/4F5%%$%:;>5
RL/4JA! RL/H \D! GH \! &.8%D$%#%5PIZFSVITW34U FZ/0HTI/4 /X
?/4J_0I^F F0F
RL/H \ D! -H 22! RL/4JA! &.8%D $%#%35PIVTSI[HTI/4 34U
@/0WVH[X3
/X+W#_:$E58 SFTS/F0F
RL/H \ D! C34J?! K3/? 2! &.8%D $%##35(UF4TIXIY3TI/4 /X
UIXFSF4TI30W FQ@SFVVFU VFuHF4YF T3JV I4 S3@IU0W F0/4J3TI4J
!/,%$1.8:/,1E;A&1:&*1 I4TFS4/UFV [W VH@@SFVVIZF VH[TS3YTIZF
LW[SIUI]3TI/45?034T\/0DI/0EF@! $;"#$& $$! 6$8#5
RL/H \D! RL34JC! +34JPM5$%##[5L3S3YTFSI]3TI/4 34U @SI<3SW
XH4YTI/4303430WVIV/XC6#X_9!3
RL/H \D! RL/4JA!+34JPM5$%##Y5(V/03TI/4 34U YL3S3YTFSI]3TI/4 /X
VFZF4TW_4I4FXH0_0F4JTL <3SI4FS_0I^FTS34V@/V3VFJF4FVI4 TLF
D3<[HV/IUF3FVH[X3
!
!"#$
7
_
7
@
"
>>#