全 文 :!24" !1#
Vol.24 No.1
! " # ACTAAGRESTIASINICA 2016 1%
Jan. 2016
犱狅犻:10.11733/j.issn.10070435.2016.01.015
¾
、
ó¿qWÞcbR5ÀÁe2|u w4q , , x,0yz,){v ( -_()9\]^¬^CË ÈYY+-_()*¾+, , 67 ÈY730020) 78 : »/0<=íS,°BC , h"6@×AT MnÓùDÜ(E+)6Ü29Þ(E-)É-(犃犮犺 狀犪狋犺犲狉狌犿犻狀犲犫狉犻犪狀狊)Gmh°T)cdj|eQW(salicylicacid,SA)D Mnú5Tcd。efgh, ¼î MnÓù¸,E+É-Gm°Ý6°»ë©
(犘<0.05)séE-É-Gm,Àì MnÓù×A TíY ,E+É-GmTh°Ý©
(犘<0.05),nE-É-Gm°Ý6°»ëï
غ;Àì MnÓù×ATíî,E+6E-É-}©h
TóI,nï
غ;LeSAÜÓù×A, E+É-}©
(犘<0.05)séE-É-},L10mg·L-1MnÓùá&=¾¼î6@×ATeSA, ef
eSAD MnÓùÞJcd。ÓS1ð,29Þ»BUVÉ-GmTh°,¸¬ú5DÉ -Gmh°ÞJß©T)cd , neSALjK-_.ßú5。 9:; :Mn;SA;É-;29Þ;Gmh° <=>?@ :S452 ABCDE:A AFG@:10070435(2016)01010707 犈犳犳犲犮狋狊狅犳犕犪狀犵犪狀犲狊犲犪狀犱犛犃狅狀犌犲狉犿犻狀犪狋犻狅狀狅犳犃犮犺狀犪狋犺犲狉狌犿犻狀犲犫狉犻犪狀狊 犛犲犲犱犆狅狀狋犪犻狀犻狀犵狅狉犉狉犲犲狅犳犈狀犱狅狆犺狔狋犲 WANZhiwen,WANGPing,ZHANGXingxu,LIChunjie (KeyLaboratoryofGrasslandFarmingSystems,ColegeofPastoralAgricultureScienceandTechnology, LanzhouUniversity,Lanzhou,GansuProvince730020,China) 犃犫狊狋狉犪犮狋:Variousconcentrationsofmanganese(Mn)andsalicylicacid (SA)affectedgerminationof drunkenhorsegrass(犃犮犺狀犪狋犺犲狉狌犿犻狀犲犫狉犻犪狀狊)seedsthatwereinfectedwith(E+)orfree(E-)ofa犈狆犻 犮犺犾狅犲/犖犲狅狋狔狆犺狅犱犻狌犿endophyte.TheresultsshowedthatE+specimensshowedasignificantly(犘<0.05) highergerminationrateandindexthanthatofunderMnfreetreatment.ThegerminationrateofE+speci mensshowedattendofdecreasingongerminationratewiththeincreasingofMnconcentration.Therewere significantdifferencesexistedbetweentreatments(犘<0.05);butthegerminationrateandindexofE- specimenshadnosignificantdifferences.E+specimensshowedasignificantly(犘<0.05)higherroot lengthwhenvariousconcentrationsofexogenousSAwereadded.TheresultsshowedthatexogenousSA couldrelieftheeffectsofMnonseedgermination. 犓犲狔狑狅狉犱狊:Mn;SA;犃犮犺狀犪狋犺犲狉狌犿犻狀犲犫狉犻犪狀狊;Endophyte;Germination HIJK :20141224;LMJK:20151013 NOPQ : ¬ ”973”´ µ(2014CB138702);¬¶·*+®¸´µ(31372366);öJúÁÂSÊ°±²³´µ(IRT13019)Å 5RST : u¹ (1992),È,67áQÉ,Ë/09,/0Òcnq-29Þt9N,Email:wanzhw14@lzu.edu.cn;<Ôc Authorforcorrespondence,Email:chunjie@lzu.edu.cn q-29Þ(Fungalendophyte)Úßà§B Lq-&A=àúYú=9ý¹#
,
I6ËÌ
JK
Geú;TÞ
[1]。
µvþÀLû
&%Fq-&°P¡犖犲狅狋狔狆犺狅犱犻狌犿 ¬|J1
,犈狆犻犮犺犾狅ё¬T29Þ[2],.à29ÞT+L
kJKThi§¥J1í
,
Ì«5ns¡
ÌÌJKDÜG9K
[3]
6È9Kú5
[4]
Tc
,
~øö
[56]、
ø
[78]、
ø
[910]、
)
[1112]
6D¸¬ú5
[13]
Tc
。
É-
(犃犮犺狀犪狋犺犲狉狌犿犻狀犲犫狉犻犪狀狊)Úq»*ÍÍ
-¬
(犃犮犺狀犪狋犺犲狉狌犿)9-»JK,ÚÕ:Ò á·-.Ì«T]Ú-ãß , LÂÜ , 234 、 ª 、 、 67? ( ) ©J=> [14]。1994V
+Bruehl?[15]LÔ¶ÕÂÜTÉ-G
m&°P¡29Þ
。2000Z¹F6eö|[16] L67É-2°P¡29Þ 犖犲狅狋狔狆犺狅犱犻 ! " # !24"
um,2004eö|?[17]L67É-2°P¡ 29ÞÂG犖.犵犪狀狊狌犲狀狊犲,}ÏÝÂwÞý ï⣠, PÂýïn犈狆犻犮犺犾狅犲犵犪狀狊狌犲狀狊犻狊[18]。2 9Þs¡É-Thi§¥ [19], Ls×AT ¸¬ú5É-TcJß©Ts [20]。 ÀìÃ)T°± 、 ETi> [21]、 (Î [22] 6ð [23] TY`d , ¸¬´µ [24] òô , n- ·Ó"TßYHlþ [25]。 jÜGïÂiOT ¸¬ô´µ¡ÙC6QF ; LJK&Y`Ö¹T ¸¬67ôUV¡JKPT9°J69ù 9ðoØ , nò]Lj<=&KàD!K6Éಠ³+ß©ÄAT°± [26]。 ¸¬ø (Mn)ÚJK 91ò3T?`üu , JKGÂtoØ= Ä , Ì«cncd&øÁð½T5m [27]。 K `T MnL[\JrP9Æíî¤`,I-× AQ=ß©<= , ÿDJK¤9Ú±cd [2829]。 Ï i½ , QW (salicylicacid,SA)LJKøHlú 5 [3031]( ~& 、 p 、 ) 、 、 ¸¬? ) 6± [32] Ò ÓÞJh
T:f , Lj¸¬TÚ± [33] 6) ú5DGmh°|ÈÉT)cd [34], íîJrT ¼69K` [35]。 Gmh°2#T9ö"#UVJKT9 °J6¤` [36], 5. , »BC/0L¸¬ Mn ÓùÍÖÜ6ÜÞÉ-GmTh°xö , W XSAD¸¬ú5Tcd,n\ß/0É -D¸¬ú5Tø 、 9ù9ð|=mÁù0 ©¡ß©®¯ 。 1 UVWXY 1.1 jkUV É-Gmé2011Ô¶67á¯0Ït(E
102°52',N37°12)ÆbGéÈYY+Ð&,
201210%_TGm5℃Ð+é()ú?- --GmF`|Ñ×C&! ( ÈY ) GmhË , 20133%ÜÞ(E+)66ÜÞ(E-)É-
u
,20147%Ð_TÉ-Gmdé»B C 。 1dBÜ MnCl2·4H2O、QW (salicylic acid,SA)©j=Úñâ¼î;QnA5mQ。 1.2 jkXY 1.2.1 Mnp[ESAE+pE-|e>34} ~E_ý Mn2+Óùá×A=¾n0,50,100,200 mg·L-1,SA Ó ù á × A = ¾ n 0,5,10,15 mg·L-1;j¼î?`A5mQnDÒ。fgÓ ²³ , h]W1òkAt`TGm , d5%TX WâÛẻ10min,jd¶çQ ¼ë,Ud °ôQ ¼3,dìíÐQ^。f"Ân9 cmTD¹,¹2jYìín°,,Ð`T ¸¬Ûáî7mLÂéD¹&。¿¹©Q> 30]É-Gm,¿MÓù3M*,MNGm Th°ë ( æ}Gm? 、 æAWdGmß , 34nGmþÀ° )。GZ025Fà¶!Ò Dà&ÛAn (20±1)℃,Ò2%n16D/8N。 °#%¿r24hMNß,!10d^²°Ý, Ʋ߰»ë 。 °Ý (%)=BGmT°ë/BGmë ×100。 °»ë (犌犐)= ∑犌狋/犇狋,â&犌狋nL狋ò T°ë ,犇狋n°áë。 1.2.2 [ESA MnWXdE+pE|e>3 4pE_ý GmÀ°ôQºG24h _ , bGé¼J2ìíTD¹,î10mg·L-1 Mn2+Óùá,LÓùá&¼î6@×A@ATSA, SA×A=¾n0,5,10,15mg·L-1,>ÂéÎÛà 228±1℃Íh°,h°7d_^²æ}6æ 。 1.2.3 Lê 1JëÏ©d Microsoft Excel2003N,Æcë。ÔdSPSS16.0forwin dows^²=ÕÖ\غ
=,dDun canâ\¸t。 2 pqW>r 2.1 MnnoE+wE-ÀÁe2|u
èg1Lã,Àì MnÓù×ATíî,E+É
-Gm°Ý6°»ë©h
TóI
,
nE-É-Gm°Ý6°»ëï
غ。
-MnÓù×An200mg·L-12,E+É-Gm
°Ý
(犘<0.05)é MnDÒ,¡×
12%;50,100,200mg·L-1MnÓùE+É-
Gm°»ë©
(犘<0.05)é MnDÒ,=
¾¡×8%,9%615%;L MnDÒ,E+É
-Gm°Ý6°»ë©
(犘<0.05)sé
E-É-Gm,Uh29ÞLj[\Gmh°;
L50mg·L-1MnÓùE+É-Gm°»ë
(犘<0.05)séE-É-Gm,èg3Lã,
801
!1# u¹?:ø、QW29ÞöcDÉ-Gmh°TUV
Mn629ÞDÉ-GmT°Ý6°»ë ÞJæöcd。
`1 c~x 犕狀noPÀÁe2
Table1 Seedsgerminationof犃犮犺狀犪狋犺犲狉狌犿犻狀犲犫狉犻犪狀狊underthedifferentconcentrationofMn
Mn
/mg·L-1
°Ý
Germinationrate/%
°»ë
Germinationindex
E+ E- E+ E-
0 96.67±3.33A 84.44±3.85a 21.58±0.08A 18.83±0.73a
50 88.89±1.92AB 86.67±5.77a 19.92±0.30B 17.94±0.49a
100 86.67±8.82AB 85.33±3.33a 19.61±1.04B 18.94±0.97a
200 85.56±5.09B 85.11±5.09a 18.42±0.46C 18.05±0.93a
ü:g&@j6@YU;ÇgÜÞJr(E+)%غ
(犘<0.05),uU;Çg6ÜÞJr(E-)%غ
(犘<0.05),gE
+6E-ã%غ
(犘<0.05),@
Note:Differentcapitalletterwithinthesamecolumnmeanssignificantdifferencewheninfectedendophyteatthe0.05level;differentlow
ercaseletterswithinthesamecolumnmeanssignificantdifferenceoffreeendophytetreatmentatthe0.05level. meanssignificantdifference
atthe0.05levelbetweenE+andE-.Thesameasbelow
2.2 SAE+wE-ÀÁe2|u èg2Lã,ÀìSAÓù×ATíî,E+ É-Gm°Ý6°»ë©jês_ TóI , @: ,E-É-Gm°Ý6° »ëÒjês_T:<ðóI 。 L0, 5,10,15mg·L-1SA Óù×A,E+É- Gm°Ý6°»ë©Øº6
; L10 mg·L-1SA Óù×A,E-É-Gm° Ý
(犘<0.05)séSADÒ,/Óù×A غ©6
; èg4Lã,29Þ6SA DÉ-T°Ý©J[\cd , IñîJ æöcd 。 `2 c~x犛犃noPÀÁe2 Table2 Seedsgerminationof犃犮犺狀犪狋犺犲狉狌犿犻狀犲犫狉犻犪狀狊underdifferentconcentrationofSA SA /mg·L-1 °Ý Germinationrate/% °»ë Germinationindex E+ E- E+ E- 0 92.22±1.92A 83.22±3.34b 19.03±0.05A 18.25±0.82ab 5 94.44±5.09A 88.90±1.92ab 19.08±0.96A 19.28±1.04a 10 94.44±5.09A 92.22±1.92a 19.86±0.59A 19.02±0.45a 15 88.89±5.09A 86.67±6.67ab 19.11±1.08A 16.88±0.35b 2.3 MnnoÀÁìwÂ2|u
~ë11,Àì MnÓù×ATíî,E+
6E-É-}©h
TóI。L Mn
n100,200mg·L-1Óù,E+É-}
(犘<0.05),0mg·L-1¸,=¾
¡18%645%,L Mnn50,100,200mg·L-1
Óù
,E-É-}
(犘<0.05),
0mg·L-1¸,=¾¡18%,34%653%;
E+É-À Mn×ATíîR,L
Mnn100mg·L-1Óù
(犘<0.05)é
DÒ
,
¡11%,E-É-À Mn×AT
íîÒRóI
,
L Mnn200mg·L-1
Óù
(犘<0.05)éDÒ,¡12%。
èg2Lã,Mn629ÞDÉ-GmTæ
}6æTUVغ
,
òñã%+L
æöcd
。
=1 犕狀noÀÁìwÂ2|u Fig.1 EffectsofMnonrootandshootlengthof犃犮犺狀犪狋犺犲狉狌犿犻狀犲犫狉犻犪狀狊seedlings 901 ! " # !24"
`3 犕狀wÞcbR5ÀÁewìÂ2|u Table3 EffectsofMnandendophyteon犃犮犺狀犪狋犺犲狉狌犿犻狀犲犫狉犻犪狀狊germinationandrootandgerminallength ÒØçSource ¶èAdf 犉
Sig. °Ý Germinationrate 29ÞEndophyte 1 0.07 0.79 Mn×A Mnconcentration 3 0.41 0.75 29Þ×Mn×AEndophyte×Mnconcentration 3 4.49 0.01 °»ë Germinationindex 29ÞEndophyte 1 31.08 <0.01 Mn×A Mnconcentration 3 9.10 <0.01 29Þ×Mn×AEndophyte×Mnconcentration 3 4.83 0.01 æ} Rootlength 29ÞEndophyte 1 0.71 0.41 Mn×A Mnconcentration 3 43.87 <0.01 29Þ×Mn×AEndophyte×Mnconcentration 3 1.64 0.22 æ Germinallength 29ÞEndophyte 1 14.04 <0.01 Mn×A Mnconcentration 3 4.06 0.03 29Þ×Mn×AEndophyte×Mnconcentration 3 2.19 0.13 2.4 SAnoÀÁìwÂ2|u
~ë21,ÀìeSA×ATíî,E+É
-}6©:SêóI
。
LeSA×
An15mg·L-12,E+É-}6©
(犘<0.05)séDÒ,©íî¡12%;E-É-
}6<ðóIR
,
©ï
غ
。
-e
SAÓù×An0,5,15mg·L-12,E+É-}
(犘<0.05)séE-É-}。èg3L
ã
,SA629ÞDÉ-æ}6æT9Þ
J
T[\cd
,
òñã%ÞJæöcd
。
=2 ÃA犛犃ÀÁìwÂ2|u Fig.2 EffectsofexogenousSAonrootandshootlengthof犃犮犺狀犪狋犺犲狉狌犿犻狀犲犫狉犻犪狀狊seedlings `4 犛犃wÞcbR5ÀÁewìÂ2|u
Table4 EffectsofSAandendophyteon犃犮犺狀犪狋犺犲狉狌犿犻狀犲犫狉犻犪狀狊germinationandlengthofrootandshoot
ÒØçSource ¶èAdf 犉
Sig.
°Ý
Germinationrate
29ÞEndophyte 1 7.99 0.01
SA×ASAconcentration 3 3.71 0.03
29Þ×SA×AEndophyte×SAconcentration 3 0.44 0.73
°»ë
Germinationindex
29ÞEndophyte 1 1.64 0.22
SA×ASAconcentration 3 2.74 0.08
29Þ×SA×AEndophyte×SAconcentration 3 0.22 0.89
æ}
Rootlength
29ÞEndophyte 1 285.43 <0.01
SA×ASAconcentration 3 6.90 <0.01
29Þ×SA×AEndophyte×SAconcentration 3 1.26 0.32
æ
Germinallength
29ÞEndophyte 1 15.09 <0.01
SA×ASAconcentration 3 7.17 <0.01
29Þ×SA×AEndophyte×SAconcentration 3 0.82 0.50
2.5 SAWÞcbR5 MnnoPÀÁe ìÂ2|u èë3Lã,L@×AT MnÓù,E +É-}ÀìSA×ATíîj_ êsTóI 。 DÒ¸ , -SA×An5mg ·L-12,E+É-}
(犘<0.05), 25%,E-É-}ÀìSA×ATíî :Sê , غ6
;E+É-ÀìSA 011 !1# u¹?:ø、QW29ÞöcDÉ-Gmh°TUV ×ATíîRóI<ð , غ6
,E- É-ÀìSA×ATíî:Sê,غ 6
。 èg5Lã,L MnÓù,29Þ 6SADÉ-æ}TæT9ÞJ
(犘 <0.05)T[\cd,òñDæ}T9ÞJ æöcd 。 =3 ÃA犛犃 犕狀z{PÀÁìwÂ2|u
Fig.3 EffectsofexogenousSAonrootandshootlengthofMntreated犃犮犺狀犪狋犺犲狉狌犿犻狀犲犫狉犻犪狀狊seedlings
`5 犛犃wÞcbR5 犕狀noPÀÁeìwÂ2|u Table5 TheinteractionofSAandendophyteonrootandgerminallengthof犃犮犺狀犪狋犺犲狉狌犿犻狀犲犫狉犻犪狀狊byMnstressed ÒØçSource ¶èAdf 犉
Sig. æ} Rootlength 29ÞEndophyte 1 40.67 <0.01 SA×ASAconcentration 3 4.17 0.02 29Þ×SA×AEndophyte×SAconcentration 3 6.36 <0.01 æ Germinallength 29ÞEndophyte 1 2.25 0.15 SA×ASAconcentration 3 1.59 0.23 29Þ×SA×AEndophyte×SAconcentration 3 3.08 0.06 3 Wp »BC/0¸¬ MnÓùE+6E- É-GmTh°xö , j|eSAD MnÓù Gmh°)Tcd , ÆòXy¡ Mn62 9Þ 、SA629Þ| MnÓùSA629 ÞTæöcd 。 3.1 MnwSAnoE+wE-ÀÁe
2|u
MnÚJK91ò3TEDüu,×AY
uÚg©cK¤`61FT5uãß
[37]。
}
Ï{L
[38]
³PBCLã
,
É-P913 Mn
TÝK×An0.01mmol·L-1,è.Lã,13
`ÿ
,
nÀìÃ)ðT6ÿ°±
,
ÙC6Y&
T¸¬»`4Yé.×A
[39],
=`T Mn(D
JK+9ùú5
,
)5^ÆòUVó¦Fo
Ø
[40],
DÉ-9K`TÖ¹6ÈÉTP9
°JJ)cd
[38]。
/0gh
,
q-29Þ§
sËÌJKDEDKFTÏd:Ý
,
Ín[JK9
[41],
q-29Þ]Lj<=UVJK
ã%Tð0cd[$)GmTh°
、
}6
É
[40,42]。
jSefQ
,
»/0°P
,
L Mn
DÒ
,E+É-Gm°Ý6°»ë©
séE-É-Gm,Uh29ÞT+LDÉ
-GmTh°ÞJ[\cd
,
.Zhang[19]/0
Ù
。
·n
,Hesse?[43]T/0gh,犖犲狅狋狔狆犺狅犱
犻狌犿¬T29ÞJs9^-(犔狅犾犻狌犿 狆犲狉犲狀狀犲)h°|Gm¤`T§¥,IDÞr\ J2°P , .G§¥Æ6Úß"+L , @}®5F T29ÞD^-Gmh°6¤`îJUV 。 .ßiyT+LL§BC&1dT29Þj |ÌÌJKT®5F9\FJ
。
Àì MnÓ
ù×ATíî
,E+É-Gm°Ý6°»ë
©h
TóI
,E-É-Gm°Ý6°
»ëÒóI
,
Iغ6
,
É-Gm
T°Ý6°»ëcNSE+séE-,D Mn
629Þ\=
,
ef
ñ+Læöc
d
。
UhL×A Mn(<50mg·L-1)Óù,2
9ÞLj MnDGmTú5,Is×A(>
100mg·L-1)T Mn67DÉ-Gm»BÞJ
)cd
,
]Lj\Gm2ú)29Þ@
}ÂtoØ
,
É-29Þt9NT¸
¬§¥
,
)ÁùJR\ß/0
。
111
! " # !24" 3.2 犕狀,犛犃wÞcbR5ÀÁeì
Â2|u
/0gh
,
eQWD¸¬¦ú5JK
Gmh°6ÈÉT9ÞJß©T:
[44],
SAÓù§CdDGmh°6ÈÉ9T-±
JrN2CAT,APX,SOD6POD?[45]øÁð
½É<ðJ
,SA]Ljs
WT»`
iÓ$)ú5DJKTÚ±
[46],
;Ÿ¬6¥
DH.T1B
[47]。
»BC&
,
Àì MnÓù×A
Tíî
,E+6E-É-}©h
Tó
I
,
nï
غ
,
Uhæ}D¸¬ MnT/
0séæ
,
ªö
[36]
/0Ù
,
.L§Úè
éæ}×Ø^QoÆÝjÑ1GÍ
,
k}L
Mnú5T2%\Ä6Ö¹`SYé,Íng
Pnc±ó
。
LeSAÜÓù×A,E+É
-}©
séE-É-},À=,SA
29Þ+Læöcd
,
UheSADÉ-2
9ÞJß©TUV
,
cdÁùJR\ß/0
。
»/0&
,
eSAD Mnú5ÞJi?Tc
d
,
.SAîØøÁð½TÉ6
WTÖ¹
Jß©]
[48]。
I:f6Úûh
,
.L§ Mn
Óù×AtJß©]
,
ÞN.5JR\ßT
/0
。
ÓS1ð
,
¸¬ MnDÉ-GmTh°Þ
J)cd
,
n29ÞT+LLj MnDG
m|ææ}TÚ±cd
。
K`TeSAD Mn
ú5AJß©Tcd
,
n29ÞSADÉ
-æ6æ}T9+Læöcd
。
Dé29
ÞDGmh°cdÁj|SAD29ÞTcd
¢R\ßT/0
。
AB
[1] SiegelMR,LatchGCM,JohnsonMC.Fungalendophytesof
grasses[J].AnnualReviewofPhytopathology,1987,(25):
293315
[2] WhiteJF,MorganJG.Endophytehostassociationsinforage
grasses.VII.犃犮狉犲犿狅狀犻狌犿犮犺犻狊狅狊狌犿,anewspeciesisolated
from犛狋犻狆犪犲犿犻狀犲狀狊inTexas[J].Mycotaxon,1987,28(1):
179189
[3] MoonCD,GuilauminJJ,RavelC,犲狋犪犾.New犖犲狅狋狔狆犺狅犱犻狌犿
endophytespeciesfromthegrasstribes犛狋犻狆犲犪犲and犕犲犾犻犮犲犪犲
[J].Mycologia,2007,99(6):895905
[4] ZhangXX,LiCJ,NanZB.Effectsofcadmiumstressonseed
germinationandseedlinggrowthof犈犾狔犿狌狊犱犪犺狌狉犻犮狌狊infected
withthe犖犲狅狋狔狆犺狅犱犻狌犿endophyte[J].ScienceChinaLifeSci
ences,2012,55(9):793799
[5] ªü,tª,eö|,?.qÔøÕ29ÞöcDÉ
-ÈÉT9ùUV
[J].-_+i,2008,16(3):244261
[6] {+Û,&6,{º.JK29Þt9NDÖöGOT
UV
[J].9\+i,2009,29(10):56185626
[7] Z¹F,eö|.q-29Þt9NL-_()]^&T
cd
[J].9\+i,2004,24(3):605616
[8] eö|.É-29Þt9N9K+9\+8T/0
[D].ÈY:ÈYY+,2005:25
[9] SabzalianMR,MirlohiA,SharifnabiB.Reactiontopowdery
mildewfungus,犅犾狌犿犲狉犻犪犵狉犪犿犻狀犻狊inendophyteinfectedand
endophytefreetalandmeadowfescues[J].AustralasianPlant
Pathology,2012,41(5):565572
[10]é.q-29Þt9ND--|.ÞTUV
[D].ÈY:ÈYY+,2011:38
[11]{O,eö|,¸ \,?.29ÞD¾Y^)TUV
[J].-_+i,2009,17(1):8892
[12]YinL,RenA,WeiM,犲狋犪犾.犖犲狅狋狔狆犺狅犱犻狌犿犮狅犲狀狅狆犺犻犪犾狌犿
infectedtalfescueanditspotentialapplicationinthephytore
mediationofsalinesoils[J].InternationalJournalofPhytore
mediation,2014,16(3):235246
[13]ªü,Z¹F,eö|.29Þsq-¸¬ú5T
/0\±
[J].-)*+,2014,31(8):14661474
[14]ü¹×.&-_«JÚJK[M].:;:&()O
Ó,
1997,166176
[15]BruehlG W,KaiserWJ,KleinRE.Anendophyteof犃犮犺
狀犪狋犺犲狉狌犿犻狀犲犫狉犻犪狀狊,anintoxicatinggrassofnorthwestChina
[J].Mycologia,1994,86(6):773776
[16]NanZB,LiCJ.犖犲狅狋狔狆犺狅犱犻狌犿innativegrassesinChinaand
observationsonendophyte/hostinteraction.In:VHPauland
PDDappriched.犘狉狅犮犲犲犱犻狀犵狊狅犳狋犺犲4狋犺犐狀狋犲狉狀犪狋犻狅狀犪犾犖犲狅狋狔
狆犺狅犱犻狌犿/犌狉犪狊狊犐狀狋犲狉犪犮狋犻狅狀狊犛狔犿狆狅狊犻狌犿.Soest,Germany,
2000:4150
[17]LiCJ,NanZB.Anew犖犲狅狋狔狆犺狅犱犻狌犿speciessymbioticwith
drunkenhorsegrass(犃犮犺狀犪狋犺犲狉狌犿犻狀犲犫狉犻犪狀狊)inChina[J].
Mycotaxon,2004,90(1):141147
[18]LeuchtmannA,BaconCW,SchardlCL,犲狋犪犾.犖狅犿犲狀犮犾犪狋狌狉犪犾
realignmentof犖犲狅狋狔狆犺狅犱犻狌犿 specieswithgenus犈狆犻犮犺犾狅ё
[J].Mycologia,2014,106(2):202215
[19]ZhangXX,FanXM,LiCJ,犲狋犪犾.Effectsofcadmiumstress
onseedgermination,seedlinggrowthandantioxidativeen
zymesin犃犮犺狀犪狋犺犲狉狌犿犻狀犲犫狉犻犪狀狊plantsinfectedwitha犖犲狅狋狔
狆犺狅犱犻狌犿endophyte[J].PlantGrowthRegulation,2010,60
(2):9197
[20]ZhangXX,LiCJ,NanZB.Effectsofcadmiumstresson
growthandantioxidativesystemsin犃犮犺狀犪狋犺犲狉狌犿犻狀犲犫狉犻犪狀狊
symbioticwith犖犲狅狋狔狆犺狅犱犻狌犿犵犪狀狊狌犲狀狊犲[J].JournalofHaz
ardousMaterials,2010,175(13):703709
[21]HirataH.Recentresearchprogressinautomotiveexhaustgas
purificationcatalyst[J].CatalysisSurveysfromAsia,2014,18
(4):128133
[22]PalmaP,KoeckSchulmeyerM,AlvarengaP,犲狋犪犾.Riskas
sessmentofpesticidesdetectedinsurfacewateroftheAlqueva
211
!1# u¹?:ø、QW29ÞöcDÉ-Gmh°TUV
reservoir(Guadianabasin,southernofPortugal)[J].Science
oftheTotalEnvironment,2014,488:208219
[23]RadicS,GregorovicG,StipanicevD,犲狋犪犾.Assessmentofsur
facewaterinthevicinityoffertilizerfactoryusingfishand
plants[J].EcotoxicologyandEnvironmentalSafety,2013,96:
3240
[24]BaiJM,LiuXP.Heavymetalpolutioninsurfacesoilsof
PearlRiverDelta,China[J].EnvironmentalMonitoringand
Assessment,2014,186(12):80518061
[25]{»®,t1.ÕÙCHlF`¸¬UV/0&ß}
küTþ
[J].()Hl*++i,2013,32(7):1289
1293
[26]WagnerGJ.Accumulationofcadmiumincropplantsandits
consequencestohumanhealth[J].AdvancesinAgronomy,
1994,51:173212
[27]MilaleoR,ReyesDíazM,IvanovAG,犲狋犪犾.Manganeseases
sentialandtoxicelementforplants:transport,accumulation
andresistancemechanisms[J].JournalofSoilScienceand
PlantNutrition,2010,10(4):470481
[28]DucicT,PoleA.Transportanddetoxificationofmanganese
andcopperinplants[J].BrazilianJournalofPlantPhysiology,
2005,17(1):103112
[29]BaldisserottoC,FerroniL,PantaleoniL,犲狋犪犾.Comparisonof
photosynthesisrecoverydynamicsinfloatingleavesofTrapa
natansafterinhibitionbymanganeseormolybdenum:Effects
onPhotosystemII[J].PlantPhysiologyandBiochemistry,
2013,70:387395
[30]YuanS,LinHH.Roleofsalicylicacidinplantabioticstress
[J].ZeitschriftfurNaturforschungc,2008,63(5/6):313320
[31]PalM,KovacsV,GSzalai,犲狋犪犾.Salicylicscidandabiotic
stressresponsesinrice[J].JournalofAgronomyandCrop
Science,2014,200(1):111
[32]RahmanA,KuldauGA,UddinWakar.Inductionofsalicylic
acidmediateddefenseresponseinperennialryegrassagainst
Infectionby犕犪犵狀犪狆狅狉狋犺犲狅狉狔狕犪犲[J].Phytopathology,2014,
104(6):614623
[33]wS¡,ÜOµ.QWDØú5»zÈÉ9ù8TUV
[J].Gm,2009,28(3):9093
[34]CavusogluK,TepeB,KilicS.Effectsofsalicylicacidpretreat
mentontheseedgermination,seedlinggrowthandLeafanato
myofbarleyundersalineconditions[J].CerealResearchCom
munications,2014,42(2):229238
[35]Khodary.Effectofsalicylicacidonthegrowth,photosynthesis
andcarbohydratemetabolisminsaltstressedmaizeplants[J].
InternationalJournalofAgricultureandBiology,2004(6):58
[36]ªö,eº,ù,?.¦、ÙD_xìíGmh°|ÈÉ
TUV
[J].¡:(+i,2005,20(1):9699
[37]íîø,«º.?`üuD^-¤`61FTUV[J].-
)*+
,2008,25(10):6669
[38]{L,ªü,«¾Ç,?.¸¬ú5DÉ-9|9ù9
ð»FTUV
[J].-)*+,2014,31(6):10801086
[39]LiuGN,YuYJ,HouJ,犲狋犪犾.Anecologicalriskassessmentof
heavymetalpolutionoftheagriculturalecosystemnearalead
acidbatteryfactory[J].EcologicalIndicators,2014,47:210
218
[40]DemirevskaKepovaK,SimovaStoilovaL,StoyanovaZ,犲狋犪犾.
Biochemicalchangesinbarleyplantsafterexcessivesupplyof
copperandmanganese[J].EnvironmentalandExperimental
Botany,2004,52(3):253266
[41]MalinowskiDP,AloushGA,BeleskyDP.Leafendophyte
犖犲狅狋狔狆犺狅犱犻狌犿犮狅犲狀狅狆犺犻犪犾狌犿 modifiesmineraluptakeintal
fescue[J].PlantandSoil,2000,227:115126
[42]é,eö|,Ï,?.Ú*-29ÞDñG--Gm
GÉTð0:
[J].-)+i,2010,19(4):3340
[43]HesseU,HahnH,AndreevaK,犲狋犪犾.Investigationsonthein
fluenceof犖犲狅狋狔狆犺狅犱犻狌犿endophytesonplantgrowthandseed
yieldof犔狅犾犻狌犿狆犲狉犲狀狀犲genotypes[J].CropScience,2004,44
(5):16891695
[44]PopovaLP,MaslenkovaLT,YordanovaRY,犲狋犪犾.Exoge
noustreatmentwithsalicylicacidattenuatescadmiumtoxicity
inpeaseedlings[J].PlantPhysiologyandBiochemistry,2009,
47(3):224231
[45]MetwalyA,FinkemeierI,GeorgiM,犲狋犪犾.Salicylicacidalevi
atesthecadmiumtoxicityinbarleyseedlings[J].PlantPhysi
ology,2003,132(1):272281
[46]YusufM,FariduddinQ,VarshneyP,犲狋犪犾.Salicylicacidmini
mizesnickeland/orsalinityinducedtoxicityinIndianmustard
(犅狉犪狊狊犻犮犪犼狌狀犮犲犪)throughanimprovedantioxidantsystem
[J].EnvironmentalScienceandPolutionResearch,2012,19
(1):818
[47]MishraA,ChoudhuriM A.Effectsofsalicylicacidonheavy
metalinducedmembranedeteriorationmediatedbylipoxygen
aseinrice[J].BiologiaPlantarum,1999,42(3):409415
[48]ParasharA,YusufM,FariduddinQ,犲狋犪犾.Salicylicaciden
hancesantioxidantsysteminBrassicajunceagrownunderdif
ferentlevelsofmanganese[J].InternationalJournalofBiologi
calMacromolecules,2014(70):551558
(
%&( / 0)
311