作 者 ：王乃江,张文辉*,陆元昌,范少辉,王勇
期 刊 ：生态学报 2010年 30卷 1期 页码：67~78
Keywords:interspecific association, Ziwuling, the Loess Plateau, natural secondary forest,
摘 要 ：用方差比率（VR）和基于2×2联列表,通过x2统计量检验、共同出现百分率（PC）、联结系数(AC)的方法，对地处黄土高原中部的陕西子午岭森林植物群落乔木层12个树种、灌木层20个树种和草本层21种植物的种间联结性进行了研究。结果表明，乔木层、灌木层总体种间关联性表现为显著正关联，草本层为不显著正关联。乔木层主要种群表现出相互独立的特点，但辽东栎(Quercus wutaishanica)和油松(Pinus tabulaefomis)，油松和茶条槭(Acer ginnal)，茶条槭和白桦(Betula platyphylla)，杜梨(Pyrus betulaefolia)与辽东栎、油松、茶条槭、白桦等7个种对存在一定正关联。正关联的种对数是副关联的2倍。灌木层南蛇藤(Celastrus orbiculatus)和悬钩子(Rubus corchorifolius)、水栒子(Cotoneaster maltiflorus)和红瑞木(Cornus alba)、多花胡枝子(Lespedeza floribund)和华北丁香(Syringa oblate)之间存在极显著的正联结；华北绣线菊(Spiraea fritschiana)和野葡萄(Vitis amurensis)、冻绿(Rhamuns utilis)和栓翅卫矛(Euonymus phellomanes)、南蛇藤和野葡萄、南蛇藤和葱皮忍冬(Lonicera ferdinandii)之间存在显著的正联结。正关联和副关联种对数相当。草本层种间联结表现出较为松散的特征，无关联的种对数多达21个。正副关联的种对数约为3∶2。最好将多个指标配合使用能更好地说明种间联结程度。
Abstract:We examined interspecific association of 12 tree species, 20 shrub species, and 21 herbacious species of the natural secondary forests at Ziwuling Bbased on the use of variance ratio (VR), x2 test and 2×2 contingency table, the overall interspecific association of the main 12 tree species, 20 shrub species and 21 herb species of natural secondary forest at Ziwuling were studied. There were positive associations among tree species and among shrub species. No relationship was detected among the herb species.The result showed that the interspecific association in tree layer or shrub layer were significantly positive while not significantly positive in herb layer. Among the tree species, seven pairs of species including The primary species in the tree layer were independent from each other, whereas seven pairs of species like Quercus wutaishanica and Pinus tabulaefomis, P. tabulaefomis and Acer ginnal, A. ginnal and Betula platyphylla, Pyrus betulaefolia and Q. wutaishanica, P. tabulaefomis, A. ginnal, and B. platyphylla showedhad positive associations; the number of species pairs having of positive association was twice two times that showing of negative association. in number. In shrub layer, three pairs of species includingsuch as Celastrus orbiculatus and Rubus corchorifolius, Cotoneaster maltiflorus and Cornus alba, and Lespedeza floribunda and Syringa oblate had highlyextremely significantly positive association;, and four pairs of shrub species including like Spiraea frischiana and Vitis amurensis, Rhamuns utilis and Euonymus phellomanes, C. orbiculatus and Vitis amurensis, and C. orbiculatus and Lonicera ferdinandii had were in significantly positive association. The number of pairs of species having The species pairs of positive association were approximately equivalent to those having of that of negative association. The associations among In herbaceous species were weak and inconsistant. layer, the interapecific association showed the dispersive feature, Twenty-one21 species pairs had nowithout interspecific associations, and the The ratio of species pairs of having positive association to species pairs of having negative association was 3∶2. The The several indicators applied together were proved proven to be more rational for describing the interspecific association.
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